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Transcript of Identification and Characterization of Shared Duplications between Rice and Wheat Provide New...
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
1/15
]OVO@]FH @][DFBOV
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Co}lgo V bvo"` V{ouh`jdo Mlbl{"` Gdfh`ob [h}lko"` ^djfoj{ Clnno"` Mojl{ Udoa"m _g } G`vllk P} dvhd"`
[hlg`v F`bf`ajl"` ]dfh }k Fllio"m Gdfhob Kobvojt"m `jk F`{ho}djo Nodbbo{`"0
` Djv{d{{ J`{dlj`b ko b` ]ofho}fho @a}ljlgdpo,_jd~o}vd{o Mb`dvo U`vf`b _jd{o Gdx{o ko ]ofho}fho 054?" @gobdl}`{dlj o{ V`j{o
kov Ub`j{ov" >6055 Fbo}glj{'No}}`jk" N}`jfom _jd{o Gdx{o ko ]ofho}fho ?54>" Foj{}o J`{dlj`b ko b` ]ofho}fho Vfdoj{dffpo,_jd~o}vd{o ko Uo}udaj`j,Djv{d{{ ko ]ofho}fho
ul} bo Ko~obluuogoj{" B`ml}`{ld}o Aojlgo o{ Ko~obluuogoj{ kov Ub`j{ov ?=" >>8>5 Uo}udaj`j Fokox" N}`jfo
[ho a}`vv n`gdbt flgu}dvov {ho glv{ dgul}{`j{ fo}o`b f}luv `jk dv ` allk vtv{og nl} v{ktdja" qd{h flgu`}`{d~o aojlgdfv"
gofh jdvgv ln o~lb{dlj" vuofd {dlj" `jk klgov{df {dlj* Ho}o" qo dkoj{dffok `jk fh`}`f{o}dsok {ho o~lb{dlj ln vh`}okkubdf`{dljv dj {ho }dfo (L}ts` v`{d~ $ `jk qho`{ ([}d{dfug ov{d~ug$ aojlgov mt flgu`}dja ;=">?; }dfo aojo vopojfov qd{h
>;=> g`uuok qho`{ OV[v vdja dgu}l~ok vopojfo `bdajgoj{ f}d{o}d` `jk v{`{dv{df`b `j`btvdv* Dj{}`vuofdfff flgu`}dvljv
dkoj{dffok =4 dj{o}fh}lglvlg b kubdf {dljv fl~o}dja 1=/ ln {ho }dfo aojlgo `jk 05 kubdf {dlj mblfiv fl~o}dja >1*?/ ln {ho
qho { aojlgo* _vdja {ho v`go go{hlklblat" qo `vvovvok l}{hlblalv }ob {dljvhduv mo{qooj {ho {ql aojlgov `jk ko{of{ok
06 mblfiv ln flbdjo`}d{t{h`{ }ou}ovoj{86*0 `jk 45*;/ ln {ho}dfo `jkqho`{ aojlgov"}ovuof{d~obt* Dj{oa}`{dljln {hodj{}`vuofdfff
kubdf`{dljv k`{` qd{h flbdjo`}d{t }ob`{dljvhduv }o~o`bok vo~oj kubdf`{ok voagoj{v fljvo}~ok `{ l}{hlblalv ulvd{dljv* @
ko{ dbok`j`btvdv ln {hoboja{h" flgulvd{dlj"`jk kd~o}aojfo {dgo ln {hovo kubdf`{dljv `jkflgu`}dvljv qd{hvl}ahg (Vl}ahug
mdflbl}$ `jk g dso (So` g`tv$ djkdf`{ok flgglj `jk bdjo`ao'vuofdfff u`{{o}jv ln fljvo}~`{dlj mo{qooj {ho kdnno}oj{ aojlgov*
[hdv `bblqok v {l u}lulvo ` glkob dj qhdfh {ho a} vv aojlgov h`~o o~lb~ok n}lg ` flgglj `jfov{l} qd{h ` m`vdf jgmo} ln
ff~o fh}lglvlgov {h}lah ` vo}dov ln qhlbo aojlgo `jk voagoj{`b kubdf`{dljv" fh}lglvlgo nvdljv" `jk {}`jvblf`{dljv*
DJ[]LK_F[DLJ
[ho a}`vv n`gdbt dv {ho nl}{h b`}aov{ `glja lqo}dja ub`j{v `jk
flgu}dvov vlgo ln {ho `a}ljlgdf`bbt glv{ dgul}{`j{ f}lu vuo'
fdov" vfh `v qho`{ ([}d{dfg vvu$" g`dso (So` g`tv$" `jk }dfo
(L}ts` v`{d~ $* A}`vv aojlgov kdnno} a}o`{bt dj vdso" ubldkt bo~ob"
`jkfh}lglvlgo jgmo}* M}o`k qho`{ ([}d{dfg `ov{d~g9 =j
;=$ mobljav {l {ho Ulldko`o n`gdbt `jk h`v ` hox`ubldk aojlgo
(@@MMKK$ ln 01 Am {h`{ l}dadj`{ok {h}lah {ql ulbtubldkds`{dlj
o~oj{v (Nobkg`j o{ `b*" 044?9 Mb`io o{ `b*" 04449 H`ja o{ `b*"
=55=$* ]dfo (=j =;$ dv kdubldk `jk mobljav {l {ho Oh}h`}{ldko`o
n`gdbt* Qd{h ` vdso ln 5*; Am" d{v aojlgo dv ;5 {dgov vg`bbo} {h`j
{h`{ ln m}o`k qho`{* Nlvvdb k`{` `jk uhtblaojo{df v{kdov ov{d'
g`{ok {h`{ {ho kdnno}oj{ a}`vv n`gdbdov kd~o}aok n}lg ` flgglj
`jfov{l} ?5 {l 15 gdbbdlj to }v `al (GT@$ (nl} }o~doqv" voo
Iobblaa" =5509 A`{" =55=$*Flgu`}`{d~o v{kdov mo{qooj a}`vvov" glv{bt fo}o`bv vfh
`v m`}bot (Hl}kog ~ba`}o$" qho`{" g`dso" }dfo" `jk vl}ahg
(Vl}ahg mdflbl}$" h`~o mooj {honlfvln dj{ojvo }ovo }fh dj {ho
u`v{ kof`ko (nl} ` }ofoj{ }o~doq" voo V`bvo `jk Nodbbo{" =551$*
O`}bt flgu`}`{d~o v{kdov }obdok lj f}lvv}ov{}df{dlj n}`agoj{
boja{h ulbtgl}uhdvg (]NBU$ g`uudja `j`btvov ln fblvobt }ob`{ok
vuofdov* [hot }o~o`bok vdajdfff`j{ g`f}lflbdjo`}d{t mo{qooj {ho
fo}o b aojlgov `jkbok {l {hofljv{}f{dlj ln ` fljvojvv a}`vv
g`u m`vok lj =? }dfo bdji`ao mblfiv (}o~doqok dj Ko~lv `jk
A`bo" =5559 Nodbbo{ `jk Iobbo}" =55=9 Ko~lv" =55?$* [hovo }ovb{v"
hlqo~o}" qo}o lm{ djok n}lg blq'}ovlb{dlj aojo{df g`uv qd{h `j
`~o} ao ln ljo g`}io} o~o}t 05 foj{dgl}a`j {h`{ `bblqok {ho
ko{of{dlj ln ljbt b`}ao }o`}}`jaogoj{v* Gl}ol~o}" {ho g`uv qo}o
fljv{}f{ok qd{h blq'flut ]NBU g`}io}v {h`{ qo}o vobof{ok nl}
{hod} `mdbd{t {l u}l~dko ` vdaj b dj f}lvv'htm}dkds {dljv" {ho}omt
bdgd{dja {ho ko{of{dlj ln qhlbo l} u`}{d b aojlgo kubdf {dlj
o~oj{v* D{ `bvl h`v mooj kdnfffb{ {l `vvovv l}{hlblalv `jk u`}`b'
lalv }ob`{dljvhduv dj aojo n`gdbdov" vdjfo flgu`}`{d~o g`uudja
mt ]NBU ln{oj dkoj{dffok u`}`b`alv }`{ho} {h`j l}{hlblalv vo'
pojfov" bo`kdja {l `j jko}ov{dg`{dlj ln flbdjo`}d{t*
Dj {ho u`v{ ? to`}v" dj{o}j`{dlj`b djd{d {d~ov h`~o bok {l {ho
ko~oblugoj{ ln `kkd{dlj`b aojlgdf }ovl}fov {h`{ `bblq flg'
u`} {d~o aojlgdf v{kdov mo{qooj {hoa}`vv aojlgov `{ ` hdaho}
bo~ob ln }ovlb{dlj (gdf}lflbdjo`}d{t$* [ho Dj{o}j`{dlj`b [}d{dfo`o
OV[ Flluo}`{d~o (h{{u3,,qho`{*uq*vk`*al~,aojlgo,$ onnl}{v
h`~o }ovb{ok dj {ho u}lkf{dlj ln :0 gdbbdlj qho`{ OV[v (h{{u3,,
qqq*jfmd*jbg*jdh*al~,kmOV[,kmOV[\vgg`}t*h{gb" 54,=8,51
}obo`vo$" 1051 ln qhdfh" koffjdja 0>"544 blfd" h`~o mooj ft{lao'
jo{df`bbt g`uuok dj kobo{dlj mdjv (Pd o{ `b*" =55;$* Dj }dfo" {ho
0 @kk}ovv fl}}ovuljkojfo {l f`{ho}djo*nodbbo{Efbo}glj{*dj}`*n}*
[ho `{hl} }ovuljvdmbo nl} kdv{}dm{dlj ln g`{o}d bv dj{oa} b {l {ho
ffjkdjav u}ovoj{ok dj {hdv `}{dfbo dj `ffl}k`jfo qd{h {ho ulbdft kovf}dmok
dj {ho Djv{}f{dljv nl} @{hl}v (qqq*ub`j{fobb*l}a$ dv3 F`{ho}djo Nodbbo{
(f`{ho}djo*nodbbo{Efbo}glj{*dj}`*n}$*QLjbdjo ~o}vdlj flj{`djv Qom'ljbt k`{`*
qqq*ub`j{fobb*l}a,fad,kld,05*005?,{uf*051*5?>654
[ho Ub`j{ Fobb" ^lb* =53 00=;" C`j`}t =558" qqq*ub`j{fobb*l}a =558 @go}df`j Vlfdo{t ln Ub`j{ Mdlbladv{v
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
2/15
Dj{o}j`{dlj`b ]dfo Aojlgo Vopojfdja U}lcof{ (D]AVU$ }ofoj{bt
flgubo{ok {ho vopojfo ln {ho L* v`{d~` vvu c`uljdf` f~
Jduuljm }o (Dj{o}j`{dlj`b ]dfo Aojlgo Vopojfdja U}lcof{"
=55?$* [qob~o uvoklglbofbov fl}}ovuljkdja {l 61="511"850
mu ln ffjdvhok vopojfo ([ho Djv{d{{o nl} Aojlgo ]ovo`}fh
Y[DA]W ~o}vdlj ;9 680"0?5"4;? mu dj D]AVU ~o}vdlj ;$ qo}o `v'vogmbok `jk fh`}`f{o}dsok {h}lah vo~o}`b }ljkv ln `jjl{`'
{dlj* [hdv }ovb{ok dj ov{dg`{ov nl} {ho }dfo aojo jgmo} }`jadja
n}lg 96="555 (]dfo @jjl{`{dlj U}lcof{" =551$ nl} {ho glv{ }o'
foj{ `jjl{`{dlj ln {ho dj{o}j`{dlj`b fljvl}{dg {l ;=">?; aojov
dj {ho [DA] ~o}vdlj ; (h{{u3,,qqq*{da}*l}a,{da}'vf}du{v,lv`0\qom,
am}lqvo,}dfo,9 T`j o{ `b*" =556$* [hovo }ovl}fov h`~o mooj
vok {l uo}nl}g b`}ao'vf`bo dj{}`vuofdfff `jk dj{o}vuofdfff vo'
pojfo flgu }dvljv mo{qooj {ho {ql aojlgov `jk h`~o hobuok
{l }offjo l} jko}v{`jkdja ln flbdjo`}d{t mo{qooj {hod} fh}lgl'
vlgov* Vl}}obbv o{ `b* (=556$" Vl}}obbv (=55;$" `jk Vdjah o{ `b*
(=551$ h`~o flgu`}ok {ho vopojfov ln ;;8? `jk 614= ft{lao'
jo{df`bbt g`uuok qho { OV[v`a`djv{{ho }dfoaojlgo vopojfo*
V{kdov nlfvdja lj vdjabo fh}lglvlgo a}luv l} }oadljv h`~o
mooj uo}nl}gok }ofoj{bt `v qobb nl} }dfo fh}lglvlgo 6 flg'u`}ok qd{h qho { `jk g`dso OV[v (Mobb o{ `b*" =55?9 ]dfo
Fh}lglvlgo 6 Vopojfdja Fljvl}{dg" =55?$ `jk nl} }dfo fh}l'
glvlgo 00 flgu`}ok qd{h qho`{ OV[v (Vdjah o{ `b*" =55;$*
[hovo v{kdov djf}o`vok {ho }ovlb{dlj ln flgu`}`{d~o g`uudja
mo{qooj {ho {ql vuofdov mt =?' {l 65'nlbk" }o~o`bdja gl}o }o'
`}}`jaogoj{v {h`j u}o~dlvbt lmvo}~ok `{ {ho aojo{df g`u bo~ob*
Dj `kkd{dlj {l {ho `vvovvgoj{ ln flbdjo`}d{t mo{qooj {ho ao'
jlgov" flgu`}`{d~o `j`btvov f`j }o~o`b `jfov{}`b aojlgo k'
ubdf`{dljv* O`}bt v{kdov qd{h {ho ff}v{ aojo}`{dlj ln glbofb`}
g`}io}v djkdf`{ok {ho u}ovojfo ln kubdf`{ok blfd lj {ho aojo{df
g`uv dj kdnno}oj{ fo}o`bv" vaaov{dja `jfov{}`b aojlgo kubd'
f`{dljv `jk ulbtubldkds`{dlj o~oj{v dj {ho hdv{l}t ln vuofdov {h`{
`}o jlq fljvdko}ok kdubldkv* ]NBU `jk dvlstgo v{kdov dj {ho
o`}bt 0445v h`k `b}o`kt vaaov{ok {h`{ g`dso fh}lglvlgov
vh`}okubdf`{ok voagoj{v (Qojkob o{ `b*" 04849@hj `jk [`jivbot"
0446$* Qhlbo kubdf`{dlj ln {ho g`dso aojlgo {h}lah `bbl{o'
{}`ubldkds`{dlj q`v dkoj{dffok `jk fh`}`f{o}dsok n}{ho} {h}lah
{ho o~lb{dlj`}t `j`btvdv ln kubdf`{ok aojov (A`{ `jk Klombot"
0441$ `jk mt dj{o}vuofdfff flgu`}dvljv mo{qooj l}{hlblalv
blfd dj }dfo" vl}ahg" `jk g`dso (Vqdaljl~` o{ `b*" =55;$* Dj }dfo"
o }bt]NBU g`uudja v{kdov vaaov{ok{h`{ fh}lglvlgov 0 `jk
? (Idvhdgl{l o{ `b*" 044;$ `v qobb `v fh}lglvlgov 00 `jk 0=
(J`a`g}` o{ `b*" 044?$ flj{`dj `jfdoj{ kubdf`{ok }oadljv* [ho
}obo`vo ln {ho aojlgo vopojfo k}`n{v n}lg c`uljdf` `jk djkdf`
}dfo vmvuofdov `bblqok qhlbo aojlgo vopojfo flgu`}dvljv
`jk n}{ho} fh`}`f{o}ds {dlj ln kubdf`{dljv dj }dfo (T o{ `b*"
=55=" =55?9 U`{o}vlj o{ `b*" =556" =55;9 ^`jkoulobo o{ `b*" =5569
Atl{ o{ `b*" =55;9 Dj{o}j`{dlj b ]dfo Aojlgo Vopojfdja
U}lcof{" =55?9 Q`ja o{ `b*" =55?$* [ho glv{ }ofoj{bt umbdvhok
`j`btvdv (T o{ `b*" =55?$ }o~o`bok ` qhlbo aojlgo kubdf`{dlj
(QAK$ {h`{ lff}}ok mo{qooj ?6 `jk 4; GT@ (d*o*" monl}o {ho
kd~o}aojfo ln {ho fo}o`b aojlgov$" ` }ofoj{ voagoj{`b kubdf`'
{dlj mo{qooj fh}lglvlgov 00 `jk 0=" `jk jgo}lv djkd~dk`b
aojo kubdf`{dljv* [lao{ho}" {hovo kubdf {dljv fl~o} `j ov{d'
g`{ok >?*1/ln {ho }dfoaojlgo* ]dfoaojlgo kubdf`{dljv h`~o
mooj v{kdok `bvl mt [DA] vdja d{v b`{ov{ aojlgo `jjl{`{dlj
(;0"5;> jlj{} jvulv`mbo obogoj{}ob {ok }dfo u}l{odj vopojfov9
h{{u3,,qqq*{da}*l}a,{km,o=i0,lv`0,voagoj{`b\ku,djkox*vh{gb$*
[ho }ovb{vdjkdf`{o` b`}ao jgmo} ln voagoj{ b kubdf {dljv" m{
{ho blq v{}djaojft ln {ho `j`btvdv kdk jl{ `bblq fbo`} fljfbvdljv
{l mo k}`qj lj {ho ox`f{ j`{}o ln {ho }dfo kubdf`{dljv* Ndj`bbt"
flgu`} {d~o `j`btvov mo{qooj }dfo `jk l{ho} a}`vvov h`~o mooj
hobunb dj }o~o bdja kubdf`{dljv dj {ho }dfo aojlgo* Nl} ox`gubo"
mt flgu`}dja =>55 g`dso g`uuok vopojfo g`}io}v qd{h {ho}dfo vopojfo" V`bvo o{ `b* (=55;$ dkoj{dffok vdx kubdf`{dljv
mo{qooj fh}lglvlgov 8'0=" ='>" >'05" 0'?" `jk 6'> {h`{ h`k
jl{ mooj ko{of{ok u}o~dlvbt*Gl}o }ofoj{bt" Qod o{ `b* (=551$ mdb{
` hdah'}ovlb{dlj flgu`} {d~ouhtvdf b g`u mo{qooj {ho }dfo`jk
g`dso aojlgov {h { hobuok{l }offjo g`dso `jk }dfo kubdf {dljv*
[hv" ` jgmo} ln v{kdov h`~o vaaov{ok {h`{ {ho a}`vv ao'
jlgov qo}o vmcof{ok {l kdnno}oj{ }ljkv ln qhlbo aojlgo `jk
voagoj{ b kubdf {dljv k}dja {hod} o~lb{dlj n}lg ` flgglj
`jfov{l} ?5 {l 15 GT@* Qhdbo {ho gofh`jdvgv mt qhdfh {ho
aojlgov h`~o o~lb~ok h`~o moflgo gl}o o~dkoj{ l~o} {ho u`v{
kof`kov" {ho m`vdf fh}lglvlgo jgmo} ln {ho a}`vv `jfov{l}
`jk {ho o~lb{dlj`}t u`{h {h`{ bok {l {ho b`}ao ~`}do{t ln m`vdf
fh}lglvlgo jgmo} nljk dj {lk`tv a}`vv aojlgov }og`dj
jfo}{`dj* Kdnno}oj{ `{hl}v h`~o u}lulvok m`vdf jgmo}v }`ja'dja n}lg ? {l 0= fh}lglvlgov (nl} ` }o~doq" voo A`{" =55=9 Qod
o{ `b*" =551$" m{ {l k`{o jl glkob {h`{ qlbk }ofljfdbo `bb ln {ho
k`{` h`v mooj u}lulvok nl} {ho v{}f{}o `jk o~lb{dlj ln {ho
`jfov{}`b a}`vv aojlgo*
Mof vo d{ dv kdnfffb{ {l djno} l}{hlblalv (ko}d~ok n}lg `
flgglj `jfov{l} mt vuofd`{dlj$ `jk u`} blalv (ko}d~ok mt
kubdf`{dlj qd{hdj ljo aojlgo$ }ob {dljvhduv n}lg vopojfo
flgu`}dvljv" v{}djaoj{ `bdajgoj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk`'
{dlj `}o ovvoj{d`b {l o~`b`{o `ff}`{obt qho{ho} {ho `vvlfd`{dlj
mo{qooj {ql l} gl}o aojov nljk dj {ho v`go l}ko} lj {ql
fh}lglvlg`b voagoj{v dj kdnno}oj{ aojlgov lff}v mt fh`jfo
l} }oof{v {}o flbdjo`}d{t* Vo~o}`b }ofoj{bt ko~obluok vln{q`}o
u}la}`gv" vfh `v Bdjo_U (H`guvlj o{ `b*" =556$" @KHl]O
(@{lg`{df Ko{of{dlj ln Hlglblalv ]oadljv9 ^`jkoulobo
o{ `b*" =55=$" NDVH (N`v{ Dkoj{dfff`{dlj ln Voagoj{`b Hlglblat9
F`b m}ovoo{ `b*" =556$"`jk Fblvo_u (H`guvlj o{ `b*" =55?$"hobu
{l `kk}ovv {hovo pov{dljv* @ jgmo} ln l{ho} u}la}`gv" vfh
`v Fg`u (N`ja o{ `b*" =556$" `jk qomvd{ov" vfh `v A}`gojo
(C`dvq`b o{ `b*" =55>$ l} {ho [DA] vtj{ojt u}lcof{v (h{{u3,,qqq*
{da}*l}a,{km,vtj{ojt,qho`{,kovf}du{dlj*vh{gb$" h`~o mooj ov{`m'
bdvhok {l ~dv bdso vopojfo'm`vok flbdjo`}d{t k`{` lm{ djok
n}lg flgu`}`{d~o vopojfo `j`btvov mo{qooj {ho a}`vv ao'
jlgov* Qhdbo {hovo qomvd{ov u}l~dko vo}'n}dojkbt a}`uhdf b
kdvub`tv ln g`f}lflbdjo }d{t" {hot }obt lj k`{ lm{ djok qd{h blq'
v{}djaojft `bdajgoj{ f}d{o}d` `jk qd{hl{ v{`{dv{df`b ~`bdk`{dlj* Dj
`kkd{dlj"{hot kl jl{{`iodj{l `fflj{ {hokojvd{t`jk blf {dlj ln
fljvo}~ok aojov {l dkoj{dnt u}ofdvobt u`}`blalv `jk l}{hlbl'
alv }oadljv9 {ho}onl}o" {hot aojo}`bbt l~o}ov{dg`{o flbdjo`}d{t
mo{qooj kdnno}oj{ voagoj{v ln {ho aojlgov*
Dj {hdv v{kt" qo `uubdok joq `jk v{}djaoj{ `bdajgoj{ f}d{o}d`
`jk uo}nl}gok v{`{dv{df`b {ov{v vtv{og`{df`bbt {l }okoffjo dj{o}'
fh}lglvlg`b kubdf`{dljv dj }dfo" dkoj{dnt qho`{ aojlgo kubd'
f`{dljv" `jk }o`vvovv flbdjo`}d{t }ob`{dljvhduv mo{qooj {ho {ql
aojlgov* [hdv `bblqok v {l ko{of{ `jk fh`}`f{o}dso vh`}ok
kubdf`{ok }oadljv mo{qooj }dfo `jk qho`{" {l flgu`}o {hog
qd{h g`dso `jk vl}ahg k`{`" `jk {l ov{`mbdvh ` glkob nl} {ho
o~lb{dlj ln {ho a}`vv aojlgov n}lg ` flgglj `jfov{l} qd{h
j ? fh}lglvlgov*
0= [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
3/15
]OV_B[V
Dgu}l~ok Vopojfo @bdajgoj{ F}d{o}d` {l Djno} Vdajdfff`j{
L}{hlblalv `jk U`}`blalv ]ob {dljvhduv
Qhoj {ql vopojfov `}o `bdajok" MB@V[J (@b{vfhb o{ `b*" 0445"
0441$ u}lkfov hdah'vfl}dja u`d}v (HVUv$ {h`{ fljvdv{ ln {ql
vopojfo n}`agoj{v ln `}md{}`}t m{ op`b boja{h" {ho `bdajgoj{
ln qhdfh dv blf`bbt g`xdg`b `jk nl} qhdfh {ho `bdajgoj{ vfl}o
goo{v l} oxfookv ` {h}ovhlbk l} f{lnn vfl}o*HVUv `}om`vok lj
v{`{dv{df`b f}d{o}d` vfh `v {ho o'~`bo" vfl}o" `jk uo}foj{`ao ln
dkoj{d{t* Ko{of{dja fljvo}~ok }oadljv dv bdgd{ok qd{h vopojfo
`bdajgoj{v lm{ djokmt MB@V[qd{h {hovokon`b{u`}`go{o}v* [l
djf}o`vo {ho vdajdfff`jfo ln dj{}`vuofdfff `jk dj{o}vuofdfff vo'
pojfo `bdajgoj{v nl} djno}}dja o~lb{dlj`}t }ob`{dljvhduv qd{hdj
`jk mo{qooj aojlgov" qo koffjok {h}oo joq u`}`go{o}v nl}
MB@V[ `j btvdv3@B nl}`bdajok boja{h" FDUnl} fgb {d~o dkoj{d{t
uo}foj{`ao" `jk F@BU nl} fgb`{d~o `bdajgoj{ boja{h uo}foj{'
`ao (voo Go{hlkv nl} koffjd{dljv$* Qd{h {hovou`}`go{o}v" MB@V[
u}lkfov {ho hdahov{ fgb`{d~o uo}foj{`ao ln dkoj{d{t l~o} {hobljaov{ fgb`{d~o boja{h" {ho}omt djf}o`vdja {ho v{}djaojft dj
koffjdja fljvo}~ {dlj mo{qooj vopojfov* Umbdfbt `~`db`mbo
}dfo `jk qho`{ OV[ vopojfov qo}o }o j`btsok mt MB@V[ vdja
{hovo u`}`go{o}v" nlbblqok mt ` v{`{dv{df`b {ov{ qd{h {ho Fblvo_u
vln{q`}o (H`guvlj o{ `b*" =55?$ {h`{ ~`bdk`{ov jlj}`jklg `v'
vlfd`{dljv mo{qooj a}luv ln vopojfov* Qo `uubdok kdnno}oj{
bo~obv ln v{}djaojfdov {l {ho dj{}`vuofdfff `jk dj{o}vuofdfff vo'
pojfo flgu`}dvljv dj {ho Fblvo_u `j`btvdv {l {`io dj{l `f'
flj{ kdnno}ojfov mo{qooj {ho jgmo}v ln vopojfov dj o`fh
k {` vo{* Vdjfo {ho qhlbo }dfo aojlgo dv `~ db`mbo (;=">?;
`jjl{`{ok aojov$" qo `uubdok hdaho} v{}djaojft `bdajgoj{ f}d{o'
}d` `jk v{`{dv{df`b ~`bdk`{dlj dj }dfo {h`j dj qho`{" nl} qhdfh ljbt
` bdgd{ok vmvo{ ln g`uuok aojo vopojfov (>;=> OV[v$ dv
`~`db`mbo* Mt flgmdjdja {hovo }ovb{v qd{h k`{` lj {ho fh}lgl'vlg`b blf`{dlj ln {ho vopojfov" qo qo}o `mbo {l kdv{djadvh
l}{hlblalv `jk u`}`blalv }oadljv dj ml{h aojlgov qd{h hdah
fljffkojfo `jk" vmvopoj{bt" {l dkoj{dnt vh`}ok kubdf`{dljv
mo{qooj {ho {ql aojlgov*
Dkoj{dfff`{dlj `jk Fh`} f{o}ds`{dlj ln Kubdf`{ok ]oadljvdj {ho ]dfo Aojlgo
Kubdf`{ok }oadljv dj }dfo qo}o dkoj{dffok mt `uubtdja {ho joq
`bdajgoj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk`{dlj kovf}dmok `ml~o {l
{ho ;=">?; jlj'{}`jvulv`mbo obogoj{'}ob`{ok aojov ko{of{ok dj
{ho nl}{h }obo`vo ln {ho [DA] }dfo aojlgo `jjl{`{dlj (h{{u3,,
qqq*{da}*l}a,{km,o=i0,lv 0,uvoklglbofbov,djnl*vh{gb$* [ho
aojo vopojfov qo}o `bdajok `a`djv{ {hogvob~ov (MB@V[J$
vdja 15/ FDU `jk 15/ F@BU* [ho }ovb{v vhlq {h`{ 6;"456"
=;05" 00=0" `jk ;==5 aojov g`{fhok so}l" ljo" {ql" `jk gl}o
{h`j {ql vopojfov" }ovuof{d~obt" obvoqho}odj {ho}dfo aojlgo*
Qo fljvdko}ok `v }obd`mbo djkdf`{l}v ln u{`{d~o kubdf`{dljv dj
{ho aojlgo {ho =;05 (?*>/$ vopojfov {h`{ dkoj{dffok ` vdjabo
hlglbla (qd{h `j `~o}`ao ~`bo ln 84*;/ FDU `jk 40*=/ F@BU$*[hot qo}o `j btsok n}{ho} qd{h Fblvo_u (kojvd{t }`{dl"=9 fbv{o}
boja{h" ;59 g`{fh" ?$ {l kdv{djadvh mo{qooj {ho }oadljv fl}}o'
vuljkdja {l aojov {h`{ }ovb{ n}lg `jfdoj{ kubdf`{dljv `jk
{hlvo {h`{ `}o `vvlfd`{ok }`jklgbt* @glja {ho =;05 u{`{d~o
kubdf`{ok blfd" ?64 (==*;/$ qo}o v{`{dv{df`bbt vdajdfff`j{* [hot
koffjok =4 kubdf`{ok }oadljv {h`{ `}o kdv{}dm{ok `glja {ho 0=
}dfo fh}lglvlgov `v nlbblqv3 }0'}=,6,?,05,0=" }='};,>,1,8,0="
}6'}1,4,05,00,0=" };'}?,8,05" }?'}4,00,0=" }>'}1,8,0="}1'}8" }8'}4,00"
}4'}00" `jk }00'}0= (Nda}o 0@9 vooVuubogoj{ b [`mbo 0 ljbdjo$*
[oj ln {ho =4 kubdf`{dljv (djkdf`{ok mt `v{o}dviv dj Vuubo'
goj{`b [`mbo 0 ljbdjo$ fl}}ovuljk {l {ho kubdf`{dljv dkoj{dffok
u}o~dlvbt mt T o{ `b* (=55?$ (dj }ok dj Nda}o 0@$ `jk fl~o}
;1*8/ ln {ho }dfo aojlgo* [ho 05 kubdf`{dljv fl}}ovuljk {l
ox`f{bt {ho v`go }oadljv `v {ho 08 kubdf`{ok }oadljv }oul}{okmt T o{ `b* (=55?$" m{ dj l} `j`btvdv qo h`~o fljvdko}ok `
Nda}o 0* Dj{}`vuofdfff Kubdf`{dljv ln {ho ]dfo `jk Qho`{ Aojlgov*
(@$ Vfhog {df }ou}ovoj{ {dlj ln {ho ?64u`d}v ln u`} blalv aojov (bdjiok mt {hdj mbo bdjov$ koffjdja =4 kubdf {dlj mblfiv lj {ho 0= }dfo fh}lglvlgov*
[ho 05 kubdf`{ok }oadljv dkoj{dffok u}o~dlvbt (T o{ `b*" =55?$ `}o hdahbdah{ok dj }ok" {ho 6 joqbt ko{of{ok kubdf`{dljv `}o djkdf`{ok dj a}ooj" `jk {ho
0> kubdf`{ok }oadljv nljk qd{hdj {ho 06 voagoj{v u}o~dlvbt dkoj{dffok `}o dj a}`t*
(M$ Vfhog`{df }ou}ovoj{`{dlj ln {ho 05 kubdf`{ok }oadljv `jk = {}`jvblf`{dljv dkoj{dffok lj {ho vo~oj qho`{ fh}lglvlgo a}luv* Kubdf`{ok
voagoj{v `}o vhlqj dj a} t" qd{h {hdj mbo bdjov }ou}ovoj{dja {hokubdf`{okaojov qd{hdj o`fhvoagoj{* [ho {} jvblf`{dljv mo{qooj q;'q? `jk q;'q1
`}o hdahbdah{ok dj }ok*
O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 06
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
4/15
vdjabo kubdf`{dlj o~oj{ qhoj {ql kubdf`{dljv qo}o uhtvdf`bbt
fblvo lj {hov`go vdv{o} fh}lglvlgov* @glja {ho04 `kkd{dlj`b
kubdf`{ok }oadljv (mo{qooj fh}lglvlgov }0'}=,6,05,0=" }='1,
8,0=" }6'}4,00" };'}?" }?'}4,00,0=" }>'}1,8,0=" }1,}8" }8'}00" `jk
}4,}00$" 6 fl}}ovuljk {l kubdf`{ok }oadljv jl{ dkoj{dffok u}o~d'
lvbt (dj a}ooj dj Nda}o 0@$* [hot fl~o} 05*;/ ln {ho aojlgo`jk koffjo jl~ob }ob`{dljvhduv mo{qooj fh}lglvlgov }? `jk
}00" }8 `jk }00" `jk }0 `jk }6* [ho }og`djdja 0> kubdf`{dljv (dj
a}`t dj Nda}o 0@$ `}o vuo}dgulvok lj {ho u}o~dlv 06 (05 6$
kubdf`{dljv* [hot koffjo jl~ob }ob`{dljvhduv mo{qooj {ho fh}l'
glvlgov `jk}ou}ovoj{ 0;*8/ ln {hoaojlgo* [hv" dj {l{ b"{ho
=4 kubdf`{dljv fl~o} 1=/ (=>1 Gm$ ln {ho }dfo aojlgo" qd{h `j
`~o}`ao kojvd{t ln ljo aojo uo} 5*8 Gm* Qo fljfbko {h`{ {ho
dkoj{dfff {dlj mt l} go{hlk ln 05 ijlqj `jk 04 `kkd{dlj`b
kubdf`{dlj mblfiv dj }dfo ~`bdk`{ov {ho }obd`mdbd{t ln l} `uu}l`fh
nl} ko{o}gdjdja dj{o}fh}lglvlg`b kubdf`{dljv `jk koglj'
v{}`{ov d{v vonbjovv nl} n}{ho} dj{}`aojlgdf `jk dj{o}aojlgdf
flgu`}dvljv*
Dkoj{dfff`{dlj `jk Fh`} f{o}ds`{dlj ln Kubdf`{ok
]oadljv dj {ho Qho`{ Aojlgo
[l dkoj{dnt kubdf`{dljv dj {ho qho`{aojlgo" qo ff}v{ ov{`mbdvhok
` vo{ ln jdpo OV[ vopojfov qd{h {ho hdahov{ ulvvdmbo boja{h
`jk nl} qhdfh fh}lglvlg`b blf`{dljv `}o ijlqj* [ho >;=> OV[
vopojfov {h`{ h`~o mooj ft{laojo{df`bbt g`uuok lj qho`{
kobo{dlj mdjv (Pd o{ `b*" =55;$ qo}o `bdajok `a`djv{ jlj}okjk`j{
OV[ fbv{o}v u}lkfok dj {ho n}`goql}i ln Aojlub`j{o u}lcof{v
(`~`db`mbo `{ h{{u3,,}ad*~o}v`dbbov*dj}`*n},k`{`,m`jiv,$* _vdja FDU
`jk F@BU ~`bov ln 4? `jk 8?/" }ovuof{d~obt" 45*>/ (?8=6$ ln
{ho g`uuok OV[v qo}o fljvdko}ok {l mo dkoj{df`b (qd{h `~o}`ao
FDU `jk F@BU ~`bov ln 44*= `jk 48*8/" }ovuof{d~obt$ {l ?151
jlj}okjk`j{OV[ flj{dav `jkqo}o f bbok QOFv (nl} qho`{ OV[
flj{dav$* [ho }og`djdja >56 vopojfov {h { qo}o jl{ `vvlfd {ok
qd{h ` flj{da qo}o f bbok QOVv (nl} qho`{ g`}io}vdjabo{ljv$* Qo
{hoj ko{o}gdjok {ho jgmo} ln QOFv (`glja ?8=6$ {h`{ `}o
`vvlfd`{ok qd{h ` jdpo OV[ flj{da (`glja ?151$ vdja {ho
v`go FDU `jk F@BU ~`bov* [ho }ovb{v vhlqok {h`{ 4>*0/
(??4>$ ln {ho OV[ flj{dav `}o `vvlfd`{ok qd{h ` vdjabo QOF"
qho}o`v 6*4/ (000$ `}o `vvlfd`{ok qd{h {ql (051$ l} {h}oo (;$
QOFv* [hovo b`{{o} fl}}ovuljk {l hlglolblalvQOF vopojfov
qd{h hdah vopojfodkoj{d{t" {ho}omt }oof{dja {ho hox`ubldk j`{}o
ln {ho qho`{ aojlgo* Qhoj {ql l} {h}oo hlglolblalv QOFv
qo}o dkoj{dffok" qo vok {ho vopojfo ln {ho OV[ flj{da {h {
vhlqok {ho hdahov{ FDU ~`bo `v ` }ou}ovoj{`{d~o ln {ho hlglo'
lblalv vopojfo a}luv dj {ho }ov{ ln {ho flgu`}`{d~o `j`btvov*
[hv" `glja {ho djd{d`b >;=> g`uuok OV[" 48*=/ (?"151
>56 >605$ `}o `vvlfd`{ok qd{h `j OV[ flj{da {h`{ }ou}ovoj{v `
vdjabo blfv lj ` a}lu ln hlglolblalv fh}lglvlgov* G`u'
udja djnl}g`{dlj nl} {ho ?151 QOFv `jk >56 QOVv (Pd o{ `b*"
=55;$ vhlqok {h`{ ?556" 4;>" ==;" `jk 061 qo}o `vvdajok {l
ljo" {ql" {h}oo" `jk gl}o {h`j {h}oo blfd dj qho`{" }ovuof{d~obt*
[ho ?556 QOFv `jk QOVv g`uudja `{ ` vdjabo blfv qo}o vok
nl} n}{ho} `j btvdv ln {ho flbdjo`}d{t qd{h {ho }dfo aojlgo"
qho}o`v {hlvo g`uudja {l {ql kdv{djf{ blfd (4;>$ qo}o vok {l
v{kt dj{}`aojlgdf kubdf`{dljv dj qho`{*
@glja {ho 4;> kubdf`{ok QOFv `jk QOVv" >68 qo}o `vvl'
fd`{ok qd{h ` aojo{df ulvd{dlj qd{hdj ` vuofdfff kobo{dlj mdj `v
koffjok mt Pd o{ `b* (=55;$* Qo dajl}ok {ho }og`djdja 658 vo'
pojfov" `v {ho djnl}g`{dlj q`v bdgd{ok {l {hod} u}ovojfo lj `
fh}lglvlgo l} ` fh}lglvlgo `}g* U{`{d~o dj{o}fh}lglvlg`b
kubdf`{dljv qo}o dkoj{dffok {h}lah {ho v{`{dv{df b `j`btvdv ln {ho
=0 ulvvdmbo u`d}qdvo flgmdj`{dljv nl}gok mt kubdf`{ok QOV
`jk QOF blfd `glja {ho vo~oj qho`{ fljvojvv fh}lglvlgoa}luv (nl} ` a}`uhdf`b kdvub`t ln `bb ln {ho }ob`{dljvhduv" voo
Vuubogoj{`b Nda}o 0 ljbdjo$* [ho jgmo} ln u{`{d~obt kubd'
f`{ok QOF l} QOV blfd }`jaok n}lg 06 {l 4; uo} fh}lglvlgo
u`d}" qd{h ` {l{`b ln >68* Qo lm{`djok {ho b`}aov{ jgmo}v nl} {ho
q;'q? `jk q;'q1 flgmdj`{dljv (voo Vuubogoj{`b Nda}ov
0'0> `jk 0'08 ljbdjo$* Ljbt =0> (66*4/$ ln {ho >68 u{`{d~o
kubdf`{dljv qo}o ~`bdk`{ok vdja Fblvo_u (kojvd{t }`{dl" 5*?9
fbv{o} boja{h" =?9 g`{fh" ?$ `jk {ho djnl}g`{dlj lj {ho ulvd{dlj
ln {ho QOV `jk QOF vopojfov lj fljvojvv fh}lglvlgov
(voo Go{hlkv `jk Vuubogoj{`b [`mbo = ljbdjo$* [hot koffjo 0=
v{`{dv{df`bbt vdajdfff`j{ kubdf`{dlj mblfiv {h`{ fl~o} >1*?/ ln {ho
aojlgo `jk fl}}ovuljk {l {ho nlbblqdjafh}lglvlgo u`d}v3 q0'
q= (0>OV[v$" q0'q6 (00OV[v$" q0'q; (? OV[v$"q0'q1 (1 OV[v$"
q='q; (> OV[v$" q='q1 (4 OV[v$" q6'q? (0; OV[v$" q6'q1 (0;OV[v$"q;'q? (>4 OV[v$"q;'q1 (;?OV[v$" q?'q1 (05OV[v$" `jk
q>'q1 (05 OV[v$ (Nda}o 0M$* Jl v{`{dv{df`bbt vdajdfff`j{ kubdf`'
{dljv qo}o ko{of{ok mo{qooj q0'q?" q0'q>" q='q>" q6'q>"
`jk q;'q> (voo Vuubogoj{`b [`mbo = `jk Vuubogoj{`b Nda}o
0 ljbdjo$* Qo nljk {ho hdahov{ jgmo} ln kubdf`{ok aojov
mo{qooj fh}lglvlgov q;'q? (Jl* 4 dj Vuubogoj{`b [`mbo =
ljbdjo9 >4 u`}`blav$ `jk q;'q1 (Jl* 05 dj Vuubogoj{`b [`mbo =
ljbdjo9 ;? u`}`blav$* [hovo }oadljv fl}}ovuljk {l {ql ijlqj
{}`jvblf`{dljv dj qho`{ (Gdfiobvlj'Tlja o{ `b*" 044?9 Gdn{`hkdj
o{ `b*" =55;$* [hot `uuo } `v kubdf {dljv dj l} `j btvdv mof vo
qo vok fljvojvv fh}lglvlgov" `jk qhoj ` hlglolblalv
}oadlj h`v mooj {}`jvblf {ok {l `jl{ho} fh}lglvlgo" d{ qdbb
vhlq vdgdb`}d{t {l {ho }og`djdja hlglolblalv }oadlj lj {ho
fh}lglvlgo a}lu ln l}dadj* [l kdv{djadvh mo{qooj kubdf`'
{dljv `jk {}`jvblf {dljv" qo vtv{og`{df`bbt }o`j btsok o~o}t
kubdf`{ok }oadlj lj o`fh ln {ho hlglolblalv fh}lglvlgo
a}luv (vooVuubogoj{`b Nda}ov 0'0 {l 0'=0 ljbdjo$* [ho }ovb{v
fljff}gok {h`{ ljbt {ho kubdf`{dljv dkoj{dffok mo{qooj q; `jk
q? `v qobb `v mo{qooj q; `jk q1 fl}}ovuljk {l {}`jvblf`{dljv*
[hv" qd{h {hdv aojlgo'qdko `j`btvdv" qo dkoj{dffok 05 kubd'
f`{ok }oadljv `jk = {}`jvblf`{dljv lj {ho vo~oj qho`{ fh}lgl'
vlgo a}luv* [ho dkoj{dfff`{dlj ln 05 qobb'koffjok kubdf`{ok
}oadljv `bblqok v {l n}{ho} v{kt {hod} l}dadj {h}lah flgu`}'
dvljv qd{h {ho }dfo kubdf`{dljv*
Dkoj{dfff {dlj ln L}{hlblalv ]oadljv mo{qooj
]dfo `jk Qho`{
[l dkoj{dnt l}{hlblalv }oadljv mo{qooj {ho }dfo `jk qho {
aojlgov" qo `bdajok {ho ?556 QOF `jk QOV vopojfov {h`{
g`uuok `{ ` vdjabo blfv dj qho { `a`djv{ {ho ;=">?; }dfo aojov*
_vdja >5/ FDU `jk 15/ F@BU nl} {ho vopojfo `bdajgoj{
(MB@V[J$"6>/ (085?$ ln {hoqho`{vopojfov vhlqokvdgdb }d{t
(`~o}`ao ~`bo ln 86*4/ FDU `jk 45*>/ F@BU$ {l ` jdpo aojo
dj }dfo* Vmvopoj{ Fblvo_u `j`btvdv (kojvd{t }`{dl" =9 fbv{o}
boja{h" =59 g`{fh" ?$ djkdf`{ok {h`{ 0058 ln {hog f`j mo fljvdk'
o}ok {}o l}{hlblav* [hot `}o u}ovoj{ dj 06 l}{hlblalv }oadljv
{h { fl~o} 86*0 `jk 45*;/ ln {ho }dfo `jk qho { aojlgov"
0; [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
5/15
}ovuof{d~obt" `jk fl}}ovuljk {l {ho nlbblqdja fh}lglvlgo
u`d}v3 q0'}? (05= aojov$" q0'}05 (;6 aojov$" q='}; (005 aojov$"
q='}1 (?0 aojov$" q6'}0 (=51 aojov$" q;'}6 (0;8 aojov$" q;'}00
(01 aojov$" q?'}6 (;5 aojov$" q?'}4 (6> aojov$" q?'}0= (;=
aojov$" q>'}= (0?> aojov$" q1'}> (4? aojov$" `jk q1'}8 (>0
aojov$ (Nda}o =@9 voo Vuubogoj{`b [`mbo 6 ljbdjo$* Dj n`f{" {hoq?'}6 }ob`{dljvhdu klovjl{ }oof{ {}o flbdjo`}d{t mo{qooj {hovo
fh}lglvlgov m{ {ho qho`{ {}`jvblf`{dlj mo{qooj q; `jk q?
`jk {ho l}{hlblat mo{qooj qho`{ fh}lglvlgo ; `jk }dfo
fh}lglvlgo 6* [hv" dj {l{`b" 0= {}o l}{hlblalv }ob`{dljvhduv
f`j mo koffjok (Nda}o 6$*
Qo {hoj flgu`}ok {ho bdjo`} l}ko} ln {ho }dfo aojov qd{h {ho
ulvd{dljv ln {ho l}{hlblalv OV[v dj {ho qho`{ kobo{dlj mdjv* [ho
}ovb{v djkdf`{o {h`{ nl} =1*=/ ln {hog" {ho dkoj{dffok qho`{ l}'
{hlbla dv jl{ blf`{ok dj {ho l}{hlblalv qho`{ kobo{dlj mdj"
{ho}omt djkdf`{dja }o`}}`jaogoj{v qd{hdj l}{hlblalv }oadljv*
Nl} ox`gubo" ln 0;4 l}{hlblav u}ovoj{ lj }dfo fh}lglvlgo
0 `jk qho`{ fh}lglvlgo 6M" =0 (0;*0/$ `}o jl{ nljk dj {ho
l}{hlblalvqho`{ mdjv (Nda}o =M$* Dj `kkd{dlj"qo lmvo}~ok {ql
b`}ao dj~o}vdljv dj~lb~dja kobo{dlj mdjv 6MV4" 6MV0" `jk f'6MV0mo{qooj q6M `jk }0 (Nda}o =M$* [hv" l} }ovb{v vhlq {h`{"
o~oj dn {ho f}}oj{ umbdf vo{ ln g`uuok qho`{ OV[ u}ovoj{v
vlgo bdgd{v dj flgu`}`{d~o `j`btvdv" vdjfo {ho bdjo`} l}ko} ln {ho
OV[v qd{hdj ` qho`{ kobo{dlj mdj dv jl{ ijlqj" }o`}}`jaogoj{v
f`j mo dkoj{dffok `jk{ho o~ b {dlj ln flbdjo`}d{t mo{qooj qho`{
`jk}dfo f`jmo dgu}l~ok {h}lah `j `ff} {o`vvovvgoj{ln {ho
vdsov `jk ulvd{dljv ln {ho l}{hlblalv }oadljv*
Dkoj{dfff`{dlj ln Vh`}ok Kubdf`{dljv mo{qooj
]dfo `jk Qho`{
Dj {hdv v{kt" qo dkoj{dffok =4 kubdf`{dljv dj }dfo" 05 dj qho`{"
`jk 0= }oadljv ln l}{hlblat mo{qooj {ho {ql aojlgov* Vo~oj ln
{ho dj{}`vuofdfff kubdf`{dljv `}o fljvo}~ok `{ l}{hlblalv ul'
vd{dljv mo{qooj }dfo `jkqho`{(Nda}o;@$* [hot `}onljklj {ho
nlbblqdja fh}lglvlgo u`d} flgmdj`{dljv3 q0'q=,}?'};" q0'q6,
}?'}0" q0'q;,}05'}6" q='q;,}1'}6" q='q1,};'}8" q?'q1,}4'}8"
`jk q>'q1,}='}> (ko{`dbok dj Vuubogoj{ b [`mbo ; ljbdjo$*
@b{lao{ho}" {hot }ou}ovoj{ >8*6/ ln {ho }dfo aojlgo `jk >?*4/
ln {ho qho`{ aojlgo* Ljo ln {ho b }aov{ vh`}ok kubdf`{dlj (Jl* =
dj Vuubogoj{`b [`mbo ; ljbdjo$ fl}}ovuljkv {l ` kubdf {dlj
mo{qooj q0 `jk q6 {h`{ dv l}{hlblalv {l {ho }0 `jk }? kubdf`'{dlj* [ho fljvo}~ {dlj ln kubdf {dljv mo{qooj {ho }dfo `jk qho {
aojlgov djkdf`{ov {h`{ {hot u}lm`mbt l}dadj`{ok n}lg `j `jfdoj{
kubdf`{dlj o~oj{ {h`{ u}ok`{ok {ho kd~o}aojfo mo{qooj {ho {ql
vuofdov" ?5 {l 15 GT@* Jl{ `bb fh}lglvlgov vhlq }og`djv ln
`jfdoj{ vh`}ok kubdf`{dljv* Jl l}{hlblalv kubdf`{dljv qo}o
dkoj{dffok mo{qooj }dfo fh}lglvlgov 00 `jk 0= `jk {hod} qho`{
l}{hlblavq; `jk q? (Nda}o6$*[hdvdv u}lm`mbt ko {l{ho n`f{{h`{
q; `jk q? h`~o mooj dj~lb~ok dj }ofoj{ {}`jvblf`{dljv" {ho}omt
kdv}u{dja {ho l}{hlblalv }ob`{dljvhduv*
[l v{kt {ho l}dadj `jk no`{}ov ln {ho vh`}ok kubdf {ok
}oadljv dj gl}o ko{`db" qo `j`btsok n}{ho} ljo ln {ho b`}aov{
vh`}ok kubdf`{dljv {h`{ dj~lb~ov qho`{ fh}lglvlgov 6M `jk
0M `jk }dfo fh}lglvlgov 0 `jk ?* [ho }0'q6M l}{hlblalv
}oadljv `}o }ob`{ok {h}lah 0;4 aojov" qhdbo >> aojov `}o flj'vo}~ok mo{qooj {ho l}{hlblalv }?'q0M }oadljv (Nda}o ;M$* [l
dkoj{dnt u`}`blalv vopojfov qd{hdj {ho dj{}`vuofdfff kubdf`'
{dljv" qo vok {ho =;> `jk 00? QOF `jk QOV vopojfov {h`{
g`u `{ jdpo ulvd{dljv dj {ho kubdf`{ok }oadljv ln q6 (kobo{dlj
mdj" 6MB=" 6MB05" `jk 6MB1 nl} q6M YNda}o ;M9 voo Vuubo'
goj{`bNda}o 0'=ljbdjoW$`jk q0 (0MB0"0MB=" `jk0MB6 nl} q0M
YNda}o ;M9 voo Vuubogoj{`b Nda}o 0'= ljbdjoW$ {l uo}nl}g `
MB@V[J `bdajgoj{ qd{h 15/ FDU `jk 15/ F@BU ~`bov* [hdv
dkoj{dffok odah{ u{`{d~o u`} blalv aojov qd{hdj {ho q0'q6
kubdf`{ok }oadlj* Dj }dfo" {ho l}{hlblalv }oadljv lj fh}lgl'
vlgov 0 `jk ? flj{`dj ;= u } blav (Nda}o ;M$* Qo{hojvok{ho
u`{{o}j ln jfbol{dko vmv{d{{dlj qd{hdj {ho vh`}ok kubdf`{ok
}oadljv {l ov{dg`{o {ho kubdf`{dlj {dgo dj {ho `jfov{}`b }dfo `jk
qho`{ aojlgov* [ho 8 qho`{ `jk ;= }dfo vopojfov qo}o vm'
cof{ok {l ` vtjljtglv jfbol{dko vmv{d{{dlj `j`btvdv (voo
Vuubogoj{`b [`mbo ? ljbdjo$* [l ~`bdk`{o {ho }ovb{v" qo `bvl
Nda}o =* Dkoj{dfff`{dlj ln 06 L}{hlblalv ]oadljv mo{qooj ]dfo `jk Qho`{*
(@$ Vfhog`{df }ou}ovoj{`{dlj ln {ho 06 l}{hlblalv }oadljv dkoj{dffok mo{qooj }dfo (}0 {l }0=$ `jk qho`{ (q0 {l q1$ fh}lglvlgov* [ho 0058 u`d}v ln
l}{hlblalv aojov `}o koudf{ok `v {hdj mbo bdjov*
(M$ Vfhog`{df }ou}ovoj{ {dlj ln {ho 0;4 l}{hlblav (~o}{df`b bdjov$ dkoj{dffok mo{qooj qho { fh}lglvlgo 6M (q6M$ jk }dfo fh}lglvlgo 0 (}0$* Kdnno}oj{
flbl}ok mblfiv }ou}ovoj{ {ho flbdjo`} }oadljv dkoj{dffok mo{qooj }0 `jk q6M* ]o`}} jaogoj{v mo{qooj l}{hlblalv aojov `}o hdahbdah{ok qd{h }ok
bdjov* [ql b`}ao dj~o}vdljv ln flbdjo`} }oadljv `}o djkdf`{ok qd{h `}}lqv `ml~o `jk moblq {ho qho`{ `jk }dfo fh}lglvlgov" }ovuof{d~obt* Qho`{
kobo{dlj mdjv `}o djkdf`{ok `ml~o {ho 6M fh}lglvlgo" qho}o v }dfo fh}lglvlgo 0 dv kd~dkok dj{l 05'Gm voagoj{v*
O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 0?
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
6/15
`j`btsok =5 u`}`blalv vopojfov }`jklgbt vobof{ok n}lg {ho
}00'}0= kubdf`{dlj* _vdja ` g{`{dlj }`{o ln >*? 6 054 vmv{d'
{{dljv uo} vtjljtglv vd{o uo} to`} (A`{ o{ `b*" 044>$" {ho
}ovb{v djkdf {o {h`{ {ho }00'}0= kubdf`{dlj o~oj{ dj }dfo lff}}ok
mo{qooj 0; `jk =1*6 GT@" qhdfh dv fljvdv{oj{ qd{h {ho =0 GT@
vaaov{ok mt T o{ `b* (=55?$* [ho kubdf`{dlj mo{qooj }0 `jk
}? dv ov{dg`{ok {l h`~o lff}}ok ?6*= {l 1>*6 GT@ dj }dfo9 {ho
qho`{ q6'q0 kubdf`{dlj {dgo dv ov{dg {ok`v 84*4 {l 0=8*6 GT@*
[hv" l} ov{dg`{ov `}o dj `a}oogoj{ qd{h umbdvhok kd~o}aojfo
{dgov nl} }dfo `jk qho`{ n}lg {hod} flgglj `jfov{l} (?5 {l 15
GT@$ `jk vuul}{ {ho dko` {h`{ kubdf {dljv lff}}ok dj {ho
a}`vv aojlgo `jfov{l} monl}o {ho kd~o}aojfo ln {ho kdnno}oj{
a}`vv vuofdov*
@ Glkob nl} {ho V{}f{} b O~lb{dlj ln ]dfo `jk Qho`{
n}lg ` Flgglj Fo}o`b @jfov{l}
Qo flgmdjok {ho k`{` lm{`djok dj {hdv v{kt lj vh`}ok kubd'
f`{dljv mo{qooj qho { `jk }dfo qd{h u}o~dlv flgu`} {d~o
`j`btvov uo}nl}gok mo{qooj }dfo `jk g`dso (V`bvo o{ `b*" =55;9
Qod o{ `b*" =551$ `jk mo{qooj }dfo `jk vl}ahg (U`{o}vlj o{ `b*"
=55;$* [ho g`dso `jk vl}ahg fh}lglvlgov nobb dj{l {ho 0=
a}luv ln l}{hlblat {h { qo h`k koffjok mo{qooj }dfo `jk qho`{
(Nda}o 6$* @j`btvdv ln {ho fljvo}~`{dlj u`{{o}j }ovb{ok dj {ho
koffjd{dljln ff~o`jfov{} b mblfiv(@?" @1"@00" @8"`jk @;9 Nda}o
6$ flj{ djdja l}{hlblalv fh}lglvlgov {h`{ oxhdmd{ vh`}ok
`jfov{}`b kubdf`{dljv* [ho ko{`dbok `j`btvdv ln {ho kubdf`{dlj
Nda}o 6* L}{hlblalv ]ob`{dljvhduv `jk Vh`}ok Kubdf`{dljv mo{qooj {ho ]dfo" G`dso" Qho`{" `jk Vl}ahg Aojlgov*
Flbdjo } qho`{" }dfo" vl}ahg" `jk g`dso fh}lglvlgov `}o kdvub`tok lj {ho v`go bdjo dj {ho ffa}o* Kubdf`{dljv `}o djkdf`{ok qd{h vlbdk bdjov* [ho ff~o
mblfiv ln vh`}ok kubdf`{dljv dkoj{dffok dj {ho nl} aojlgov `}o kdvub`tok lj {ho }dah{ vdko ln {ho `jfov{} b fh}lglvlgov (@?" @1" @00" @8" `jk @;$ {h`{
{hot koffjo* [ho `}{on`f{`b vtj{ojdf }ob`{dljvhdu dkoj{dffok mo{qooj q? `jk }6 {h`{ }oof{v {ho qho`{ q;'q? {}`jvblf`{dlj dv djkdf`{ok dj u`}oj{hovov*
[ho {ql kubdf`{dljv vh`}ok mo{qooj qho`{ `jk }dfo fh}lglvlgov {h`{ kl jl{ vh`}o ` flgglj `jfov{}t (q0'}05,q='}1 `jk q1'}>,q='};$ m{ `}o
nljk lj l}{hlblalv fh}lglvlgov dj ml{h vuofdov `}o djkdf {ok qd{h klmbo `}}lqv*
0> [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
7/15
u`{{o}jv mo{qooj }dfo" qho`{" vl}ahg" `jk g`dso qd{hdj o`fh
mblfi (Nda}o 6$ bok v {l u}lulvo ` glkob (Nda}o ?$ nl} {ho
o~lb{dlj ln {hovo nl} a}`vv aojlgov n}lg ` flgglj `jfov{l}
qd{h ff~o fh}lglvlgov {h`{ qo}o j`gok @;" @?" @1" @8" `jk
@00" nlbblqdja {ho f}}oj{ jgmo}dja ln {ho }dfo fh}lglvlgov*
[ho ff}v{ mblfi (`jfov{}`b fh}lglvlgo @?$ fl}}ovuljkv {l {ho
}?'}0 `jk q0'q6 vh`}ok kubdf`{dlj* Dj vl}ahg" {ho fl}}ovuljk'
dja kubdf`{dlj dv nljk mo{qooj {ho vA `jk v@ fh}lglvlgov
(U`{o}vlj o{ `b*" =55;$* Dj g`dso" d{ dv blf`{ok lj fh}lglvlgov
g6 `jk g8 `v qobb `v lj fh}lglvlgov g> `jk g8 (Nda}ov 6
`jk?$* [hokubdf {dlj vh }okmo{qooj g6 `jkg8 `jkmo{qooj
g> `jk g8 }oof{v {ho }ofoj{ {o{}`ubldkds`{dlj ln {ho g`dso
aojlgo" qho}o v {ho fljvo}~ {dlj mo{qooj g6'g8 `jk g>'g8
k`{ov m`fi {l ` QAK ln {ho `jfov{}`b a}`vv aojlgo qd{h ff~o
fh}lglvlgov (Nda}o ?" o~oj{ 0$* [hdv u`{{o}j ln kubdf`{dljv
q`v }ofoj{bt fljff}gok mt Qod o{ `b* (=551$ dj ` }ofljv{}f{dlj ln
{ho g`dso aojlgo o~lb{dlj`}t hdv{l}t {h}lah ` flgu`}`{d~o
`j`btvdv mo{qooj ` hdah'}ovlb{dlj dj{oa}`{ok uhtvdf`b g`u ln
g`dso `jk {ho }dfo `jk vl}ahg aojlgov*
[ho vofljk mblfi (`jfov{}`b fh}lglvlgo @1$ fl}}ovuljkv {l
{ho }6'}1'}05 `jk q;'q='q0 vh`}ok kubdf`{dljv (Nda}ov 6 `jk
?$* Ho}o" {ho }6'}1 `jk }6'}05 kubdf`{ok }oadljv kl jl{ l~o}b`u
`jk fl~o} ?=/ ln fh}lglvlgo 6 (Nda}o 0@$* [hdv u`{{o}j ln
kubdf`{dlj }oof{v {ho l}dadj ln {ho }6 fh}lglvlgo {h}lah
{}`jvblf`{dljv `jk nvdljv mo{qooj {ho `jfov{}`b fh}lglvlgov
@1 `jk @05" `v vaaov{ok u}o~dlvbt mt Q`ja o{ `b* (=55?$
(Nda}o ?" o~oj{ =$* [ho v`go {ql kubdf`{dljv `}o fljvo}~ok dj
qho`{ mo{qooj fh}lglvlgo q; `jk {ho q= `jk q0 fh}lgl'
vlgov" vaaov{dja {h`{ qho { fh}lglvlgo ; l}dadj`{ov n}lg {ho
v`go `jfov{}`b o~oj{* Dj vl}ahg" ` kubdf`{dlj q`v lmvo}~ok
mo{qooj {ho vM `jk vF fh}lglvlgov (U`{o}vlj o{ `b*" =55;$"
`jk d{ q`v kogljv{} {ok {h { vF l}dadj`{ok n}lg ` fh}lglvlg`b
nvdlj mo{qooj {ho `jfov{}`b }6 `jk }05 fh}lglvlgov (@6 `jk
@05 dj l} glkob9 Nda}o ?$* [hdv l}dadj oxub`djv qht ljbt ljo
kubdf`{dlj f`j mo nljk dj vl}ahg (Nda}o 6$* [ho nvdlj
mo{qooj @6 `jk @05 lff}}ok dj {ho `jfov{}`b aojlgo {l g`dso
`jk vl}ahg (Nda}o ?$9 {ho}onl}o" {ho vh`}ok kubdf`{dlj nljk
dj g`dso mo{qooj {ho g0'g?'g4 `jk g1'g= fh}lglvlgov
(Nda}o 6$ }oof{v {ho v`go u`{{o}j* [hdv q`v `bvl fljff}gok mt
{ho g`dso aojlgo `j`btvdv ln Qod o{ `b* (=551$*
Mblfi 6 (`jfov{}`b fh}lglvlgo @00$ fl}}ovuljkv {l {ho bovv
fljvo}~ok kubdf`{dlj" vdjfo d{ dv ko{of{ok ljbt mo{qooj }dfo }00
`jk }0=" vl}ahg vH `jk vO" `jk g`dso g='g; `jk g6'g05
(Nda}o 6$* Dj qho`{" {hdv kubdf`{dlj f`jjl{ mo ko{of{ok" mo'
f`vo ln {ho u}o~dlvbt goj{dljok {}`jvblf {dljv {h`{ h`~o
`nnof{ok {ho l}{hlblalv q; `jk q? fh}lglvlgov*
Mblfi ; (`jfov{}`b fh}lglvlgo @8$ fl}}ovuljkv {l {ho kubd'
f`{dljv vh`}ok mo{qooj }dfo }8'}4 `jk qho`{ q1'q? (Nda}o ?$*
[hovo kubdf`{dljv qo}o `bvl lmvo}~ok mt U`{o}vlj o{ `b* (=55;$
lj {ho l}{hlblalv vl}ahg fh}lglvlgov vC `jk vM (Nda}o 6$
`jk mt Qod o{ `b* (=551$ mo{qooj g0 `jk g; `jk mo{qooj g=
`jk g1 (}oof{dja {o{}`ubldkds`{dlj$ `v qobb `v mo{qooj g0'g;
`jk g='g1 (}oof{dja {ho `jfov{}`b kubdf`{dlj$ (Nda}o 6$*
Mblfi ? (`jfov{} b fh}lglvlgo @;$ flj{ djv kubdf`{dljv
vh`}ok mo{qooj {ho }dfo }>'}='};" qho`{ q>'q= (Nda}o ?@$" `jk
vl}ahg vD" vN" `jk vK (U {o}vlj o{ `b*" =55;$ l}{hlblalv
fh}lglvlgov* [hdv u`{{o}j }oof{v {ho l}dadj ln {ho @= fh}lgl'
vlgo {h}lah ` nvdlj mo{qooj {ho @; `jk @> `jfov{}`b fh}l'
glvlgov (Nda}o ?" o~oj{ =$* [ho kubdf`{dlj mo{qooj q> `jk
q1 q`v jl{ ko{of{ok dj qho`{ mof`vo ln {ho u}o~dlvbt goj'
{dljok {}`jvblf`{dljv mo{qooj q;'? `jk q1* [hovo kubdf`{dljv
`}o fljvo}~ok dj g`dso (Qod o{ `b*" =551$ mo{qooj fh}lglvlgov
g>" g?" `jk g4" mo{qooj fh}lglvlgov g; `jk g?" `jk mo'
{qooj fh}lglvlgov g= `jk g05 `v ` }ovb{ ln {o{}`ubldkds`{dlj
ln {ho aojlgo* [ho lmvo}~ok kubdf`{dljv mo{qooj g>'g?'g4"
g;'g?" `jk g='g05 }oof{ {ho `jfov{}`b }ob`{dljvhduv mo{qooj
{ho @=" @;" `jk @> fh}lglvlgov (Nda}o ?$* Dj {ho }dfoqho`{
flgu`}dvlj" qo h`k dkoj{dffok vo~oj vh`}ok kubdf`{dljv" m{
ljbt ff~oln {hog fl}}ovuljk {l {ho`jfov{} b vh }okkubdf {dljv
{h`{ qo h`~o dkoj{dffok mo{qooj {ho nl} a}`vv aojlgov* Dj n`f{"
Nda}o ;* Vo~oj Kubdf`{dljv @}o Vh`}ok mo{qooj ]dfo `jk Qho`{*
(@$ Vfhog`{df }ou}ovoj{`{dlj ln {ho vo~oj kubdf`{ok }oadljv vh`}ok mo{qooj qho`{ `jk }dfo* [ho u`}`blalv }oadljv `}o }ou}ovoj{ok qd{h {ho v`go
flbl}v dj qho`{ ({lu$ `jk }dfo (ml{{lg$ `v qobb `v {ho fl}}ovuljkdja l}{hlblalv }oadljv dkoj{dffok mo{qooj {ho {ql vo{v ln fh}lglvlgov (foj{o}$* [ho
flbl} flko dv `v nlbblqv3 q0'q=,}?'}; (}ok$" q0'q6,}?'}0 (l}`jao$" q0'q;,}05'}6 (a}ooj$" q='q;,}1'}6 (bdah{ mbo$" q='q1,};'}8 (u}ubo$" q?'q1,}4'}8
(m}lqj$" `jk q>'q1,}='}> (k`}i mbo$* ]dfoqho`{ l}{hlblav jl{ dj~lb~ok dj vh`}ok kubdf`{dljv `}o djkdf`{ok dj a}`t*
(M$ Vfhog`{df }ou}ovoj{`{dlj ln {ho kubdf`{dljv vh`}ok mo{qooj }dfo fh}lglvlgov ? {l 0 (;= u`}`blav bdjiok mt hl}dslj{`b bdjov `{ ml{{lg$ `jk qho`{
fh}lglvlgov 0M {l 6M (ff~o u`} blav bdjiok mt hl}dslj{ b bdjov `{ {lu$* [ho }dfo `jk qho`{ flbdjo } }oadljv }0'q6M jk }?'q0M`}o bdjiok mt 0;4`jk >>l}{hlblav (~o}{df`b bdjov$" }ovuof{d~obt*
O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 01
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
8/15
Nda}o ?* Glkob nl} {ho V{}f{}`b O~lb{dlj ln {ho ]dfo" Qho`{" Vl}ahg" `jk G`dso Aojlgov n}lg ` Flgglj @jfov{l} qd{h j ? Fh}lglvlgov*
08 [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
9/15
{ho l{ho} {ql (}?'};,q='q0 `jk };'}8,q='q1$ `}o nljk lj
fh}lglvlgov {h`{ kl jl{ vh`}o ` flgglj `jfov{}t `jk `}o
vuo}dgulvoklj }oadljv {h { `}o`b}o`kt dj~lb~ok dj QAK* [hot
ljbt `uuo`}ok dj {ho flgu`}`{d~o `j`btvdv mof`vo {hot `}o
blf`{ok dj vtj{ojdf }oadljv*
[ho `j`btvdv ln kubdf`{dljv {h`{ (0$ `}o vh`}ok mo{qooj {ho}dfo" qho`{" g`dso" `jk vl}ahg aojlgov" (=$ kl jl{ l~o}b`u
qd{h o`fh l{ho}"`jk (6${dbo{lao{ho}{l fl~o} {hoaojlgov`bglv{
oj{d}obt bok{l {hoglkob ln o~lb{dlj {h { dv u}ovoj{okdj Nda}o ?"
dj qhdfh `j `jfov{}`b aojlgo qd{h ff~o fh}lglvlgov (@?" @1"
@00" @8" `jk @>$ jko}qoj{ ` QAK (o~oj{ 0$ {h`{ }ovb{ok dj ff~o
`kkd{dlj`b fh}lglvlgov3 @0" @05" @0=" @4" `jk @>* [hdv {o{}`'
ubldkds {dlj q`v nlbblqok mt {ql {}`jvblf {dljv,nvdljv {h`{
h`~o }ovb{ok dj {ql joq fh}lglvlgov" @6 (@05 @1$ `jk
@=(@;@>$ (o~oj{=$" `jk`j dj{o}gokd {o`jfov{l} qd{hj0=
(? ? =$ fh}lglvlgov* Vmvopoj{bt" {ho kdnno}oj{ a}`vv
aojlgov qlbk h`~o o~lb~ok kdnno}oj{d`bbt n}lg {hdv `jfov{}`b
aojlgdf v{}f{}o* Dj }dfo" `kkd{dlj b voagoj{ b kubdf`{dljv
lff}}ok qd{hl{ glkdntdja {ho m`vdf v{}f{}o ln 0= fh}lgl'
vlgov* [hv" {ho =4 kubdf`{dlj o~oj{v dkoj{dffok dj {ho }dfoaojlgo }ovb{ n}lg {h}oovffovvd~o }ljkv ln kubdf`{dljv3 (0$ `
QAK (o~oj{ 0$ `jk {ql fh}lglvlgo {}`jvblf {dljv (o~oj{ =$9 (=$
== `kkd{dlj b voagoj{`b kubdf {dljv l~o}b`uudja qd{h {ho
QAKv9 `jk (6$ }ofoj{ kubdf`{dljv l~o} 96 Gm `{ {ho {o}gdj`b
ojkv ln fh}lglvlgov }00 `jk }0=*
N}lg {ho `jfov{}`b aojlgo qd{h 0= fh}lglvlgov" qho`{
jko}qoj{ ff~o fh}lglvlg`b nvdljv (o~oj{ 6 dj Nda}o ?$ mo'
{qooj @? `jk @05" @> `jk @8" @4 `jk @0=" @6 `jk @00" `jk @;
`jk @1" }ovb{dja dj fh}lglvlgov q0" q1" q?" q;" `jk q="
}ovuof{d~obt* Fh}lglvlgov q6 `jk q> l}dadj`{ok kd}of{bt n}lg
{ho `jfov{}`b fh}lglvlgov @0 `jk @=" }ovuof{d~obt* [hdv
}ovb{ok dj `j `jfov{}`b qho`{ aojlgo qd{h j 1 fh}lglvlgov
(? ? = ?$* [hv" {ho 05 kubdf`{ok }oadljv lmvo}~ok ho}o dj
{ho qho`{ aojlgo }oof{ {ho `jfov{}`b QAK (o~oj{ 0" nl} vo~oj
ln {ho kubdf`{dljv$ `jk {h}oo `kkd{dlj`b voagoj{`b kubdf`{dljv
{h`{ h`~o lff}}ok vdjfo {ho fh}lglvlg`b nvdljv*
Nl} g`dso `jk vl}ahg" l} ffjkdjav `jk glkob `}o dj flg'
ubo{o `a}oogoj{ qd{h {ho }ofoj{ `j btvdv ln Qod o{ `b* (=551$" qhl
vhlqok {h`{ ml{h h`~o o~lb~ok n}lg `j `jfov{}`b aojlgo qd{h
0= fh}lglvlgov `n{o} {ql fh}lglvlg`b nvdljv (mo{qooj @6
`jk @05 `jk mo{qooj @1 `jk @49 Nda}o ?$" }ovb{dja dj `j dj'
{o}gokd`{o `jfov{l} qd{h j 05 fh}lglvlgov (? ? = =$
(o~oj{ ; dj Nda}o ?$* [hoj" g`dso `jk vl}ahg o~lb~ok djko'
uojkoj{bt n}lg {hdv `jfov{l}* Qhdbo {ho vl}ahg aojlgo v{}f'
{}o }og`djok vdgdb`} {l {ho `jfov{}`b aojlgo" g`dso jko}qoj{
` QAK }ovb{dja dj{l `j dj{o}gokd`{o qd{h j =5 fh}lglvlgov
(o~oj{ ? dj Nda}o ?$* [hdv fl}}ovuljkv {l {ho {o{}`ubldkds`{dlj
o~oj{ kovf}dmok dj u}o~dlv v{kdov (A`{ `jk Klombot" 04419
Vqdaljl~` o{ `b*" =55;$* Nlbblqdja {hdv o~oj{" jgo}lv fh}l'
glvlg`b nvdljv h`~o bok {l ` aojlgo v{}f{}o qd{h 05 fh}l'
glvlgov (j 05 Y? ? = =W 6 = 05$* @{ bo`v{ 01
fh}lglvlg`b nvdljv (o~oj{ > dj Nda}o ?$ gv{ h`~o lff}}ok {l
oxub dj {ho }ob {dljvhduv {h`{ f`j mo lmvo}~ok{lk`t mo{qooj {ho
kdnno}oj{ g`dso fh}lglvlgov*
[hv" qd{h {hdv glkob {h`{ u}lulvov ` }ofljv{}f{dlj ln {ho}dfo" qho`{" vl}ahg" `jkg`dso aojlgov n}lg `j `jfov{l} qd{h
j ? fh}lglvlgov" d{ dv ulvvdmbo {l dggokd`{obt dkoj{dnt {ho
`jfov{} b }ob`{dljvhduv jk {ho l}dadj (QAK" m}o`i`ao" nvdlj$ ln
{ho kdnno}oj{ fh}lglvlgov dj o`fh ln {ho nl} aojlgov*
KDVF_VVDLJ
Dj{} aojlgdf `jk Dj{o}aojlgdf Flgu`}dvljv _vdja
Dgu}l~ok Dj{oa}`{d~o @bdajgoj{ F}d{o}d` ]o~o b Vh`}ok
Kubdf`{dljv mo{qooj ]dfo `jk Qho {
Dj {hdv v{kt" qo vok dgu}l~ok dj{oa}`{d~o vopojfo `bdajgoj{
f}d{o}d` (FDU `jk F@BU$ flgmdjok qd{h ` v{`{dv{df`b ~`bdk`{dlj {l
}o`j`btso {ho umbdfbt `~`db`mbo k`{` vo{ ln }dfo `jk qho`{ vo'pojfov* [hdv `bblqok v {l dkoj{dnt l}{hlblav `jk u`}`blav qd{h
fljffkojfo `jk {l djno} joq }ob`{dljvhduv flgu`}ok qd{h u}o~d'
lv v{kdov {h { qo}o od{ho} m`vok ljbt lj vg bb'vf bo vopojfo
flgu`}dvljv (Atl{ o{ `b*" =55;9 Vdjah o{ `b*" =55;9 Mobb o{ `b*"
=55?9 ]dfo Fh}lglvlgo 6 Fljvl}{dg" =55?$ l} b`fiok v{`{dv{d'
f`b ~`bdk {dlj qhoj uo}nl}gok `{ {ho qhlbo aojlgo bo~ob (Vl}}obbv
o{ `b*" =5569 B` ]l{` `jk Vl}}obbv" =55;9 Vdjah o{ `b*" =551$*
Nl}go} flgu`}`{d~o v{kdov qo}o uo}nl}gok qd{h ` vopojfo
`bdajgoj{ {h}ovhlbk ln 85/ dkoj{d{t l~o} 055 m`vov nl} {ho mov{
HVU (B` ]l{` `jk Vl}}obbv" =55;$ l} qd{h ` vfl}o ln =55 nl}
MB@V[J `j`btvdv (Vdjah o{ `b*" =551$* Qo h`~o vhlqj u}o~dlvbt
{h { {hovo ~`bovg`t bo`k {l {hogdvdkoj{dfff`{dlj ln l}{hlblalv
}oadljv (V`bvo o{ `b*" =55=" =55;$" `v {hot `bblq {ho ko{of{dlj ln
gogmo}v ln aojo n`gdbdov {h`{ `}o jl{ {}bt l}{hlblalv l} ln
hlglbladov {h`{ `}o m`vok ljbt lj fljvo}~ok klg`djv*
[l ~`bdk`{o l} v{}`{oat" qo ff}v{ `uubdok l} v{}djaoj{ `bdaj'
goj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk`{dlj {l {ho }dfo aojlgo* Qo
fljff}gok `bb ln {ho kubdf`{dljv }oul}{ok u}o~dlvbt (T o{ `b*"
=55?$ `jk dkoj{dffok 04 `kkd{dlj`b }oadljv kubdf`{ok mo{qooj
kdnno}oj{ }dfo fh}lglvlgov* T o{ `b* (=55?$ h`k ov{dg`{ok {h`{
{ho kubdf`{dljv ln {ho }dfo aojlgo fl~o} >?*1/ ln {ho aojlgo*
Dj l} `j`btvdv" {ho v`go }oadljv (05 kubdf`{dljv$ fl~o} ;1*8/
ln {ho aojlgo* [ho kdnno}ojfo dv ko {l {ho `ff}`ft ln {ho
go{hlkvok {l ko{o}gdjo{ho bdgd{vln {hokubdf`{ok }oadljv* T
o{ `b* (=55?$ `klu{ok ` a}`uhdf`b `uu}l`fh mof`vo ln {ho m`fi'
a}ljk jldvo u}lkfok mt MB@V[* _vdja gl}o v{}djaoj{ f}d{o}d`
`bblqok v {l ko{o}gdjo {ho bdgd{v ln {ho kubdf {ok }oadljv
qd{h hdah fljffkojfo `jk `ff}`ft" {ho}omt }okfdja {ho vdso ln
{ho }oadljv flgu`}ok qd{h {hod} `j`btvdv* [ho `ff}`ft `jk
Nda}o ?* (flj{djok$*
Fh}lglvlgov `}o }ou}ovoj{ok qd{h flbl} flkov {l dbbgdj`{o {ho o~lb{dlj ln voagoj{v n}lg ` flgglj `jfov{l} qd{h ff~o fh}lglvlgov* [ho ff~o
fh}lglvlgov `}o j`gok `ffl}kdja {l {ho }dfo jlgojfb`{}o* Kdnno}oj{ o~oj{v {h`{ h`~o vh`uok {ho v{}f{}o ln {ho kdnno}oj{ a}`vv aojlgov k}dja
{hod} o~lb{dlj n}lg {ho flgglj `jfov{l} `}o djkdf`{ok qd{h &* Nl} o`fh vuofdov" fh}lglvlgo jgmo}v `}o nlbblqok mt ` nl}gb` djkdf`{dja {ho
o~lb{dlj }t l}dadj ln {hdv jgmo} (o*a*"dj qho {"j 1 ? Y`jfov{l}W ? YQAKW = Y`joubldk voagoj{ b kubdf {dljW ? Yfh}lglvlgo nvdljW$*Qho}o
ulvvdmbo" kd~o}aojfo {dgov `}o djkdf`{ok m`vok lj u}o~dlv ov{dg`{ov mt A`{ (=55=$" Vqdaljl~` o{ `b* (=55;$" `jk U`{o}vlj o{ `b* (=55;$*
O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv 04
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
10/15
}lmv{jovv ln l} go{hlk `}o vuul}{ok mt }ovb{v lm{`djok mt
{ho ]dfo Fh}lglvlgov 00 `jk 0= Vopojfdja Fljvl}{d` (=55?$"
qhl ov{dg`{ok {ho boja{h ln {ho glv{ }ofoj{ kubdf`{dlj lj {ho
~o}t kdv{`b ojk ln fh}lglvlgov 00 `jk 0= `v 6*6 Gm* [hdv
fl}}ovuljkv ox f{bt {l {ho boja{h {h { qo nljk qd{h l} go{hlk
(voo Vuubogoj{`b [`mbo 0 ljbdjo$ `jk dv vg`bbo} {h`j {ho ov{d'g`{ov (>*? `jk;*8 Gm lj fh}lglvlgov 00 `jk 0=" }ovuof{d~obt$
ln T o{ `b* (=55?$ nl} {ho v`go }oadlj* Gl}ol~o}" l} `j`btvdv
`bblqok v {l ko{of{ 0> jl~ob kubdf`{dljv {h`{ `}o vuo}dgulvok
lj }oadljv `b}o`kt dkoj{dffok `v kubdf`{ok" {ho}omt koffjdja `k'
kd{dlj`b `jfov{}`b }ob`{dljvhduv mo{qooj {ho }dfo fh}lglvlgov*
Dj{o}ov{djabt" {hovo }ob {dljvhduv qo}o jl{ lmvo}~ok mo{qooj {ho
l}{hlblalv fh}lglvlgov dj {ho l{ho} a}`vv aojlgov* [hdv
vaaov{v {h`{ {ho vuo}dgulvok kubdf`{dljv lmvo}~ok ho}o dj
}dfo `}o{}`fov ln ~o}t `jfdoj{ kubdf`{dljv {h { u}ok`{o {hoQAK
dkoj{dffok dj {ho a}`vv `jfov{l}* @kkd{dlj`b a}`vv aojlgo vo'
pojfov `v qobb `v `kkd{dlj`b {llbv {h`{ `bblq {ho ov{dg`{dlj ln
kd~o}aojfo {dgov mo{qooj ~o}t `jfdoj{ kubdf`{dljv qd{h hdah
`ff}`ft `}o jookok {l vuul}{ {hdv htul{hovdv*
Dj qho`{" qo dkoj{dffok 05 kubdf`{ok }oadljv {h`{ }ou}ovoj{>1*?/ ln {ho aojlgo* U}o~dlv ]NBU g`uudja v{kdov h`k
vaaov{ok `jfdoj{ kubdf`{dljv dj {ho qho`{ aojlgo* Nl} ox'
`gubo"Kmfl~vit o{ `b*(044>$ vhlqok{h`{ 60/ln {ho]NBU blfd
dj {ho kdubldk [}d{dfg gljlflffg g`u `}o u}ovoj{ gl}o {h`j
ljfo dj {ho aojlgo" `jk Pd o{ `b* (=55;$ }oul}{ok {h`{ 04/ ln {ho
OV[ g`}io}v vok nl} kobo{dlj mdj g`uudja qo}o nljk lj jlj'
hlglolblalv vo{v ln fh}lglvlgov* _j{db jlq" hlqo~o}" {ho
ox{oj{ `jk blf`bds`{dlj ln {ho kubdf {ok }oadljv qo}o jl{ koffjok
u}ofdvobt" `jk l} }ovb{v u}l~dko ` udf{}o ln kubdf`{dljv dj {ho
qho`{ aojlgo* [ho 05 dkoj{dffok kubdf`{dljv `}o u}lm`mbt `j
jko}ov{dg`{dlj" `jk d{ dv ulvvdmbo {h`{ ` jgmo} ln kubdf`{dljv
{h`{ qo}o jl{ v{ {dv{df bbt ~`bdk`{ok dj {hdv v{kt (o*a*" q='q6"
q='q?" `jk q?'q>$ qdbb mo fljff}gok dj {ho n{}o `v `kkd{dlj`b
aojo'g`uudja djnl}g`{dlj moflgov `~`db`mbo* @{ {hdv {dgo" l}
`j`btvov kogljv{}`{o `b}o`kt {h`{ glv{ ln {ho aojlgo dv kubd'
f`{ok `jk v{}ljabt vaaov{ `j `jfdoj{ kubdf`{dlj ln {ho kdubldk
qho`{ aojlgov monl}o {hod} htm}dkds`{dlj dj{l ulbtubldk qho`{*
Nlbblqdja {ho v`go `uu}l`fh" qo }oox`gdjok {ho flbdjo`}d{t
mo{qooj }dfo `jk qho`{ `jk dkoj{dffok 06 flbdjo`} }oadljv {h`{
fl~o} 86*0 `jk 45*;/ ln {ho }dfo `jk qho`{ aojlgov" }ovuof'
{d~obt* U}o~dlv flgu`}`{d~o `j`btvov mo{qooj {ho }dfo k}`n{
vopojfov `jk qho`{ OV[v (Vl}}obbv o{ `b*" =5569 Fljbot o{ `b*"
=55;9 B` ]l{ `jk Vl}}obbv" =55;9 Bdjidoqdfs o{ `b*" =55;9
Gji~lbk o{ `b*" =55;9 ]`jkh`q` o{ `b*" =55;$ `bvl nljk 06 l}'
{hlblalv voagoj{v mo{qooj {ho {ql aojlgov* Ho}o" qo qo}o
`mbo {l dkoj{dnt gl}o u}ofdvobt {ho bdgd{v ln {ho l}{hlblalv
}oadljv `jk {l ko{of{ gl}o }o`}}`jaogoj{v {h`j u}o~dlvbt
kovf}dmok vdja ` flgu`}dvlj mo{qooj {ho flgubo{o }dfo
aojlgo vopojfo (Dj{o}j`{dlj`b ]dfo Aojlgo Vopojfdja
U}lcof{" =55?$ `jk ` jdpo vo{ ln g`uuok qho`{ OV[v qd{h
{ho hdahov{ ulvvdmbo boja{h* Qd{h {hdv bo~ob ln }ovlb{dlj" d{ dv jl
bljao} ulvvdmbo {l vubd{ qho`{ fh}lglvlgov dj{l ` glv`df ln
}dfol}{hlblalv }oadljv (Vl}}obbvo{ `b*" =5569Vl}}obbv" =55;$" `jk
{ho flbdjo`}d{t mo{qooj {ho {ql aojlgov dv gl}o n}`agoj{ok
{h`{ u}o~dlvbt `vvgok mt flgu`}`{d~o g`uudja* Ho}o" {ho
ox{oj{ ln ko{of{`mbo }o`}}`jaogoj{v dv fblvo} {l {h`{ lmvo}~ok
{h}lah flgu }dvljv mo{qooj qho`{ M@F vopojfov `jk
l}{hlblalv }dfo aojlgo vopojfov" dj qhdfh g`jt oxfou{dljv
{l gdf}lflbdjo`}d{t h`~o mooj lmvo}~ok (}o~doqok dj V`bvo `jk
Nodbbo{" =551$*
Mt flgmdjdja k`{` n}lg {ho dj{}`vuofdfff `jk dj{o}vuofdfff
flgu`}dvljv" qo dkoj{dffok vo~oj kubdf`{dlj o~oj{v {h`{ `}o
fljvo}~ok `{ l}{hlblalv ulvd{dljv mo{qooj }dfo `jk qho`{* Dj
{hdv v{kt" qo vok ` flgmdj`{dlj ln djkd~dk`b `j`btvdv ln kubd'f`{dljv qd{hdj o`fh aojlgo `jk flbdjo`}d{t `vvovvgoj{ mo'
{qooj {ho {ql aojlgov vdja dkoj{df`b `bdajgoj{ f}d{o}d` `jk
v{`{dv{df`b ~`bdk`{dlj* U}o~dlv `{{ogu{v {l dkoj{dnt vh`}ok kubd'
f`{dljv mo{qooj qho`{ `jk }dfo qo}o ljbt m`vok lj {ho j`btvdv
ln }oadljv {h`{ fl}}ovuljk {l kubdf`{ok }oadljv dj }dfo vhlqdja
` hdah u}lul}{dlj ln aojo g`{fhov qd{h {ql kdnno}oj{ qho`{
fh}lglvlgo a}luv* [hdv k`b vtj{ojt `uu}l`fh vok }ofoj{bt
mt Vdjah o{ `b* (=551$ q`v m`vok lj }dfo aojov u}ovoj{ dj 8 ln {ho
05 }oadljv u}o~dlvbt ijlqj {l mo kubdf`{ok (T o{ `b*" =55?$
`jk {ho dkoj{dfff`{dlj ln hlglblalv qho`{ OV[v {h`{ g`u dj
kobo{dlj mdjv lj {ql kdv{djf{ qho`{ fh}lglvlgov* _vdja {hdv
`uu}l`fh lj {ho 06 l}{hlblalv }oadljv {h`{ qo}o dkoj{dffok dj
l} v{kt" qo nljk {h`{ }0 dv l}{hlblalv {l q6 (=51 fljvo}~ok
aojov$ `jk {l q0 (06 fljvo}~ok aojov$ m{ `bvl {l q= qd{h {hov`go vdajdfff`jfo (00 fljvo}~ok aojov$* Vdgdb`}bt" }? dv l}{hlb'
lalv {l q0 (05= fljvo}~ok aojov$ `jk {l q6 (4 fljvo}~ok
aojov$ m{ `bvl {l q= (8 fljvo}~ok aojov$ (k`{` jl{ vhlqj$*
[hv" dj flj{}`v{ qd{h l} v{}`{oat" d{ q`v jl{ ulvvdmbo qd{h {hdv
`uu}l`fh {l ~`bdk`{o v{`{dv{df`bbt {h`{ {ho }0'}? kubdf`{dlj dj }dfo
dv vh }ok mo{qooj q6'q0 dj qho`{* Vdjah o{ `b* (=551$ `bvl
fljfbkok {h`{ {ho }oadlj kubdf`{ok mo{qooj }00 `jk }0= dv
fljvo}~ok dj qho`{ mo{qooj q;'q?" vaaov{dja {h`{ {ho kubd'
f`{dlj u}ok`{ok {ho }dfo `jk qho`{ kd~o}aojfo* [hdv fljdf{v
qd{h l} ffjkdjav `v qobb `v qd{h u}o~dlv v{kdov djkdf`{dja {h`{
{ho}00'}0= kubdf`{dlj dv vuofdfff {l {ho}dfo bdjo`ao `jk}ob {d~obt
}ofoj{ (T o{ `b*" =55?$* [hovo ffjkdjav u}l~dko ` jl{o ln f`{dlj
nl} flgu`}`{d~o aojlgo'qdko `j`btvov `jk vaaov{ {h`{ `uubt'
dja {ho v`go v{}djaoj{ `bdajgoj{ f}d{o}d` `jk v{`{dv{df`b ~`bdk {dlj
{l o`fh ln {ho k`{` vo{v vok nl} aojlgo flgu`}dvljv dv ov'
voj{d`b {l `~ldk gdvdj{o}u}o{`{dlj ln o~lb{dlj`}t u`{{o}jv* Ho}o"
mt m`vdja l} dj{}`aojlgdf `jk dj{o}aojlgdf flgu`}dvljv lj
{ho hdahov{ dkoj{d{t l~o} {ho bljaov{ vopojfo `bdajgoj{" qo
vok ` fljvo}~`{d~o `uu}l fh {h`{ oxfbkok vopojfov {h`{
flbk bo`k ul{oj{d`bbt {l `j l~o}ov{dg`{dlj ln vopojfo fljvo}'
~`{dlj qd{hdj `jk mo{qooj {ho }dfo `jk qho`{ aojlgov* Jo~o}'
{hobovv" qo qo}o `mbo {l ko{of{ `kkd{dlj b kubdf`{dljvdj }dfo `jk
u}l~dko ` ko{`dbok `vvovvgoj{ ln aojlgo kubdf`{dljv dj qho`{"
kogljv{} {dja {ho }lmv{jovv ln {hdv `uu}l`fh* Ndj bbt" {ho
vo~oj kubdf`{dljv {h`{ `}o vh`}ok dj }dfo `jk qho`{ `}o bdiobt
jko}ov{dg`{ok" vdjfo {ho k`{` vo{ ln qho`{ aojov nl} qhdfh `
fh}lglvlg`b ulvd{dlj dv ijlqj dv f}}oj{bt bdgd{ok* N{}o b`}ao
OV[ g`uudja l} aojlgo vopojfdja u}lcof{v qdbb mo }opd}ok {l
lm{`dj ` flgubo{o `vvovvgoj{ ln {ho kubdf`{dljv dj {ho qho`{
aojlgo `jk ln {hod} fljvo}~`{dlj qd{h {ho }dfo aojlgo*
O~lb{dlj ln {ho A}`vv Aojlgov n}lg `jj? ?
Flgglj @jfov{l}
Djkd~dk`b aojlgo kubdf`{dlj `j`btvov dj }dfo" vl}ahg" `jk
g`dso h`~o `b}o`kt vaaov{ok `j `jfov{}`b QAK u}ok`{dja {ho
kd~o}aojfo mo{qooj {ho kdnno}oj{ fo}o`b aojlgov (U`{o}vlj
o{ `b*" =55;9 T o{ `b*" =55?9 Q`ja o{ `b*" =55?$" m{ {l k`{o {ho
=5 [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
11/15
l}dadj`b m`vdf jgmo} ln fh}lglvlgov }og`djok jfo}{`dj
(A`{" =55=$* ]ofoj{bt" Qod o{ `b* (=551$ {hl}lahbt v{kdok {ho
l}dadj ln {ho g`dso fh}lglvlgov `jk u}lulvok ` glkob nl} {ho
o~lb{dlj ln {ho fo}o`b aojlgov n}lg `j `jfov{l} qd{h j 0=
fh}lglvlgov qhlvo v{}f{}o dv vdgdb } {l {ho }dfo aojlgo* L}
glkob dv dj flgubo{o `a}oogoj{ qd{h {ho u}lulvok o~lb{dlj`}tu`{h n}lg {hdv `jfov{l} m{ u}l~dkov `kkd{dlj`b o~dkojfo nl} {ho
l}dadj ln {ho 0= `jfov{}`b fh}lglvlgov*
Mt dj{oa} {dja joq k`{` lj {ho kubdf`{dljv ln {ho qho`{
aojlgo dj{l ` ko{`dbok flgu`}`{d~o `j`btvdv ln {ho u`{{o}j ln
vh`}ok kubdf`{dljv mo{qooj }dfo" qho`{" g`dso" `jk vl}ahg"
qo qo}o `mbo {l u}lulvo ` m`vdf fh}lglvlgo jgmo} ln j ?
nl} {ho flgglj `jfov{l} ln {ho fo}o`b aojlgov* Qo vaaov{
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{l 15 GT@" {ho `jfov{l} qd{h j ? fh}lglvlgov jko}qoj{
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`jk nvdljv bok {l {ho fljv{}f{dlj ln {ql joq fh}lglvlgov"
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lvbt* [hdv vfoj`}dl }ofljfdbov u}o~dlv flj{}`kdf{l}t vaaov'{dljv {h { h`k u}lulvok a}`vv `jfov{l} qd{h od{ho} j ? l}j
0= fh}lglvlgov (A`{" =55=$" vdjfo {ho o~lb{dlj n}lg ? {l 0=
fh}lglvlgov qlbk h`~o h`uuojok monl}o {ho kd~o}aojfo
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h`~o }o{`djok {hdv l}dadj`b fh}lglvlgo jgmo}" qho}o`v d{
qlbk h`~o mooj }okfok dj {ho qho`{" g`dso" `jk vl}ahg
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fh}lglvlgo jgmo} o~lb{dlj* Fh}lglvlgo nvdljv" {}`jvbl'
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o~lb{dlj ln {ho @}`mdkluvdv {h`bd`j` aojlgo (j ?$ n}lg `j
`jfov{l} `{j 8* Flgu`}`{d~o `j`btvov qd{h {ho }ob {ok vuofdov
@}`mdkluvdv bt}`{` (j 8$`jk F`uvobb` }mobb` (j8$ h`~o vhlqj
{h`{ fh}lglvlgov 0" =" `jk ? l}dadj`{ok {h}lah {ho nvdlj ln
{}`jvblf`{ok `jk dj~o}{ok voagoj{v ln `jfov{}`b fh}lglvlgov
(nl} }o~doq" voo Vfhmo}{" =551$* [ho ff~o nvdljv vaaov{ok mt
l} glkob dj qho`{ bdiobtlff}}okdj {ho`jfov{} b aojlgoln {ho
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m`vdf fh}lglvlgo jgmo} ln j 1* U}obdgdj`}t o~dkojfo nl}
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ul}{ok dj ` m`}bot,}dfo flbdjo`}d{t v{kt (V{odj o{ `b*" =551$ vdja
{ho k`b vtj{ojt `uu}l`fh ogubltok mt Vdjah o{ `b* (=551$*
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` k}`g`{df }okf{dlj ln {ho g`dso fh}lglvlgo jgmo} (n}lg =5
{l 05$`n{o}{o{}`ubldkds`{dlj(Qod o{ `b*" =5519 l}k`{`$" l}glkob
vaaov{v {h`{ ljbt {ql fh}lglvlg`b nvdljv h`~o lff}}ok
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[hdv blq bo~ob ln }o`}}`jaogoj{v dv vdgdb`} {l qh`{ dv lmvo}~ok dj
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dv ulvvdmbo {h`{ {ho `jfov{}`b a}`vv aojlgo flj{`djok ` aojo
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ln ` qhlbo aojlgo klmbdja" vdgdb`} {l {ho Uh0 aojo dj qho`{
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`bvl u`}{dfdu {o dj b`}ao }o`}}`jaogoj{v (Mojjo{soj" =55?$"
kdnno}ojfov dj {ho `glj{ ln {}`jvulv`mbo obogoj{v g`t dj'
ojfo {ho koa}oo ln aojlgo }o`}}`jaogoj{v `v qobb* [hv" ljo
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vopojfov `}o flgubo{ok* D{ qdbb mo dj{o}ov{dja `bvl {l voo
qho{ho} l} glkob }og`djv flgu`{dmbo qd{h {ho o~lb{dlj ln {ho
ogo}adja {oguo}`{o a}`vv aojlgo glkob" M}`fhtulkdg kdv'
{`fhtlj (j ?$" qhdfh dv `{ `j dj{o}gokd`{o o~lb{dlj`}t v{`ao
mo{qooj }dfo `jk qho {(K}`uo} o{ `b*" =550$ `jk nl} qhdfh `j 86
qhlbo aojlgo vhl{aj vopojfo qdbb mo flgubo{ok qd{hdj {ho
jox{ to`} (Hl o{ `b*" =55>$* Ndj`bbt" {l vuul}{ l} htul{hovdv ln
`j `jfov{l} qd{h j ? fh}lglvlgov" `kkd{dlj`b flgu`}`{d~o
v{kdov `}o jookok qd{h aojlgov n}lg vuofdov l{vdko ln {ho
Ul`bov* Gogmo}v ln {ho @}of`bov (flflj{ YFlflv jfdno}`W"
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vfh v{kdov*
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`jt u`d}qdvo `bdajgoj{* [hovo u`}`go{o}v qo}o `uubdok {l `bb ln {ho
MB@V[ `bdajgoj{v {h { qo}o uo}nl}gok dj {hdv v{kt* @n{o} {ov{dja
kdnno}oj{ flgmdj`{dljv ln FDU `jk F@BU ~`bov dj {ho kdnno}oj{ `j`btvov"
v{}djaoj{ ~`bov (4?/ FDU" 8?/ F@BU$ qo}o vok nl} {ho dkoj{dfff`{dlj ln
{}o u`} blav `glja {hoqho`{ ([}d{dfg `ov{d~g$ OV[flj{dav" qho}o v
bovv v{}djaoj{ u`}`go{o}v qo}o `uubdok nl} {ho `j`btvov ln flbdjo`}d{t
(>5/ FDU" 15/ F@BU$ `jk kubdf`{dlj (15/ FDU" 15/ F@BU$*
Qho`{`jk ]dfo Vopojfo K`{`m`vov
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`vvdajok {l kobo{dlj mdjv mt Pd o{ `b* (=55;$ qo}o klqjbl`kok n}lg {hoA}`djAojo qomvd{o (h{{u3,,qho`{*uq*vk`*al~,$* Vdjfo vlgo OV[v g`t
fl}}ovuljk {l jljl~o}b`uudja 64 `jk ?4 ojkv ln {ho v`go fKJ@ vo'
pojfo" qo ko~obluok ` jdaojo vo{ ln g`uuok qho`{ OV[v mt `bdajdja
{hovo vopojfov `a`djv{ {ho umbdf OV[ fbv{o}v {h`{ qo}o u}lkfok dj
{ho n}`goql}i ln Aojlub`j{o u}lcof{v (h{{u3,,}ad*~o}v`dbbov*dj}`*n},k`{`,
m`jiv,$* [l ojv}o {h`{ o`fh OV[ dv `vvlfd`{ok qd{h ` jdpo OV[ flj{da
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`jk F@BU (8?/$ ~`bov qo}o `uubdok {l {ho `j`btvdv* [hdv bok {l {ho
dkoj{dfff`{dlj ln QOVv (d*o*" OV[v {h`{ `}o jl{ `vvlfd`{ok qd{h `jt OV[
fbv{o}$`jk QOFv (d*o*"OV[v`vvlfd {ok qd{h `j OV[flj{da qd{h 4?/FDU
`jk 8?/ F@BU$* QOF `jk QOV vopojfov qo}o vok nl} {ho dkoj{dff'
f`{dlj ln kubdf`{dljv qd{hdj {ho qho`{ aojlgo `jk nl} ov{`mbdvhdja
O~lb{dlj ln @jfov{} b A}`vv Aojlgo Kubdf {dljv =0
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
12/15
l}{hlblalv }ob`{dljvhduv mo{qooj }dfo (L}ts` v`{d~ $ `jk qho`{* Vdjfo
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qho`{ `jk l}{hlblav qd{h }dfo" {ho `j`btvdv q`v uo}nl}gok lj {ho vo~oj
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fh}lglvlgov ln ` ad~oj fh}lglvlgo a}lu q`v fljvdko}ok qd{h `
jdpo ulvd{dlj lj ` vdjabo fljvojvv fh}lglvlgo uo} a}lu* [ho
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JA]$$,FVd W 6 055" qho}o MV{ mdj v{`}{ fll}kdj`{o" Mvd mdj vdso (dj
uo}foj{ ao ln {ho}og`djdja fh}lglvlgo YPdo{ `b*" =55;W$" JA jgmo}
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lalv fh}lglvlgo `}gv qo}o dajl}ok*
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qo}o klqjbl kok n}lg {ho [DA] qomvd{o (n{u3,,n{u*{da}*l}a,um,k`{ ,
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vuljkdja {l OV[v {h`{ qo}o g`uuok lj {ql kdnno}oj{ hlglolblalv
fljvojvv fh}lglvlgov nl} o`fh ln {ho vo~oj fh}lglvlgo a}luv* Dj
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ln kubdf`{ok }oadljv*
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lalv Aojov U}ovoj{ dj {ho Vh }ok Kubdf`{dlj mo{qooj ]dfo
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== [ho Ub j{ Fobb
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
13/15
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
14/15
U`{o}vlj" @*H*" Mlqo}v" C*O*" Uo{o}vlj" K*A*" Ov{dbb" C*F*" `jk
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lvouhlvuh`{o dvlgo} vo dj g`dso (So` g`tv B*$* C* Ho}ok* 63 =08==8*
T" C*" o{ `b* (=55=$* @ k}`n{ vopojfo ln {ho }dfo aojlgo (L}ts` v`{d~` B*
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8/14/2019 Identification and Characterization of Shared Duplications between Rice and Wheat Provide New Insight into Grass
15/15
[hdv djnl}g`{dlj dv f}}oj{ `v ln G`}fh 6" =554
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