1. Plant Structure and Growth

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    Last Updated:June 25, 2016 LMS SEGi education group 1

    PLANT ANATOMY AND PHYSIOLOGY

    WEEK 1 WEEK 2

    !"ant #or$ and #unction

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    LE%&'('G )*JE+(-ES

    Understand t.at t.e p"ant structure consists o/ &oots,Ste$s and Leaes

    *e aare o/ t.e /unctions o/ t.e roots, ste$s and "eaes

    Kno t.e transport o/ ater and /ood sustances in

    p"ants and .ere t.ese occur

    Kno so$e p"ant respond to t.e eniron$ent 3

    p.ototropis$, geotropis$ and tig$otropis$

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    LE%&'('G )U+)MES

    %t t.e end o/ t.is topic ou s.ou"d e a"e to:

    (denti/ and di//erentiate t.e crosssections o/ $onocot

    and dicot root and ste$ under t.e $icroscope

    State t.e structure and /unctions o/ 7"e$ and p."oe$

    (ndicate .ere actie and passie transport occur in a

    p"ant8

    9escrie t.e di//erent tpes o/ responses seen in p"ants8

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    Why are plants important?

    #ood

    S.e"ter

    )7gen !rotection

    ;aitat

    %est.etics

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    !"ant $orp.o"og t.e stud o/ t.e e7terna" structure o/

    p"ants

    !"ant anato$ t.e stud o/ t.e interna" structure

    A plant can be divided into 2 basic systems:

    Terrestrial plants face a divided habitat:

    Each system depends on the other

    Shoots provide sugars for roots (viaphloem)

    Roots provide H2! minerals and support for shoots (via "ylem)

    #$ Root system

    2$ Shoot system % stems! leaves & flo'ers

    Air source of 2for *HS

    Soil % provides H2 and minerals

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    The Root System

    4 /unctions:#$ Anchor plants

    2$ Absorb and conduct H2 & nutrients

    +$ Store food

    2 ma,or types of root systems:#$ Taproot system

    Seen in many eudicots

    Some are modified to store much reserve food

    2$ -ibrous root system *revent soil erosion

    Seen primarily in monocots

    Root hairs: enhance H2 absorption by ....................increasing surface area

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    The Shoot System

    Vegetative shoot = stem + leaves

    Floral shoot - terminates in flowers

    STEMS -on the shoot tip

    -has developing leaves

    -a.k.a. axillary bud-embryonic side shoot

    -usually dormant

    where leaves

    attach to stem

    Node

    Terminal

    bud

    Lateral

    bud

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    LEAES

    .e $ain p.otosnt.etic organs o/ a p"ant

    Usua"" a /"attened blade.ic. is "ea/ ase /or$s a

    sheatharound t.e ste$?

    Monocot "eaes para""e" $a

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    (s t.e eaporatie "oss o/ ater/ro$ p"ants

    (t is t.e super.ig.a /ornutrient transport+oo"s t.e p"ant)ccurs pri$ari" on t.e "oer

    "ea/ sur/ace

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    #ig 4@ 10

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    #ig8 4@810

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    TYPES of PLANT

    CELLS Note: Structure is related to function.

    1. PARENCHYMA CELLS

    least specialized plant cells

    primary walls are thin, flexible

    function - synthesizing and storing organic

    products

    2. COLLENCHYMA CELLS

    wall is uneven in thickness

    grouped into strands or cylinders to support young

    parts of plants

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    3. SCLERENCHYMA CELLS

    have very rigid, thick walls strengthened by lignin

    function - support

    4. WATER CONDUCTING CELLS: Tracheids & Vessel

    Elementsdead at functional maturity

    Tracheids - long, thin tapered cells; lignin; has pits for

    flow of H2O; functions: support &

    conduction

    Vessel elements - wider, shorter, thinner walled,

    less tapered; aligned end to end; end walls

    are perforated for H2O flow

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    Transport in Plants

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    En!o!ermis " #asparian strip

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    Mo$ement o% &ater " Apoplast an!

    Symplast

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    Apoplast an! Symplast

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    Mem'rane Transport

    Passi$e Transport( No ener)y re*+ire!, Di%%+sion o% a s+'stan-e %rom a re)ion o%

    hi)her -on-entration to one o% lo&er-on-entration &itho+t the help o% anytransport protein,

    Osmosis,the mo$ement o% &ater into -ells.is an e/ample o% passi$e !i%%+sion( the

    !i%%+sion o% &ater a-ross a mem'rane %roma re)ion o% hi)her -on-entration 0hypotonicor !il+te sol+tion1 to one o% lo&er-on-entration 0hypertonic or -on-entrate!sol+tion1,

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    A-ti$eTransport

    18 .e transport o/ sustances /ro$ a

    region o/ "oer concentration to a region

    o/ .ig.er concentration8

    %ctie transport reAuires energ

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    A-ti$e Transport

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    A-ti$e Transport 2 Passi$e Transport

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    .e ra//ic o/ BLargeB Mo"ecu"es

    En!o-ytosis an! E/o-ytosis( Me-hanisms%or transport in or o+t

    En!o-ytosis( 3 4in!s 0in all. the

    e/tra-ell+lar %a-es o% mem'ranes %+se to%orm $esi-les &hi-h enter the -ell5 -ell

    al&ays 6sees7 the -ytoplasmi- %a-e1,

    A, pha)o-ytosis( 6-ell+lar eatin)7

    8, pino-ytosis( 6-ell+lar !rin4in)7

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    Exocytosis:

    Essentially the reverse of

    endocytosis. For releaseof cellular contentsoutside the cell (here, the

    cytoplasmic faces ofvesicle and plasmamembrane fuse, so

    secretion can occur).

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    3, E/o-ytosis(

    Essentially the re$erse o%

    en!o-ytosis, 9or release o%-ell+lar -ontents o+tsi!e the -ell

    0here. the -ytoplasmi- %a-es o%

    $esi-le an! plasma mem'rane%+se. so se-retion -an o--+r1,

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    E/o-ytosis(

    :se! to e/port materials o+t o% -ell,

    Materials in vesicles %+se &ith -ell

    mem'rane an! are release! to

    o+tsi!e, Tear )lan!s e/port salty sol+tion,

    Pan-reas +ses e/o-ytosis to se-rete ins+lin,

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    SIEVE-TUBE MEMBERS: Food-conducting cells

    Are chains of phloem cells that transport sucrose, other

    organic compounds and minerals Are alive at functional maturity

    Their end walls have pores called sieve-tubes At least one companion cell is connected to each These cells nuclei and ribosomes serve the sieve-tube

    member which lack these organelles

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    Monocot Stem (xs)

    Dicot Stem (xs)

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    Dicot Stem (xs)

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    M t R t ( )

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    Monocot Root (xs)

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    (Apoplastic)

    (Symplastic)

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    Monocot Root (xs)

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    Dicot Root (xs)

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    Dicot Root Vascular Bundle

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    Root Vascular Cylinder and Cortex -- Dicot

    Cortex

    Endodermis

    Xylem

    Phloem

    Pericycle

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    Mass Flow -1

    Cort

    Endodermis

    Xylem

    Phloem

    Pericycle

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    Mass Flow - 2

    Cort

    Endodermis

    Xylem

    Phloem

    Pericycle

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    In shrubs like this tea plant

    (Camellia sinensis), the rootsystem will be more tap root than

    fibrous root.

    Notice the diameter of this tap

    root compared to this manswaist!

    But shrubs also generally have

    some compromise for uprootingforcesfeeder roots extending

    laterally.

    Tropical soils are nutrient poor

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    Tropical soils are nutrient poor.

    Roots must traverse the surface

    for minerals, so roots grow on

    the surface (no tap root).

    So, to keep this tall baobab tree

    standing upright, the roots grow

    in diameter but only in the

    vertical dimensions to formridge rootscalled buttress

    roots.

    These roots are a meter tall!

    These roots inspired gothic

    cathedral architects to design

    buttress walls.

    C h d l hi d i b ll

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    https://reader009.{domain}/reader009/html5/0328/5abad96aab0ba/5abad987a09b8.jpg

    http://www.oxc.com.hk/raoul_nathalie/gallery/images/04%20Buttress.jpg

    Cathedral architects to design buttress walls.

    Prop roots

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    Pandanus utilis- screw pine

    Prop roots

    C h d l hi d i ll

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    http://www.contrib.andrew.cmu.edu/~ajm/Pages/Graphics/flyingbuttress.JPG

    http://williamcalvin.com/BHM/img/FlyingButtressND.jpg

    Cathedral architects to design prop walls.

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    Monocot Leaf Monocot (cross section)

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    Dicot Leaf (cross section)

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    Dicot Leaf (cross section)

    !L%' G&)W;

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    !L%' G&)W;

    *egins it. seed ger$ination andcontinues /or t.e "i/espan

    Indeterminate growthC continued grot.

    as "ong as t.e p"ant "ies

    (nd8 grot. is $ade possi"e meristems

    M i t

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    Meristems; perpet+ally em'ryoni- tiss+es

    Meriste$atic ce""s Unspecia"iDed

    9iide to generate ne ce""s near t.e groing

    point

    Apical meristems Located in root tips and s.oot uds

    Supp" ce""s /or p"ants to gro in "engt.

    Primary growthC e"ongation initiated apica" $eriste$s

    /or$s pri$ar tissues organiDed into

    t.e 4 tissue sste$s

    LATERAL MERISTEMS

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    LATERAL MERISTEMS c"inders o/ diiding ce""s e7tending a"ong

    t.e "engt.s o/ roots s.oots

    add ne "aers o/ ascu"ar tissues

    produces toug.er secondar der$a" tissues Secondary growthC increased girt. !+e to

    lateral meristems

    t.ic=ening o/ roots and s.oots

    Gro&th o% Shoots

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    Gro&th o% Shoots

    PRIMARY GROWTH O9 SHOOTS $ost s.oot e"ongation is due to grot. o/ internodes e"o t.e s.oot ape7 grot. resu"ts /ro$ and it.in t.e internode a7i""arF"atera" uds $a /or$ ranc.es "ater in t.e p"ants "i/e

    SE#ONDARY GROWTH secondary plant bodyresu"ts /ro$ secondar grot.

    >grot. in dia$eter?

    2ndar grot. resu"ts /ro$ 2 "atera" $eriste$s:

    vascular cambium produces 2ndar 7"e$ and p."oe$ cork cambium produces a toug., t.ic= coering /or roots and

    ste$sH rep"aces epider$is

    accu$u"ated "aers o/ 2ndar 7"e$ produces wood.ic. consists $ost" o/ trac.eids, esse" e"e$ents

    and /ier

    Root Tip Di%%erent

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    Root Tip " Di%%erent

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    Geotropism

    )sere t.e picture e"o8

    .e ste$ disp"as negatie geotropis$ecause it gros opposite to t.e pu"" o/ grait

    Geotropism

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    Root Tip Senses Gravity

    Auxin Hormone ProducedAuxin Accumulates on Lower

    Growth Inhibited on Lower

    Relative to Upper

    Root Curves Downward

    Geotropism

    Root Hairs Increase Surface

    AreaRoot Hairs Secrete Acid (H+)

    H+Cation Exchange w/Minerals

    Mineral Uptake into Roots

    Water and Mineral

    Uptake

    Geotropism

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    ;or$one au7in rings positie geotropis$

    %u7in in.iits grot. o/ root ce""s so t.ece""s in t.e upper sur/ace e"ongate and t.e

    root cures donard

    %u7in sti$u"ates t.e grot. o/ ste$ ce""sso t.e ce""s e"ongate and cures upard

    Geotropism

    Phototropism

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    W.at causes ce""s in s.ad area o/ ste$ to

    e"ongateI .e p"ant .or$one au7in $igrates to s.ad

    area causing t.e ce""s to e"ongate

    Phototropism

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    Thi)motropism

    .is is caused uneAua" grot. due to

    contact it. so"id o

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    Thigmotropism

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    &E#E&E'+ES

    (ntroductor !"ant *io"og Stern, *id"ac= and Jans=

    200, 11t.Ed8, McGra ;i""

    (ntegrated *io"og ri"", Enger and &oss 200, 4 rd

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