Download - Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass Genome Evolution

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    2/15

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    3/15

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    4/15

    vdjabo kubdf`{dlj o~oj{ qhoj {ql kubdf`{dljv qo}o uhtvdf`bbt

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    5/15

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    `jk q>'q1,}='}> (ko{`dbok dj Vuubogoj{ b [`mbo ; ljbdjo$*

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    kobo{dlj mdjv `}o djkdf`{ok `ml~o {ho 6M fh}lglvlgo" qho}o v }dfo fh}lglvlgo 0 dv kd~dkok dj{l 05'Gm voagoj{v*

    O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 0?

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    6/15

    `j`btsok =5 u`}`blalv vopojfov }`jklgbt vobof{ok n}lg {ho

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    Flbdjo } qho`{" }dfo" vl}ahg" `jk g`dso fh}lglvlgov `}o kdvub`tok lj {ho v`go bdjo dj {ho ffa}o* Kubdf`{dljv `}o djkdf`{ok qd{h vlbdk bdjov* [ho ff~o

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    nljk lj l}{hlblalv fh}lglvlgov dj ml{h vuofdov `}o djkdf {ok qd{h klmbo `}}lqv*

    0> [ho Ub j{ Fobb

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    7/15

    u`{{o}jv mo{qooj }dfo" qho`{" vl}ahg" `jk g`dso qd{hdj o`fh

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    k`{ov m`fi {l ` QAK ln {ho `jfov{}`b a}`vv aojlgo qd{h ff~o

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    q`v }ofoj{bt fljff}gok mt Qod o{ `b* (=551$ dj ` }ofljv{}f{dlj ln

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    (Nda}o 6$ }oof{v {ho v`go u`{{o}j* [hdv q`v `bvl fljff}gok mt

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    vlgo {h}lah ` nvdlj mo{qooj {ho @; `jk @> `jfov{}`b fh}l'

    glvlgov (Nda}o ?" o~oj{ =$* [ho kubdf`{dlj mo{qooj q> `jk

    q1 q`v jl{ ko{of{ok dj qho`{ mof`vo ln {ho u}o~dlvbt goj'

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    g>" g?" `jk g4" mo{qooj fh}lglvlgov g; `jk g?" `jk mo'

    {qooj fh}lglvlgov g= `jk g05 `v ` }ovb{ ln {o{}`ubldkds`{dlj

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    g;'g?" `jk g='g05 }oof{ {ho `jfov{}`b }ob`{dljvhduv mo{qooj

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    flgu`}dvlj" qo h`k dkoj{dffok vo~oj vh`}ok kubdf`{dljv" m{

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    (@$ Vfhog`{df }ou}ovoj{`{dlj ln {ho vo~oj kubdf`{ok }oadljv vh`}ok mo{qooj qho`{ `jk }dfo* [ho u`}`blalv }oadljv `}o }ou}ovoj{ok qd{h {ho v`go

    flbl}v dj qho`{ ({lu$ `jk }dfo (ml{{lg$ `v qobb `v {ho fl}}ovuljkdja l}{hlblalv }oadljv dkoj{dffok mo{qooj {ho {ql vo{v ln fh}lglvlgov (foj{o}$* [ho

    flbl} flko dv `v nlbblqv3 q0'q=,}?'}; (}ok$" q0'q6,}?'}0 (l}`jao$" q0'q;,}05'}6 (a}ooj$" q='q;,}1'}6 (bdah{ mbo$" q='q1,};'}8 (u}ubo$" q?'q1,}4'}8

    (m}lqj$" `jk q>'q1,}='}> (k`}i mbo$* ]dfoqho`{ l}{hlblav jl{ dj~lb~ok dj vh`}ok kubdf`{dljv `}o djkdf`{ok dj a}`t*

    (M$ Vfhog`{df }ou}ovoj{`{dlj ln {ho kubdf`{dljv vh`}ok mo{qooj }dfo fh}lglvlgov ? {l 0 (;= u`}`blav bdjiok mt hl}dslj{`b bdjov `{ ml{{lg$ `jk qho`{

    fh}lglvlgov 0M {l 6M (ff~o u`} blav bdjiok mt hl}dslj{ b bdjov `{ {lu$* [ho }dfo `jk qho`{ flbdjo } }oadljv }0'q6M jk }?'q0M`}o bdjiok mt 0;4`jk >>l}{hlblav (~o}{df`b bdjov$" }ovuof{d~obt*

    O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 01

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    8/15

    Nda}o ?* Glkob nl} {ho V{}f{}`b O~lb{dlj ln {ho ]dfo" Qho`{" Vl}ahg" `jk G`dso Aojlgov n}lg ` Flgglj @jfov{l} qd{h j ? Fh}lglvlgov*

    08 [ho Ub j{ Fobb

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    9/15

    {ho l{ho} {ql (}?'};,q='q0 `jk };'}8,q='q1$ `}o nljk lj

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    [ho `j`btvdv ln kubdf`{dljv {h`{ (0$ `}o vh`}ok mo{qooj {ho}dfo" qho`{" g`dso" `jk vl}ahg aojlgov" (=$ kl jl{ l~o}b`u

    qd{h o`fh l{ho}"`jk (6${dbo{lao{ho}{l fl~o} {hoaojlgov`bglv{

    oj{d}obt bok{l {hoglkob ln o~lb{dlj {h { dv u}ovoj{okdj Nda}o ?"

    dj qhdfh `j `jfov{}`b aojlgo qd{h ff~o fh}lglvlgov (@?" @1"

    @00" @8" `jk @>$ jko}qoj{ ` QAK (o~oj{ 0$ {h`{ }ovb{ok dj ff~o

    `kkd{dlj`b fh}lglvlgov3 @0" @05" @0=" @4" `jk @>* [hdv {o{}`'

    ubldkds {dlj q`v nlbblqok mt {ql {}`jvblf {dljv,nvdljv {h`{

    h`~o }ovb{ok dj {ql joq fh}lglvlgov" @6 (@05 @1$ `jk

    @=(@;@>$ (o~oj{=$" `jk`j dj{o}gokd {o`jfov{l} qd{hj0=

    (? ? =$ fh}lglvlgov* Vmvopoj{bt" {ho kdnno}oj{ a}`vv

    aojlgov qlbk h`~o o~lb~ok kdnno}oj{d`bbt n}lg {hdv `jfov{}`b

    aojlgdf v{}f{}o* Dj }dfo" `kkd{dlj b voagoj{ b kubdf`{dljv

    lff}}ok qd{hl{ glkdntdja {ho m`vdf v{}f{}o ln 0= fh}lgl'

    vlgov* [hv" {ho =4 kubdf`{dlj o~oj{v dkoj{dffok dj {ho }dfoaojlgo }ovb{ n}lg {h}oovffovvd~o }ljkv ln kubdf`{dljv3 (0$ `

    QAK (o~oj{ 0$ `jk {ql fh}lglvlgo {}`jvblf {dljv (o~oj{ =$9 (=$

    == `kkd{dlj b voagoj{`b kubdf {dljv l~o}b`uudja qd{h {ho

    QAKv9 `jk (6$ }ofoj{ kubdf`{dljv l~o} 96 Gm `{ {ho {o}gdj`b

    ojkv ln fh}lglvlgov }00 `jk }0=*

    N}lg {ho `jfov{}`b aojlgo qd{h 0= fh}lglvlgov" qho`{

    jko}qoj{ ff~o fh}lglvlg`b nvdljv (o~oj{ 6 dj Nda}o ?$ mo'

    {qooj @? `jk @05" @> `jk @8" @4 `jk @0=" @6 `jk @00" `jk @;

    `jk @1" }ovb{dja dj fh}lglvlgov q0" q1" q?" q;" `jk q="

    }ovuof{d~obt* Fh}lglvlgov q6 `jk q> l}dadj`{ok kd}of{bt n}lg

    {ho `jfov{}`b fh}lglvlgov @0 `jk @=" }ovuof{d~obt* [hdv

    }ovb{ok dj `j `jfov{}`b qho`{ aojlgo qd{h j 1 fh}lglvlgov

    (? ? = ?$* [hv" {ho 05 kubdf`{ok }oadljv lmvo}~ok ho}o dj

    {ho qho`{ aojlgo }oof{ {ho `jfov{}`b QAK (o~oj{ 0" nl} vo~oj

    ln {ho kubdf`{dljv$ `jk {h}oo `kkd{dlj`b voagoj{`b kubdf`{dljv

    {h`{ h`~o lff}}ok vdjfo {ho fh}lglvlg`b nvdljv*

    Nl} g`dso `jk vl}ahg" l} ffjkdjav `jk glkob `}o dj flg'

    ubo{o `a}oogoj{ qd{h {ho }ofoj{ `j btvdv ln Qod o{ `b* (=551$" qhl

    vhlqok {h`{ ml{h h`~o o~lb~ok n}lg `j `jfov{}`b aojlgo qd{h

    0= fh}lglvlgov `n{o} {ql fh}lglvlg`b nvdljv (mo{qooj @6

    `jk @05 `jk mo{qooj @1 `jk @49 Nda}o ?$" }ovb{dja dj `j dj'

    {o}gokd`{o `jfov{l} qd{h j 05 fh}lglvlgov (? ? = =$

    (o~oj{ ; dj Nda}o ?$* [hoj" g`dso `jk vl}ahg o~lb~ok djko'

    uojkoj{bt n}lg {hdv `jfov{l}* Qhdbo {ho vl}ahg aojlgo v{}f'

    {}o }og`djok vdgdb`} {l {ho `jfov{}`b aojlgo" g`dso jko}qoj{

    ` QAK }ovb{dja dj{l `j dj{o}gokd`{o qd{h j =5 fh}lglvlgov

    (o~oj{ ? dj Nda}o ?$* [hdv fl}}ovuljkv {l {ho {o{}`ubldkds`{dlj

    o~oj{ kovf}dmok dj u}o~dlv v{kdov (A`{ `jk Klombot" 04419

    Vqdaljl~` o{ `b*" =55;$* Nlbblqdja {hdv o~oj{" jgo}lv fh}l'

    glvlg`b nvdljv h`~o bok {l ` aojlgo v{}f{}o qd{h 05 fh}l'

    glvlgov (j 05 Y? ? = =W 6 = 05$* @{ bo`v{ 01

    fh}lglvlg`b nvdljv (o~oj{ > dj Nda}o ?$ gv{ h`~o lff}}ok {l

    oxub dj {ho }ob {dljvhduv {h`{ f`j mo lmvo}~ok{lk`t mo{qooj {ho

    kdnno}oj{ g`dso fh}lglvlgov*

    [hv" qd{h {hdv glkob {h`{ u}lulvov ` }ofljv{}f{dlj ln {ho}dfo" qho`{" vl}ahg" `jkg`dso aojlgov n}lg `j `jfov{l} qd{h

    j ? fh}lglvlgov" d{ dv ulvvdmbo {l dggokd`{obt dkoj{dnt {ho

    `jfov{} b }ob`{dljvhduv jk {ho l}dadj (QAK" m}o`i`ao" nvdlj$ ln

    {ho kdnno}oj{ fh}lglvlgov dj o`fh ln {ho nl} aojlgov*

    KDVF_VVDLJ

    Dj{} aojlgdf `jk Dj{o}aojlgdf Flgu`}dvljv _vdja

    Dgu}l~ok Dj{oa}`{d~o @bdajgoj{ F}d{o}d` ]o~o b Vh`}ok

    Kubdf`{dljv mo{qooj ]dfo `jk Qho {

    Dj {hdv v{kt" qo vok dgu}l~ok dj{oa}`{d~o vopojfo `bdajgoj{

    f}d{o}d` (FDU `jk F@BU$ flgmdjok qd{h ` v{`{dv{df`b ~`bdk`{dlj {l

    }o`j`btso {ho umbdfbt `~`db`mbo k`{` vo{ ln }dfo `jk qho`{ vo'pojfov* [hdv `bblqok v {l dkoj{dnt l}{hlblav `jk u`}`blav qd{h

    fljffkojfo `jk {l djno} joq }ob`{dljvhduv flgu`}ok qd{h u}o~d'

    lv v{kdov {h { qo}o od{ho} m`vok ljbt lj vg bb'vf bo vopojfo

    flgu`}dvljv (Atl{ o{ `b*" =55;9 Vdjah o{ `b*" =55;9 Mobb o{ `b*"

    =55?9 ]dfo Fh}lglvlgo 6 Fljvl}{dg" =55?$ l} b`fiok v{`{dv{d'

    f`b ~`bdk {dlj qhoj uo}nl}gok `{ {ho qhlbo aojlgo bo~ob (Vl}}obbv

    o{ `b*" =5569 B` ]l{` `jk Vl}}obbv" =55;9 Vdjah o{ `b*" =551$*

    Nl}go} flgu`}`{d~o v{kdov qo}o uo}nl}gok qd{h ` vopojfo

    `bdajgoj{ {h}ovhlbk ln 85/ dkoj{d{t l~o} 055 m`vov nl} {ho mov{

    HVU (B` ]l{` `jk Vl}}obbv" =55;$ l} qd{h ` vfl}o ln =55 nl}

    MB@V[J `j`btvdv (Vdjah o{ `b*" =551$* Qo h`~o vhlqj u}o~dlvbt

    {h { {hovo ~`bovg`t bo`k {l {hogdvdkoj{dfff`{dlj ln l}{hlblalv

    }oadljv (V`bvo o{ `b*" =55=" =55;$" `v {hot `bblq {ho ko{of{dlj ln

    gogmo}v ln aojo n`gdbdov {h`{ `}o jl{ {}bt l}{hlblalv l} ln

    hlglbladov {h`{ `}o m`vok ljbt lj fljvo}~ok klg`djv*

    [l ~`bdk`{o l} v{}`{oat" qo ff}v{ `uubdok l} v{}djaoj{ `bdaj'

    goj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk`{dlj {l {ho }dfo aojlgo* Qo

    fljff}gok `bb ln {ho kubdf`{dljv }oul}{ok u}o~dlvbt (T o{ `b*"

    =55?$ `jk dkoj{dffok 04 `kkd{dlj`b }oadljv kubdf`{ok mo{qooj

    kdnno}oj{ }dfo fh}lglvlgov* T o{ `b* (=55?$ h`k ov{dg`{ok {h`{

    {ho kubdf`{dljv ln {ho }dfo aojlgo fl~o} >?*1/ ln {ho aojlgo*

    Dj l} `j`btvdv" {ho v`go }oadljv (05 kubdf`{dljv$ fl~o} ;1*8/

    ln {ho aojlgo* [ho kdnno}ojfo dv ko {l {ho `ff}`ft ln {ho

    go{hlkvok {l ko{o}gdjo{ho bdgd{vln {hokubdf`{ok }oadljv* T

    o{ `b* (=55?$ `klu{ok ` a}`uhdf`b `uu}l`fh mof`vo ln {ho m`fi'

    a}ljk jldvo u}lkfok mt MB@V[* _vdja gl}o v{}djaoj{ f}d{o}d`

    `bblqok v {l ko{o}gdjo {ho bdgd{v ln {ho kubdf {ok }oadljv

    qd{h hdah fljffkojfo `jk `ff}`ft" {ho}omt }okfdja {ho vdso ln

    {ho }oadljv flgu`}ok qd{h {hod} `j`btvdv* [ho `ff}`ft `jk

    Nda}o ?* (flj{djok$*

    Fh}lglvlgov `}o }ou}ovoj{ok qd{h flbl} flkov {l dbbgdj`{o {ho o~lb{dlj ln voagoj{v n}lg ` flgglj `jfov{l} qd{h ff~o fh}lglvlgov* [ho ff~o

    fh}lglvlgov `}o j`gok `ffl}kdja {l {ho }dfo jlgojfb`{}o* Kdnno}oj{ o~oj{v {h`{ h`~o vh`uok {ho v{}f{}o ln {ho kdnno}oj{ a}`vv aojlgov k}dja

    {hod} o~lb{dlj n}lg {ho flgglj `jfov{l} `}o djkdf`{ok qd{h &* Nl} o`fh vuofdov" fh}lglvlgo jgmo}v `}o nlbblqok mt ` nl}gb` djkdf`{dja {ho

    o~lb{dlj }t l}dadj ln {hdv jgmo} (o*a*"dj qho {"j 1 ? Y`jfov{l}W ? YQAKW = Y`joubldk voagoj{ b kubdf {dljW ? Yfh}lglvlgo nvdljW$*Qho}o

    ulvvdmbo" kd~o}aojfo {dgov `}o djkdf`{ok m`vok lj u}o~dlv ov{dg`{ov mt A`{ (=55=$" Vqdaljl~` o{ `b* (=55;$" `jk U`{o}vlj o{ `b* (=55;$*

    O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 04

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    10/15

    }lmv{jovv ln l} go{hlk `}o vuul}{ok mt }ovb{v lm{`djok mt

    {ho ]dfo Fh}lglvlgov 00 `jk 0= Vopojfdja Fljvl}{d` (=55?$"

    qhl ov{dg`{ok {ho boja{h ln {ho glv{ }ofoj{ kubdf`{dlj lj {ho

    ~o}t kdv{`b ojk ln fh}lglvlgov 00 `jk 0= `v 6*6 Gm* [hdv

    fl}}ovuljkv ox f{bt {l {ho boja{h {h { qo nljk qd{h l} go{hlk

    (voo Vuubogoj{`b [`mbo 0 ljbdjo$ `jk dv vg`bbo} {h`j {ho ov{d'g`{ov (>*? `jk;*8 Gm lj fh}lglvlgov 00 `jk 0=" }ovuof{d~obt$

    ln T o{ `b* (=55?$ nl} {ho v`go }oadlj* Gl}ol~o}" l} `j`btvdv

    `bblqok v {l ko{of{ 0> jl~ob kubdf`{dljv {h`{ `}o vuo}dgulvok

    lj }oadljv `b}o`kt dkoj{dffok `v kubdf`{ok" {ho}omt koffjdja `k'

    kd{dlj`b `jfov{}`b }ob`{dljvhduv mo{qooj {ho }dfo fh}lglvlgov*

    Dj{o}ov{djabt" {hovo }ob {dljvhduv qo}o jl{ lmvo}~ok mo{qooj {ho

    l}{hlblalv fh}lglvlgov dj {ho l{ho} a}`vv aojlgov* [hdv

    vaaov{v {h`{ {ho vuo}dgulvok kubdf`{dljv lmvo}~ok ho}o dj

    }dfo `}o{}`fov ln ~o}t `jfdoj{ kubdf`{dljv {h { u}ok`{o {hoQAK

    dkoj{dffok dj {ho a}`vv `jfov{l}* @kkd{dlj`b a}`vv aojlgo vo'

    pojfov `v qobb `v `kkd{dlj`b {llbv {h`{ `bblq {ho ov{dg`{dlj ln

    kd~o}aojfo {dgov mo{qooj ~o}t `jfdoj{ kubdf`{dljv qd{h hdah

    `ff}`ft `}o jookok {l vuul}{ {hdv htul{hovdv*

    Dj qho`{" qo dkoj{dffok 05 kubdf`{ok }oadljv {h`{ }ou}ovoj{>1*?/ ln {ho aojlgo* U}o~dlv ]NBU g`uudja v{kdov h`k

    vaaov{ok `jfdoj{ kubdf`{dljv dj {ho qho`{ aojlgo* Nl} ox'

    `gubo"Kmfl~vit o{ `b*(044>$ vhlqok{h`{ 60/ln {ho]NBU blfd

    dj {ho kdubldk [}d{dfg gljlflffg g`u `}o u}ovoj{ gl}o {h`j

    ljfo dj {ho aojlgo" `jk Pd o{ `b* (=55;$ }oul}{ok {h`{ 04/ ln {ho

    OV[ g`}io}v vok nl} kobo{dlj mdj g`uudja qo}o nljk lj jlj'

    hlglolblalv vo{v ln fh}lglvlgov* _j{db jlq" hlqo~o}" {ho

    ox{oj{ `jk blf`bds`{dlj ln {ho kubdf {ok }oadljv qo}o jl{ koffjok

    u}ofdvobt" `jk l} }ovb{v u}l~dko ` udf{}o ln kubdf`{dljv dj {ho

    qho`{ aojlgo* [ho 05 dkoj{dffok kubdf`{dljv `}o u}lm`mbt `j

    jko}ov{dg`{dlj" `jk d{ dv ulvvdmbo {h`{ ` jgmo} ln kubdf`{dljv

    {h`{ qo}o jl{ v{ {dv{df bbt ~`bdk`{ok dj {hdv v{kt (o*a*" q='q6"

    q='q?" `jk q?'q>$ qdbb mo fljff}gok dj {ho n{}o `v `kkd{dlj`b

    aojo'g`uudja djnl}g`{dlj moflgov `~`db`mbo* @{ {hdv {dgo" l}

    `j`btvov kogljv{}`{o `b}o`kt {h`{ glv{ ln {ho aojlgo dv kubd'

    f`{ok `jk v{}ljabt vaaov{ `j `jfdoj{ kubdf`{dlj ln {ho kdubldk

    qho`{ aojlgov monl}o {hod} htm}dkds`{dlj dj{l ulbtubldk qho`{*

    Nlbblqdja {ho v`go `uu}l`fh" qo }oox`gdjok {ho flbdjo`}d{t

    mo{qooj }dfo `jk qho`{ `jk dkoj{dffok 06 flbdjo`} }oadljv {h`{

    fl~o} 86*0 `jk 45*;/ ln {ho }dfo `jk qho`{ aojlgov" }ovuof'

    {d~obt* U}o~dlv flgu`}`{d~o `j`btvov mo{qooj {ho }dfo k}`n{

    vopojfov `jk qho`{ OV[v (Vl}}obbv o{ `b*" =5569 Fljbot o{ `b*"

    =55;9 B` ]l{ `jk Vl}}obbv" =55;9 Bdjidoqdfs o{ `b*" =55;9

    Gji~lbk o{ `b*" =55;9 ]`jkh`q` o{ `b*" =55;$ `bvl nljk 06 l}'

    {hlblalv voagoj{v mo{qooj {ho {ql aojlgov* Ho}o" qo qo}o

    `mbo {l dkoj{dnt gl}o u}ofdvobt {ho bdgd{v ln {ho l}{hlblalv

    }oadljv `jk {l ko{of{ gl}o }o`}}`jaogoj{v {h`j u}o~dlvbt

    kovf}dmok vdja ` flgu`}dvlj mo{qooj {ho flgubo{o }dfo

    aojlgo vopojfo (Dj{o}j`{dlj`b ]dfo Aojlgo Vopojfdja

    U}lcof{" =55?$ `jk ` jdpo vo{ ln g`uuok qho`{ OV[v qd{h

    {ho hdahov{ ulvvdmbo boja{h* Qd{h {hdv bo~ob ln }ovlb{dlj" d{ dv jl

    bljao} ulvvdmbo {l vubd{ qho`{ fh}lglvlgov dj{l ` glv`df ln

    }dfol}{hlblalv }oadljv (Vl}}obbvo{ `b*" =5569Vl}}obbv" =55;$" `jk

    {ho flbdjo`}d{t mo{qooj {ho {ql aojlgov dv gl}o n}`agoj{ok

    {h`{ u}o~dlvbt `vvgok mt flgu`}`{d~o g`uudja* Ho}o" {ho

    ox{oj{ ln ko{of{`mbo }o`}}`jaogoj{v dv fblvo} {l {h`{ lmvo}~ok

    {h}lah flgu }dvljv mo{qooj qho`{ M@F vopojfov `jk

    l}{hlblalv }dfo aojlgo vopojfov" dj qhdfh g`jt oxfou{dljv

    {l gdf}lflbdjo`}d{t h`~o mooj lmvo}~ok (}o~doqok dj V`bvo `jk

    Nodbbo{" =551$*

    Mt flgmdjdja k`{` n}lg {ho dj{}`vuofdfff `jk dj{o}vuofdfff

    flgu`}dvljv" qo dkoj{dffok vo~oj kubdf`{dlj o~oj{v {h`{ `}o

    fljvo}~ok `{ l}{hlblalv ulvd{dljv mo{qooj }dfo `jk qho`{* Dj

    {hdv v{kt" qo vok ` flgmdj`{dlj ln djkd~dk`b `j`btvdv ln kubd'f`{dljv qd{hdj o`fh aojlgo `jk flbdjo`}d{t `vvovvgoj{ mo'

    {qooj {ho {ql aojlgov vdja dkoj{df`b `bdajgoj{ f}d{o}d` `jk

    v{`{dv{df`b ~`bdk`{dlj* U}o~dlv `{{ogu{v {l dkoj{dnt vh`}ok kubd'

    f`{dljv mo{qooj qho`{ `jk }dfo qo}o ljbt m`vok lj {ho j`btvdv

    ln }oadljv {h`{ fl}}ovuljk {l kubdf`{ok }oadljv dj }dfo vhlqdja

    ` hdah u}lul}{dlj ln aojo g`{fhov qd{h {ql kdnno}oj{ qho`{

    fh}lglvlgo a}luv* [hdv k`b vtj{ojt `uu}l`fh vok }ofoj{bt

    mt Vdjah o{ `b* (=551$ q`v m`vok lj }dfo aojov u}ovoj{ dj 8 ln {ho

    05 }oadljv u}o~dlvbt ijlqj {l mo kubdf`{ok (T o{ `b*" =55?$

    `jk {ho dkoj{dfff`{dlj ln hlglblalv qho`{ OV[v {h`{ g`u dj

    kobo{dlj mdjv lj {ql kdv{djf{ qho`{ fh}lglvlgov* _vdja {hdv

    `uu}l`fh lj {ho 06 l}{hlblalv }oadljv {h`{ qo}o dkoj{dffok dj

    l} v{kt" qo nljk {h`{ }0 dv l}{hlblalv {l q6 (=51 fljvo}~ok

    aojov$ `jk {l q0 (06 fljvo}~ok aojov$ m{ `bvl {l q= qd{h {hov`go vdajdfff`jfo (00 fljvo}~ok aojov$* Vdgdb`}bt" }? dv l}{hlb'

    lalv {l q0 (05= fljvo}~ok aojov$ `jk {l q6 (4 fljvo}~ok

    aojov$ m{ `bvl {l q= (8 fljvo}~ok aojov$ (k`{` jl{ vhlqj$*

    [hv" dj flj{}`v{ qd{h l} v{}`{oat" d{ q`v jl{ ulvvdmbo qd{h {hdv

    `uu}l`fh {l ~`bdk`{o v{`{dv{df`bbt {h`{ {ho }0'}? kubdf`{dlj dj }dfo

    dv vh }ok mo{qooj q6'q0 dj qho`{* Vdjah o{ `b* (=551$ `bvl

    fljfbkok {h`{ {ho }oadlj kubdf`{ok mo{qooj }00 `jk }0= dv

    fljvo}~ok dj qho`{ mo{qooj q;'q?" vaaov{dja {h`{ {ho kubd'

    f`{dlj u}ok`{ok {ho }dfo `jk qho`{ kd~o}aojfo* [hdv fljdf{v

    qd{h l} ffjkdjav `v qobb `v qd{h u}o~dlv v{kdov djkdf`{dja {h`{

    {ho}00'}0= kubdf`{dlj dv vuofdfff {l {ho}dfo bdjo`ao `jk}ob {d~obt

    }ofoj{ (T o{ `b*" =55?$* [hovo ffjkdjav u}l~dko ` jl{o ln f`{dlj

    nl} flgu`}`{d~o aojlgo'qdko `j`btvov `jk vaaov{ {h`{ `uubt'

    dja {ho v`go v{}djaoj{ `bdajgoj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk {dlj

    {l o`fh ln {ho k`{` vo{v vok nl} aojlgo flgu`}dvljv dv ov'

    voj{d`b {l `~ldk gdvdj{o}u}o{`{dlj ln o~lb{dlj`}t u`{{o}jv* Ho}o"

    mt m`vdja l} dj{}`aojlgdf `jk dj{o}aojlgdf flgu`}dvljv lj

    {ho hdahov{ dkoj{d{t l~o} {ho bljaov{ vopojfo `bdajgoj{" qo

    vok ` fljvo}~`{d~o `uu}l fh {h`{ oxfbkok vopojfov {h`{

    flbk bo`k ul{oj{d`bbt {l `j l~o}ov{dg`{dlj ln vopojfo fljvo}'

    ~`{dlj qd{hdj `jk mo{qooj {ho }dfo `jk qho`{ aojlgov* Jo~o}'

    {hobovv" qo qo}o `mbo {l ko{of{ `kkd{dlj b kubdf`{dljvdj }dfo `jk

    u}l~dko ` ko{`dbok `vvovvgoj{ ln aojlgo kubdf`{dljv dj qho`{"

    kogljv{} {dja {ho }lmv{jovv ln {hdv `uu}l`fh* Ndj bbt" {ho

    vo~oj kubdf`{dljv {h`{ `}o vh`}ok dj }dfo `jk qho`{ `}o bdiobt

    jko}ov{dg`{ok" vdjfo {ho k`{` vo{ ln qho`{ aojov nl} qhdfh `

    fh}lglvlg`b ulvd{dlj dv ijlqj dv f}}oj{bt bdgd{ok* N{}o b`}ao

    OV[ g`uudja l} aojlgo vopojfdja u}lcof{v qdbb mo }opd}ok {l

    lm{`dj ` flgubo{o `vvovvgoj{ ln {ho kubdf`{dljv dj {ho qho`{

    aojlgo `jk ln {hod} fljvo}~`{dlj qd{h {ho }dfo aojlgo*

    O~lb{dlj ln {ho A}`vv Aojlgov n}lg `jj? ?

    Flgglj @jfov{l}

    Djkd~dk`b aojlgo kubdf`{dlj `j`btvov dj }dfo" vl}ahg" `jk

    g`dso h`~o `b}o`kt vaaov{ok `j `jfov{}`b QAK u}ok`{dja {ho

    kd~o}aojfo mo{qooj {ho kdnno}oj{ fo}o`b aojlgov (U`{o}vlj

    o{ `b*" =55;9 T o{ `b*" =55?9 Q`ja o{ `b*" =55?$" m{ {l k`{o {ho

    =5 [ho Ub j{ Fobb

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    11/15

    l}dadj`b m`vdf jgmo} ln fh}lglvlgov }og`djok jfo}{`dj

    (A`{" =55=$* ]ofoj{bt" Qod o{ `b* (=551$ {hl}lahbt v{kdok {ho

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    o~lb{dlj ln {ho fo}o`b aojlgov n}lg `j `jfov{l} qd{h j 0=

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    glkob dv dj flgubo{o `a}oogoj{ qd{h {ho u}lulvok o~lb{dlj`}tu`{h n}lg {hdv `jfov{l} m{ u}l~dkov `kkd{dlj`b o~dkojfo nl} {ho

    l}dadj ln {ho 0= `jfov{}`b fh}lglvlgov*

    Mt dj{oa} {dja joq k`{` lj {ho kubdf`{dljv ln {ho qho`{

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    vh`}ok kubdf`{dljv mo{qooj }dfo" qho`{" g`dso" `jk vl}ahg"

    qo qo}o `mbo {l u}lulvo ` m`vdf fh}lglvlgo jgmo} ln j ?

    nl} {ho flgglj `jfov{l} ln {ho fo}o`b aojlgov* Qo vaaov{

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    {l 15 GT@" {ho `jfov{l} qd{h j ? fh}lglvlgov jko}qoj{

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    mo{qooj {ho kdnno}oj{ a}`vv aojlgov* Dj l} glkob" }dfo qlbk

    h`~o }o{`djok {hdv l}dadj`b fh}lglvlgo jgmo}" qho}o`v d{

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    {}`jvblf`{ok `jk dj~o}{ok voagoj{v ln `jfov{}`b fh}lglvlgov

    (nl} }o~doq" voo Vfhmo}{" =551$* [ho ff~o nvdljv vaaov{ok mt

    l} glkob dj qho`{ bdiobtlff}}okdj {ho`jfov{} b aojlgoln {ho

    [}d{dfo`o (qho`{" m`}bot" `jk }to YVof`bo fo}o`boW$ {h`{ h`~o `

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    {}`fov ln {ho u}ovojfo ln `jfdoj{ kubdf`{dljv q`v }ofoj{bt }o'

    ul}{ok dj ` m`}bot,}dfo flbdjo`}d{t v{kt (V{odj o{ `b*" =551$ vdja

    {ho k`b vtj{ojt `uu}l`fh ogubltok mt Vdjah o{ `b* (=551$*

    Flgu`}ok qd{h {ho jgo}lv }o`}}`jaogoj{v {h`{ }ovb{ok dj

    ` k}`g`{df }okf{dlj ln {ho g`dso fh}lglvlgo jgmo} (n}lg =5

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    vaaov{v {h`{ ljbt {ql fh}lglvlg`b nvdljv h`~o lff}}ok

    vdjfo {o{}`ubldkds`{dlj ln {ho `jfov{l} qd{h j ? fh}lglvlgov*

    [hdv blq bo~ob ln }o`}}`jaogoj{v dv vdgdb`} {l qh`{ dv lmvo}~ok dj

    qho`{" dj qhdfh o~oj `n{o} vo~o}`b }ljkv ln ulbtubldkds {dlj

    {ho}o dv ~o}t bd{{bo kdv}u{dlj ln flbdjo`}d{t mo{qooj {ho qdbk

    kdubldk `jfov{l}v`jk {ho ulbtubldkqho`{ aojlgov* [ho}onl}o" d{

    dv ulvvdmbo {h`{ {ho `jfov{}`b a}`vv aojlgo flj{`djok ` aojo

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    qhlbo aojlgo vhl{aj vopojfo qdbb mo flgubo{ok qd{hdj {ho

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    Qho`{`jk ]dfo Vopojfo K`{`m`vov

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    `vvdajok {l kobo{dlj mdjv mt Pd o{ `b* (=55;$ qo}o klqjbl`kok n}lg {hoA}`djAojo qomvd{o (h{{u3,,qho`{*uq*vk`*al~,$* Vdjfo vlgo OV[v g`t

    fl}}ovuljk {l jljl~o}b`uudja 64 `jk ?4 ojkv ln {ho v`go fKJ@ vo'

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    f`{dlj ln kubdf`{dljv qd{hdj {ho qho`{ aojlgo `jk nl} ov{`mbdvhdja

    O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv =0

  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

    12/15

    l}{hlblalv }ob`{dljvhduv mo{qooj }dfo (L}ts` v`{d~ $ `jk qho`{* Vdjfo

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    qho`{ hlglolblalv fh}lglvlgo a}luv* [hv" `jt QOF `jk QOV{h`{ q`v g`uuok dj ` kobo{dlj mdj lj {ql l} {h}oo ln {ho hlglolblalv

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    jdpo ulvd{dlj lj ` vdjabo fljvojvv fh}lglvlgo uo} a}lu* [ho

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    JA]$$,FVd W 6 055" qho}o MV{ mdj v{`}{ fll}kdj`{o" Mvd mdj vdso (dj

    uo}foj{ ao ln {ho}og`djdja fh}lglvlgo YPdo{ `b*" =55;W$" JA jgmo}

    ln aojov `vvdajok {l ` mdj" JA] aojo }`ji qd{hdj ` mdj" `jk FVd {l{`b

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    lalv fh}lglvlgo `}gv qo}o dajl}ok*

    [ho vopojfov ln {ho 0= }dfouvoklglbofbov(mdbk ;9 61="511"850 mu$

    qo}o klqjbl kok n}lg {ho [DA] qomvd{o (n{u3,,n{u*{da}*l}a,um,k`{ ,

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    `jk {ho ;=">?; aojov dkoj{dffok mt {ho `jjl{`{dlj qo}o vok nl} {ho

    `j`btvdv ln {ho }dfo kubdf`{dljv* Qo fhlvo {l uo}nl}g {ho `j`btvdv qd{h{ho [DA] `jjl{ {dlj }`{ho} {h j {ho gl}o fljvo}~`{d~o `jjl{ {dlj

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    U}lcof{ (=551$ {l djf}o`vo {ho u}lm`mdbd{t ln ffjkdja fljvo}~ok l}{hlblav

    o~oj `glja vopojfov {h`{ qo}o jl bljao} fljvdko}ok dj {ho joq

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    Dkoj{dfff`{dlj ln Kubdf`{ok ]oadljv dj Qho`{ `jk ]dfo

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    vuljkdja {l OV[v {h`{ qo}o g`uuok lj {ql kdnno}oj{ hlglolblalv

    fljvojvv fh}lglvlgov nl} o`fh ln {ho vo~oj fh}lglvlgo a}luv* Dj

    }dfo" kubdf`{dljv qo}o dkoj{dffok {h}lah aojlgo vopojfo flgu`}dvlj

    vdja {ho ;=">?; }dfo `jjl{ {ok aojov* @n{o} `bdajgoj{ ln {ho }dfo

    `jjl{ {ok aojov vdja 15/ FDU `jk 15/ F@BU ~`bov" {ho aojov

    vhlqdja ` g`{fh qd{h `jl{ho} aojo obvoqho}o dj {ho aojlgo qo}o

    koffjok `v u{`{d~o u`}`blalv u`d}v {h`{ qo}o vok nl} {ho dkoj{dfff`{dlj

    ln kubdf`{ok }oadljv*

    Dkoj{dfff`{dlj ln L}{hlblalv ]oadljv mo{qooj ]dfo `jk Qho {

    Jlj}okjk`j{ QOFv `jk QOVv qo}o `bdajok `a`djv{ {ho }dfo `jjl{`{ok

    aojov qd{h >5/ FDU `jk 15/ F@BU {l dkoj{dnt u{`{d~o l}{hlblalv

    vopojfov* @j dj'kou{h `j`btvdv ln {ho `bdajgoj{v vhlqok {h`{ {ho

    u}ovojfo ln jfljvo}~ok ?4 l} 64 j{}`jvb`{ok }oadljv (_[]v$ fljvdko}'

    `mbt kof}o`vok {ho l~o}`bb uo}foj{`ao ln dkoj{d{t `jk kdv{}mok {ho

    dkoj{dfff`{dlj ln l}{hlblalv ~o}vv u`}`blalv aojov* [l {`io {hdv

    u}lmbog dj{l `fflj{" qo ov{dg {ok {ho `~o}`ao boja{h ln {ho ?4

    `jk 64

    _[]v dj l} k`{` vo{* [ho `~o}`ao boja{hv qo}o =?5 > >?*> `jk ;55 >

    00=*? mu ln {ho ?4 `jk 64 _[]v" }ovuof{d~obt" nl} 8?/ ln {ho QOFv l}

    QOVv fljvdko}ok dj l} v{kt* [hv" dj `kkd{dlj {l vopojfov vhlqdja

    hdah vdgdb`}d{t (FDU$ l~o} {ho qhlbo boja{h (F@BU$" qo fljvdko}ok `v

    ul{oj{d`b l}{hlblav l} u`}`blav `bb aojov nl} qhdfh u`d}qdvo vopojfo

    `bdajgoj{v vhlqok ` gdjdgg ln >5/ FDU `jk 15/ F@BU `n{o} vtv'

    {og`{df`bbt oxfbkdja {ho _[] vopojfov*

    @ aojo (A$q`v fljvdko}ok}o`}} jaok mo{qooj }dfo `jkqho`{(d*o*" jl{

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    V{`{dv{df`b @j`btvdv

    U{ {d~o u`} blalv u`d}v dkoj{dffok k}dja{ho kubdf {dlj `j btvov dj {ho

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    mblfiv ln aojov {h { `}ojl{ lm{ djok mt fh`jfo(d*o*" {h { `}ojl{ lm{ djok

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

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  • 8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass

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