Taxonomical updates for Buthidae (2008 ) - NTNU...Diplocentrus poncei Francke & Quijano-Ravell, 2009...
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2020 © Jan Ove Rein, The Scorpion Files Taxonomical updates in The Scorpion Files for Scorpionidae (2008 →) Taxa Status Distribution Comments Reference Aops New genus Australia The family Urodacidae was abolished by Soleglad, Fet & Kovarik in 2005 and the subfamily Urodacinae was transfered to Scorpionidae. Volschenk & Prendini have not accepted this decision and retained the family Urodacidae (and placed the new genus and species in this family because of its close relationship to Urodacus). The taxonomy of The Scorpion Files follows Soleglad & Fet (2005), and I have here listed Aops in Scorpionidae. This is formally not correct, but is the best choice in the current situation until a new revision on higher scorpion systematics is published. Volschenk ES, Prendini L. Aops oncodactylus, gen. et sp. nov, the frist troglobitic urodacid (Urodacidae: Scorpiones), with a re-assessment of cavernicolus, troglobitic and troglomorphic scorpions. Invertebrate Systematics. 2008;22:235. Cazierius granulosus Teruel, 2013 New sp. Cuba Teruel R. La subfamilia Diplocentrinae (Scorpiones: Scorpionidae) en Cuba. Sexta parte: Nueva especie de Cazierius Francke 1978. Revista Iberica de Arachnologia. 2013 (23):43-8. Cazierius neibae New sp. Dominican Republic Kovarik F, Teruel R. Three New Scorpion Species from the
Transcript of Taxonomical updates for Buthidae (2008 ) - NTNU...Diplocentrus poncei Francke & Quijano-Ravell, 2009...
2020 © Jan Ove Rein, The Scorpion Files
Taxonomical updates in The Scorpion Files for Scorpionidae (2008 →)
Taxa Status Distribution Comments Reference Aops New genus Australia The family Urodacidae was
abolished by Soleglad, Fet &
Kovarik in 2005 and the subfamily
Urodacinae was transfered to
Scorpionidae. Volschenk & Prendini
have not accepted this decision and
retained the family Urodacidae (and
placed the new genus and species in
this family because of its close
relationship to Urodacus). The
taxonomy of The Scorpion Files
follows Soleglad & Fet (2005), and I
have here listed Aops in
Scorpionidae. This is formally not
correct, but is the best choice in the
current situation until a new revision
on higher scorpion systematics is
published.
Volschenk ES, Prendini L. Aops
oncodactylus, gen. et sp. nov, the
frist troglobitic urodacid
(Urodacidae: Scorpiones), with a
re-assessment of cavernicolus,
troglobitic and troglomorphic
scorpions. Invertebrate
Systematics. 2008;22:235.
Cazierius granulosus Teruel,
2013
New sp. Cuba Teruel R. La subfamilia
Diplocentrinae (Scorpiones:
Scorpionidae) en Cuba. Sexta
parte: Nueva especie de
Cazierius Francke 1978. Revista
Iberica de Arachnologia. 2013
(23):43-8.
Cazierius neibae New sp. Dominican
Republic
Kovarik F, Teruel R. Three New
Scorpion Species from the
2020 © Jan Ove Rein, The Scorpion Files
Dominican Republic, Greater
Antilles (Scorpiones: Buthidae,
Scorpionidae). Euscorpius. 2014
(187):1-27.
Cazierius scaber (Pocock,
1893)
New comb. Jamaica New name: Heteronebo scaber
(Pocock, 1893)
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Cazierius tatae Santiago-
Blay, 2009
Synonymization
New sp.
Puerto Rico
Regarded as junior synonym of C.
garridoi Armas, 2005 and
synonymised.
de Armas LF. Acerca de algunos
alacranes (Scorpiones: Buthidae,
Scorpionidae) recientemente
descritos o registrados de Puerto
Rico e Isla Mona, Antillas
Mayores. Boletin Sociedad
Entomológica Aragonesa.
2009(45):298.
Santiago-Blay JA. Systematics
and some aspects of the biology
of the scorpions (Arachnida) of
the greater Puerto Rico region: A
biosystematic synopsis.
Entomological News. 2009 Jan-
Feb;120 (1):109-24.
Chersonesometrus Couzijn,
1978
Chersonesometrus bastawadei
Prendini & Loria, 2020
New status
New sp.
India
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
2020 © Jan Ove Rein, The Scorpion Files
Chersonesometrus hendersoni
Prendini & Loria, 2020
Chersonesometrus
nathanorum Prendini & Loria,
2020
Chersonesometrus
shivashankari Prendini &
Loria, 2020
Species transferred from
Heterometrus (new comb.):
Chersonesometrus
beccaloniae (Kovařik, 2004)
Chersonesometrus fulvipes
(C.L. Koch, 1837)
Chersonesometrus
madraspatensis (Pocock,
1900)
Chersonesometrus
pelekomanus (Couzijn, 1981)
Chersonesometrus tristis
(Henderson, 1919)
Chersonesometrus
wroughtoni (Pocock, 1899)
New sp.
New sp.
New sp.
India
India
India
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
2020 © Jan Ove Rein, The Scorpion Files
Cryptoiclus Teruel &
Kovarik, 2012
New genus Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions
of Cuba: Jakub Rolcik - Clarion
Production; 2012 (page 168-174)
Cryptoiclus rodriguezi Teruel
& Kovarik, 2012
New sp. Cuba Endemic to eastern Cuba. Teruel R, Kovarik F. Scorpions
of Cuba: Jakub Rolcik - Clarion
Production; 2012 (page 168-174)
Deccanometrus Prendini &
Loria, 2020
Deccanometrus obscurus
(Couzijn, 1981)
Species transferred from
Heterometrus (new comb.):
Deccanometrus bengalensis
(C.L. Koch, 1841)
Deccanometrus latimanus
(Pocock, 1894)
Deccanometrus liurus
(Pocock, 1897)
Deccanometrus phipsoni
(Pocock, 1893)
New genus
New status
Have had different statuses in the
past. See article.
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
2020 © Jan Ove Rein, The Scorpion Files
Deccanometrus ubicki
(Kovařik, 2004)
Deccanometrus xanthopus
(Pocock, 1897)
Didymocentus armasi Teruel
& Rodriguez, 2008
New sp. Cuba Teruel R, Rodriguez TM. La
subfamilia Diplocentrinae
(Scorpiones: Scorpionidae) en
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.
Boletin Sociedad Entomológica
Aragonesa. 2008(42):53-78
Didymocentus jaumei Armas,
1976
Restored
species
Species is valid and is redescribed
and updated as to their respective
diagnosis and geographical
distribution.
Teruel R, Rodriguez TM. La
subfamilia Diplocentrinae
(Scorpiones: Scorpionidae) en
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.
Boletin Sociedad Entomológica
Aragonesa. 2008(42):53-78
Didymocentus sanfelipensis
Armas, 1976
Restored
species
Species is valid and is redescribed
and updated as to their respective
diagnosis and geographical
distribution.
Teruel R, Rodriguez TM. La
subfamilia Diplocentrinae
(Scorpiones: Scorpionidae) en
Cuba. Quinta parte: el genero
Didymocentrus Kraepelin 1905.
Boletin Sociedad Entomológica
Aragonesa. 2008(42):53-78
Diplocentrus bicolor
Contreras-Felix & Santibanez-
Lopez, 2011
New sp. Mexico Contreras-Felix G, Santibanez-
Lopez CE. Diplocentrus bicolor
sp. n. (Scorpiones:
Diplocentridae) from Jalisco,
Mexico. Zootaxa.
2011(2992):61-8.
2020 © Jan Ove Rein, The Scorpion Files
Diplocentrus duende
Santibanez-Lopez &
Gonzalez-Santillan, 2017
New sp. Mexico Santibanez-Lopez CE,
Gonzalez-Santillan E. A new
species of Diplocentrus
(Scorpiones: Diplocentridae)
with punctate pedipalp surfaces,
a diagnostic character within the
"mexicanus" group. C R Biol.
2017; In Press.
Diplocentrus formosus Armas
& Martin-Frias, 2003
New status Reinstated as valid species.
Previously synonymized with
Diplocentrus tehuano Francke,
1977.
Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus franckei
Santobanez-Lopez, 2014
New sp. Mexico Santibanez-Lopez CE. A new
species of the genus
Diplocentrus Peters, 1861
(Scorpiones, Diplocentridae)
from Oaxaca, Mexico. Zookeys.
2014 (412):103-16.
Diplocentrus gladiator New sp. Mexico Species described in 1999, but I
didn’t learn about this species before
October 2008. Thanks to Oscar
Francke for informing me!
Beutelspacher CR, Trujillo
Olivera M. Una especie nueva de
Diplocentrus Peters
(Scorpionida: Diplocentridae) de
Chiapas, Mexico. Rev Nica Ent.
1999;50:1-11.
Diplocentrus insularis
Sagastume-Espinoza,
Longhorn & Santiban˜ez-
New sp. Honduras Sagastume-Espinoza KO,
Longhorn SJ, Santibanez-Lopez
CE. A new scorpion species of
2020 © Jan Ove Rein, The Scorpion Files
Lopez, 2015 genus Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae)
endemic to Islas de la Bahia,
Honduras. C R Biol. 2015 May
27.
Diplocentrus Izabal Armas &
Trujillo, 2016
New sp. Guatemala De Armas LF, Trujillo RE. A
new species of Diplocentrus
(Scorpionidae: Diplocentrinae)
from western Izabal, Guatemala.
Euscorpius. 2016(225):1-8.
Diplocentrus kraepelini
Santibanez-Lopez, Francke &
Prendini, 2013
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus lachua Armas.
Trujillo & Agreda, 2011
New sp. Guatemala de Armas LF, Trujillo RE,
Agreda EO. Nueva especie de
Diplocentrus Peters, 1851
(Scorpionidae: Diplocentrinae)
del noroeste de alta Verapaz,
Guatemala. Boletin de la SEA.
2011(49):113-7.
Diplocentrus landelinoi
Trujillo & Armas, 2012
New sp. Guatemala Trujillo RE, de Armas LF. Dos
especie nuevas de Diplocentrus
Peters, 1861 (Scorpionidae:
Diplocentrinae) de Guatemala.
Revista Iberica de Arachnologia.
2012 (21):131-8.
2020 © Jan Ove Rein, The Scorpion Files
Diplocentrus longimanus
Santibanez-Lopez, Francke &
Athanasiadis, 2011
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Athanasiadis MC. The
genus Diplocentrus Peters
(Scorpiones: Diplocentridae) in
Morelos, Mexico. Revista
Iberica de Arachnologia.
2011;19:3-13.
Diplocentrus majahuensis New sp. Mexico Species described in 2003, but I
didn’t learn about this species before
October 2008. Thanks to Oscar
Francke for informing me!
Baldazo Monsivaiz JG.
Diplocentrus majahuensis,
nueva especie de alcran del
estado de Guerrero, Mexico
(Scorpiones, Diplocentridae).
Entomologia Mexicana.
2003;2:73-7.
Diplocentrus motagua Armas
& Trujillo, 2009
New sp. Guatemala de Armas LF, Trujillo RE.
Nueva especie de Diplocentrus
Peters, 1861 (Scorpiones:
Scorpionidae) de Guatemala.
Boletin Sociedad Entomológica
Aragonesa. 2009(45):67-72.
Diplocentrus oxlajujbaktun
Trujillo & Armas, 2012
New sp. Guatemala Trujillo RE, de Armas LF. Dos
especie nuevas de Diplocentrus
Peters, 1861 (Scorpionidae:
Diplocentrinae) de Guatemala.
Revista Iberica de Arachnologia.
2012 (21):131-8.
Diplocentrus poncei Francke
& Quijano-Ravell, 2009
New comb.
New sp.
Mexico
See Kolotl below.
Authors using family Diplocentridae
Francke OF, Quijano-Ravell AF.
Una especie nueva Diplocentrus
(Scorpiones: Diplocentridae) del
estado de Michoacan, Mexico.
2020 © Jan Ove Rein, The Scorpion Files
Revista Mexicana de
Biodiversidad 2009;80:659-63.
Diplocentrus popti Trujillo &
Armas, 2016
New sp. Guatemala Trujillo RE, de Armas LF.
Nueva especie de Diplocentrus
peters, 1861 (Scorpiones:
Scorpionidae: Diplocentrinae)
del occidente de Guatemala.
Revista Iberica de Arachnologia.
2016(28):103-6.
Diplocentrus sagittipalpus
Santibanez-Lopez, Francke &
Prendini, 2013
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus sissomi
Santibanez-Lopez, Francke &
Prendini, 2013
New sp. Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Systematics of
the keyserlingii group of
Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae),
with descriptions of three new
species from Oaxaca, Mexico.
American Museum Novitates.
2013 (3777):1-47.
Diplocentrus tenango New sp. Mexico Santibanez-Lopez CE, Francke
OF. A new species of
Diplocentrus (Arachnida:
Scorpiones) from Oaxaca,
Mexico. Zootaxa.
2020 © Jan Ove Rein, The Scorpion Files
2008(1742):53-60
Gigantometrus Couzijn, 1978
Gigantometrus titanicus
(Couzijn, 1981)
Species transferred from
Heterometrus (new comb.):
Gigantometrus swammerdami
(Simon, 1872)
New status
New status
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus atrascorpius
Mirza, Joshi, Desouza &
Sanap, 2012
Synonymization
New sp.
India
Synonymized with
Chersonesometrus beccaloniae
(Kovařik, 2004).
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Mirza ZA, Joshi D, Desouza G,
2020 © Jan Ove Rein, The Scorpion Files
Sanap RV. Description of a new
species of scorpion of the genus
Heterometrus Ehrenberg, 1828
(Scorpiones: Scorpionidae) from
the Western Ghats, India. Indian
Journal of Arachnology.
2012;1(2):1-8.
Heterometrus cimrmani
Kovarik, 2004
Synonymization Synonymized with Heterometrus
laevigatus (Thorell, 1876)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus glaucus
(Thorell, 1876)
New status Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus granulomanus
Couzijn, 1981
Synonymization Synonymized with Srilankametrus
caesar (C.L. Koch, 1841)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
2020 © Jan Ove Rein, The Scorpion Files
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus keralensis
Tikader & Bastawade, 1983
Synonymization Synonymized with Sahyadrimetrus
rugosus (Couzijn, 1983)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus laevifrons
Roewer, 1943
Synonymization Synonymized with Heterometrus
glaucus (Thorell, 1876)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus laevigatus
(Thorell, 1876)
New status Prendini L, Loria SF. Systematic
revision of the Asian Forest
2020 © Jan Ove Rein, The Scorpion Files
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus liangi Zhu &
Yang, 2007
Synonymization
New sp.
Vietnam
Synonymized with Heterometrus
Silenus (Simon, 1884).
Haven’t seen this paper (in Chinese).
Info based on bibliographical data
only.
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Zhu MS, Yang X-F. Two species
of the genus Heterometrus
Ehrenberg, 1828 (Scorpionidae)
from South Vietnam sold in pet
shops in China. Acta
Arachnologica Sinica. 2007;16
(2):92-103.
Heterometrus minotaurus
Pliskova, Kovarik, Kosulic &
Stahlavsky, 2016
Synonymization
Synonymized with Heterometrus
laevigatus (Thorell, 1876)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
2020 © Jan Ove Rein, The Scorpion Files
New sp.
Thailand
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480
Pliskova J, Kovarik F, Kosulic
O, Stahlavsky F. Description of
a new species of Heterometrus
Ehrenberg, 1828 (Scorpiones:
Scorpionidae) from Thailand
with remarks about the
utilization of cytogenetic data in
taxonomy of the genus. Annales
Zoologici. 2016;66(3):467-76.
Heterometrus mysorensis
Kovařik, 2004
Synonymization Synonymized with
Chersonesometrus tristis
(Henderson, 1919)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus nepalensis
Kovařik, 2004
Synonymization Synonymized with Deccanometrus
bengalensis (C. L. Koch, 1841)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
2020 © Jan Ove Rein, The Scorpion Files
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus rolciki Kovarik,
2004
Synonymization Synonymized with Sahyadrimetrus
scaber (Thorell, 1876)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus sejnai Kovařik,
2004
Synonymization Synonymized with Javanimetrus
cyaneus (C. L. Koch, 1836)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus serratus
(Pocock, 1900)
New status Restored from synonymy with H.
indus (Geer, 1778).
Kovarik F, Lowe G, Ranawana
KB, Hoferek D, Jayarathne
VAS, Pliskova J, et al. Scorpions
2020 © Jan Ove Rein, The Scorpion Files
of Sri Lanka (Arachnida,
Scorpiones: Buthidae,
Chaerilidae, Scorpionidae) with
Description of Four New Species
of the Genera Charmus Karsch,
1879 and Reddyanus Vachon,
1972, stat. n. Euscorpius. 2016
(220):1-133.
Heterometrus silenus (Simon,
1884)
New status Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus telanganaensis
Javed, Mirza, Tampal &
Lourenco, 2010
Synonymization
New sp.
India
Synonymized with Deccanometrus
xanthopus (Pocock, 1897)
Kovarik F, Lowe G, Ranawana
KB, Hoferek D, Jayarathne
VAS, Pliskova J, et al. Scorpions
of Sri Lanka (Arachnida,
Scorpiones: Buthidae,
Chaerilidae, Scorpionidae) with
Description of Four New Species
of the Genera Charmus Karsch,
1879 and Reddyanus Vachon,
1972, stat. n. Euscorpius. 2016
(220):1-133.
Maqsood Javed SM, Mirza ZA,
2020 © Jan Ove Rein, The Scorpion Files
Tampal F, Lourenco WR. A new
species of the genus
Heterometrus Ehrenberg, 1828
(Scorpiones: Scorpionidae) from
India with notes on its natural
history. Boletin de la SEA.
2010(47):143-8.
Heterometrus tibetanus
Lourenco et al, 2005
Synonymization Synonymized with Deccanometrus
bengalensis (C. L. Koch, 1841)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Heterometrus yaleensis
Kovarik, Ranawana,
Jayarathne, Hoferek &
Stahlavsky, 2019
New sp. Sri Lanka Kovarik F, Ranawana KB,
Jayarathne VAS, Hoferek D,
Stahlavsky F. Scorpions of Sri
Lanka (Arachnida: Scorpiones).
Part III. Heterometrus yaleensis
sp. n. (Scorpionidae).
Euscorpius. 2019(283):1-13.
Heteronebo barahonae
Teruel, de Armas & Kovarik,
2015
New sp. Hispaniola,
Dominican
Republic
Teruel R, de Armas LF, Kovarik
F. Two new species of scorpions
(Scorpiones: Buthidae,
Scorpionidae) from Dominican
Republic, Greater Antilles.
Revista Iberica de Arachnologia.
2015 (27):13-33.
2020 © Jan Ove Rein, The Scorpion Files
Heteronebo franckei
Stockwell, 1985
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Heteronebo jamaicae
jamaicae Francke, 1978
New status
Synonymization
Revalidated as species.
Synonymized with Heteronebo
scaber (Pocock, 1893)
Santibanez-Lopez CE, Francke
OF, Prendini L. Phylogeny of
the North American scorpion
genus Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae)
based on morphology, nuclear
and mitochondrial DNA.
Arthropod Systematics &
Phylogeny. 2014;72(3):257-79.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Heteronebo jamaicae
occidentalis Francke, 1978
New status
Revalidated as species.
Santibanez-Lopez CE, Francke
OF, Prendini L. Phylogeny of
the North American scorpion
genus Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae)
based on morphology, nuclear
and mitochondrial DNA.
Arthropod Systematics &
Phylogeny. 2014;72(3):257-79.
2020 © Jan Ove Rein, The Scorpion Files
Synonymization Synonymized with Heteronebo
scaber (Pocock, 1893)
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Heteronebo jamaicae
portlandensis Francke, 1978
New status
Synonymization
Revalidated as species.
Synonymized with Heteronebo
scaber (Pocock, 1893)
Santibanez-Lopez CE, Francke
OF, Prendini L. Phylogeny of
the North American scorpion
genus Diplocentrus Peters, 1861
(Scorpiones: Diplocentridae)
based on morphology, nuclear
and mitochondrial DNA.
Arthropod Systematics &
Phylogeny. 2014;72(3):257-79.
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Heteronebo morenoi (Armas,
1973)
New status Cuba Raised from subspecies status.
Previously Heteronebo bermudezi
morenoi (Armas, 1973).
Teruel R, Rodriguez-Cabrera
TM. The Missing Piece of the
Puzzle Solved: Heteronebo
Pocock, 1899 (Scorpiones:
Scorpionidae) Occurs at Isla de
Pinos, Cuba. Euscorpius.
2017(240):1-4.
Heteronebo scaber (Pocock,
1893)
New comb. Jamaica Previous name: Cazierius scaber
(Pocock, 1893) + senior synonym of
Heteronebo franckei Stockwell,
Teruel R. Los escorpiones
Diplocentrinos de Jamaica
(Scorpiones: Scorpionidae:
2020 © Jan Ove Rein, The Scorpion Files
1985, H. jamaicae jamaicae
Francke, 1978, H. jamaicae
occidentalis Francke, 1978 & H.
jamaicae portlandensis Francke,
1978
Diplocentronae). Boletin
Sociedad Entomológica
Aragonesa. 2009; (44):103-10.
Javanimetrus Couzijn, 1981
Species transferred from
Heterometrus (new comb.):
Javanimetrus cyaneus (C.L.
Koch, 1836)
New status Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Kolotl Santibanez-Lopez,
Francke & Prendini, 2014
New genus Western Mexico Santibanez-Lopez CE, Francke
OF, Prendini L. Kolotl, n. gen.
(Scorpiones: Diplocentridae), a
New Scorpion Genus from
Mexico. American Museum
Novitates. 2014 (3815):1-28.
Kolotl magnus (Beutelspacher
& López-Forment, 1991)
New comb. Previous name: Diplocentrus
magnus Beutelspacher & López-
Forment, 1991.
Santibanez-Lopez CE, Francke
OF, Prendini L. Kolotl, n. gen.
(Scorpiones: Diplocentridae), a
New Scorpion Genus from
2020 © Jan Ove Rein, The Scorpion Files
Mexico. American Museum
Novitates. 2014 (3815):1-28.
Kolotl poncei (Francke &
Quijano-Ravell, 2009)
New comb. Previous name: Diplocentrus poncei
Francke & Quijano-Ravell, 2009.
Santibanez-Lopez CE, Francke
OF, Prendini L. Kolotl, n. gen.
(Scorpiones: Diplocentridae), a
New Scorpion Genus from
Mexico. American Museum
Novitates. 2014 (3815):1-28.
Oiclus ardens Ythier, 2019 New sp. Guadeloupe:
BasseTerre
Ythier E. On the genus Oiclus
Simon, 1880 (Scorpiones:
Diplocentridae) in Guadeloupe
islands, with description of three
new species. Arachnida - Rivista
Aracnologica Italiana.
2019;5(12):17-49.
Oiclus cousteaui Ythier, 2019 New sp. Guadeloupe: Ilets
Pigeon
Ythier E. On the genus Oiclus
Simon, 1880 (Scorpiones:
Diplocentridae) in Guadeloupe
islands, with description of three
new species. Arachnida - Rivista
Aracnologica Italiana.
2019;5(12):17-49.
Oiclus tipunch Ythier, 2019 New sp. Guadeloupe: Iles
des Saintes
Ythier E. On the genus Oiclus
Simon, 1880 (Scorpiones:
Diplocentridae) in Guadeloupe
islands, with description of three
new species. Arachnida - Rivista
Aracnologica Italiana.
2019;5(12):17-49.
Pandiborellius Rossi, 2015 New status Previously Pandinurus
(Pandiborellius) Rossi, 2015a.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
2020 © Jan Ove Rein, The Scorpion Files
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandiborellius arabicus
(Kraepelin, 1894)
Pandinurus arabicus
(Kraepelin, 1894)
New comb.
New comb.
Previous Pandinurus arabicus
(Kraepelin, 1894)
Previous Pandinus arabicus
(Kraepelin, 1894).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandiborellius awashensis
(Kovarik, 2012)
New comb.
Previous Pandinurus awashensis
(Kovarik, 2012)
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
2020 © Jan Ove Rein, The Scorpion Files
Pandinurus awashensis
(Kovarik, 2012)
New comb.
New sp.
Ethiopia
Previous Pandinus awashensis
(Kovarik, 2012).
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
Pandiborellius igdu Kovarik,
Lowe, Soleglad & Pliskova,
2017
New sp. Ethiopia Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
2020 © Jan Ove Rein, The Scorpion Files
Taxonomy. Euscorpius.
2017(238):1-103.
Pandiborellius insularis
Kovarik, Lowe, Soleglad &
Pliskova, 2017
New sp. Eritrea Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandiborellius lanzai (Rossi,
2015)
Pandinurus lanzai Rossi,
2015
New comb.
New sp.
Somalia
Previously Pandinurus lanzai Rossi,
2015.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A. Due ulteriori nuove
specie del sottogenere
Pandiborellius Rossi, 2015 dal
Corno d'Africa (Scorpiones,
Scorpionidae). Arachnida -
Rivista Aracnologica Italiana.
2015;1(2):2-36.
2020 © Jan Ove Rein, The Scorpion Files
Pandiborellius magrettii
(Borelli, 1901)
Pandinurus magrettii (Borelli,
1901)
New comb.
New comb.
Previous Pandinurus magrettii
(Borelli, 1901)
Previous Pandinus magrettii
(Borelli, 1901).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandiborellius meidensis
(Karsch, 1879)
New comb.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
2020 © Jan Ove Rein, The Scorpion Files
Pandinurus meidensis
(Karsch, 1879)
New comb Previous Pandinus meidensis
(Karsch, 1879).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandiborellius nistriae
(Rossi, 2014)
Pandinurus nistriae (Rossi,
2014)
New comb.
New comb.
New sp.
Djibouti
Previous Pandinurus nistriae (Rossi,
2014)
Previous Pandinus nistriae (Rossi,
2014).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Rossi A. Notes on the
distribution of Pandinus
(Pandinus) Thorell, 1876 and
2020 © Jan Ove Rein, The Scorpion Files
Pandinus (Pandinurus) Fet, 1997
with the descriptions of two new
species from Central African
Republic. Onychium. 2014;10
(2013):10-31.
Pandiborellius percivali
(Pocock, 1902)
Pandinurus percivali (Pocock,
1902)
New comb.
New comb.
Previous Pandinurus percivali
(Pocock, 1902)
Previous Pandinus percivali
(Pocock, 1902).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandiborellius somalilandus
(Kovarik, 2013)
New comb.
Previous New comb.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
2020 © Jan Ove Rein, The Scorpion Files
Pandinurus somalilandus
(Kovarik, 2013)
New comb.
New sp.
Somaliland
Previous Pandinus somalilandus
(Kovarik, 2012).
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
Pandinoides Fet, 1997 New status Raised to genus from subgenus
status within genus Pandinus
Thorell, 1876
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinoides cavimanus
(Pocock, 1888)
New comb. Previous Pandinus cavimanus
(Pocock, 1888).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
2020 © Jan Ove Rein, The Scorpion Files
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinoides duffmackayi
Prendini, 2016
New sp. Kenya and
Tanzania
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Pandinoides militaris Pocock,
1900
Pandinus militaris Pocock,
1900
New status/
New comb.
Synonymization
Revalidated and redescribed.
This taxa was judged as a junior
synonym of Pandinoides cavimanus
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Kovarik F. A checklist of
scorpions (Arachnida) in the
collection of the
Forschungsinstitut und
Naturmuseum Senckenberg,
Frankfurt am Main, Germany.
Serket. 2002 April;8 (1):1-23.
Pandinops Birula, 1913 New status Raised to genus from subgenus
status within genus Pandinus
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
2020 © Jan Ove Rein, The Scorpion Files
Thorell, 1876 revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops bellicosus (L.
Koch, 1875)
New comb. Previous Pandinus bellicosus (L.
Koch, 1875).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinus boschisi
(Caporiacco, 1937)
New comb. Previously placed in Pandinus with
a nomen dubium status.
Rossi A. Una revisione
preliminare del genere Pandinus
Thorell, 1876 con la descrizione
di un nuovo sottogenere e due
nuove specie dall'Etiopia
(Scorpiones, Scorpionisae).
Arachnida - Rivista
Aracnologica Italiana.
2015;1(3):37-52.
Pandinops colei (Pocock,
1896)
New comb. Previous Pandinus colei (Pocock,
1896).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops eritreaensis New comb. Previous Pandinus eriatreaensis Rossi A: Sui sottogeneri di
2020 © Jan Ove Rein, The Scorpion Files
(Kovarik, 2003) Kovarik, 2003. Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops friedrichi Kovarik,
2016
New sp. Somalia Kovarik F. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part VIII.
Pandinops Birula, 1913
(Scorpionidae), With
Description of Two New
Species. Euscorpius.
2016(229):1-20.
Pandinops hawkeri (Pocock,
1900)
Synonymization
New comb.
Synonymized with Pandinops
pugilator (Pocock, 1900).
Previous Pandinus hawkeri (Pocock,
1900).
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XIII. Review of Pandinops
hawkeri, P. peeli, P.platycheles,
and P. pugilator (Scorpionidae).
Euscorpius. 2017(254):1-20.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops peeli (Pocock, Synonymization Synonymized with Pandinops Kovarik F, Lowe G, Elmi HSA.
2020 © Jan Ove Rein, The Scorpion Files
1900)
New comb.
pugilator (Pocock, 1900).
Previous Pandinus peeli (Pocock,
1900).
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XIII. Review of Pandinops
hawkeri, P. peeli, P.platycheles,
and P. pugilator (Scorpionidae).
Euscorpius. 2017(254):1-20.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops platycheles
(Werner, 1916)
New comb.
New comb.
Previous Pandinurus platycheles
(Werner, 1916)
Previous Pandinoides platycheles
(Werner, 1916)
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XIII. Review of Pandinops
hawkeri, P. peeli, P.platycheles,
and P. pugilator (Scorpionidae).
Euscorpius. 2017(254):1-20
Kovarik F, Lowe G, Pliskova J,
Stahlavsky F. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part VII.
Parabuthus Pocock, 1890
(Buthidae), with Description of
P. hamar sp. n. and P. kajibu sp.
n. from Ethiopia. Euscorpius.
2016(228):1-58.
2020 © Jan Ove Rein, The Scorpion Files
New comb.
Previous Pandinus platycheles
(Werner, 1916).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops pococki (Kovarik,
2000)
New comb. Previous Pandinus pococki Kovarik,
2000
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinops pugilator (Pocock,
1900)
New status Previously synonymized with
Pandinops hawkeri (Pocock, 1900).
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XIII. Review of Pandinops
hawkeri, P. peeli, P.platycheles,
and P. pugilator (Scorpionidae).
Euscorpius. 2017(254):1-20.
Pandinops turieli Kovarik,
2016
New sp. Ethiopia and
Kenya
Kovarik F. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part VIII.
Pandinops Birula, 1913
(Scorpionidae), With
Description of Two New
Species. Euscorpius.
2016(229):1-20.
2020 © Jan Ove Rein, The Scorpion Files
Pandinopsis Vachon, 1974 New status
New status
Confirmed genus status.
Raised to genus from subgenus
status within genus Pandinus
Thorell, 1876
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinopsis dictator (Pocock,
1888)
New comb.
Transferred to new genus.
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
2020 © Jan Ove Rein, The Scorpion Files
Pandinopsis dictator (Pocock,
1888)
New comb.
New comb.
Previous Pandinopsis dictator
(Pocock, 1888). Pandinopsis is
subgenus.
Previous Pandinus dictator (Pocock,
1888).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus Fet, 1997 New status Raised to genus from subgenus
status within genus Pandinus
Thorell, 1876
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus afar Kovarik,
Lowe, Soleglad & Pliskova,
2017
New sp. Ethiopia Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
2020 © Jan Ove Rein, The Scorpion Files
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus awalei Kovarik,
Lowe & Elmi, 2020
New sp. Somaliland Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XXV. Description of Pandinurus
awalei sp. n. and the male of
Pandiborellius somalilandus
(Kovařík, 2012), with remarks
on recent synonymies
(Scorpionidae: Pandininae).
Euscorpius. 2020(322):1-21.
Pandinurus bartolozzii Rossi,
2015
Synonymization
New sp.
Democratic
Republic of the
Congo
Synonymized with Pandinus
viatoris (Pocock, 1890)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
2020 © Jan Ove Rein, The Scorpion Files
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus cianferonii Rossi,
2015
Synonymization
New sp.
Somalia
Synonymized with P. pallidus
(Kraepelin, 1894)
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A. Ulteriori commenti
sulla tassonomia dei generi
Pandinus e Pandinurus, con la
definizione di un nuovo
sottogenere ed una nuova specie
di Pandinurus dalla Somalia.
Arachnida - Rivista
Aracnologica Italiana.
2015;1(4):41-55.
Pandinurus citernii (Borelli,
1919)
New status Restored from synonymy with P.
phillipsii (Pocock, 1896).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
2020 © Jan Ove Rein, The Scorpion Files
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus exitialis (Pocock,
1888)
New comb. Previous Pandinus exitialis (Pocock,
1888).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus flagellicauda
Rossi, 2015
Synonymization
New sp.
Democratic
Republic of the
Congo
Synonymized with Pandinus
viatoris (Pocock, 1890)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus fulvipes Kovarik,
Lowe & Mazuch, 2019
New sp. Somaliland Kovarik F, Lowe G, Mazuch T.
Scorpions of the Horn of Africa
2020 © Jan Ove Rein, The Scorpion Files
(Arachnida: Scorpiones). Part
XIX. Pandiborellius meidensis
(Karsch, 1879) and Pandinurus
fulvipes sp. n. (Scorpionidae)
from Somaliland. Euscorpius.
2019(275):1-18.
Pandinurus gregoryi (Pocock,
1896)
New comb.
New status
Previous Pandinus gregoryi
(Pocock, 1888).
Considered a synonym of Pandinus
exitialis (Pocock, 1888) by Kovarik,
2003, but restored species status in
Kovarik’s (2009) Illustrated catalog
of scorpions (page 55).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Kovarik F. Illustrated catalog of
scorpions. Part I. Introductory
remarks; keys to families and
genera; subfamily Scorpioninae
with illustrated keys to
Heterometrus and Pandinus
species. Prague: Clarion
Production; 2009.
Pandinurus hangarale
Kovarik, Lowe, Mazuch,
Awale, Stundlowa &
Stahlavsky, 2017
New sp. Somaliland Kovarik F, Lowe G, Mazuch T,
Awale AI, Stundlova J,
Stahlavsky F. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part XII.
Pandinurus hangarale sp. n.
(Scorpionidae) from Somaliland
and a Review of Type Locality
and True Distribution of
2020 © Jan Ove Rein, The Scorpion Files
Pandinurus smithi (Pocock,
1897). Euscorpius. 2017(253):1-
18.
Pandinurus intermedius
(Borelli, 1919)
Pandinurus intermedius
(Borelli, 1919)
New status
Synonymization
loweiNew
status
Restored to species status.
Syn. with Pandinurus citernii
(Borelli, 1919)
Restored from synonymy with P.
phillipsii (Pocock, 1896).
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XXV. Description of Pandinurus
awalei sp. n. and the male of
Pandiborellius somalilandus
(Kovařík, 2012), with remarks
on recent synonymies
(Scorpionidae: Pandininae).
Euscorpius. 2020(322):1-21.
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
2020 © Jan Ove Rein, The Scorpion Files
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus janae Rossi, 2015 Synonymization
New sp.
Yemen
Synonymized with Pandinus
percivali (Pocock, 1902).
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus kmoniceki
Kovarik, Lowe, Mazuxh,
Pliskova & Stahlavsky, 2017
New sp. Somaliland Kovarik F, Lowe G, Mazuch T,
Pliskova J, Stahlavsky F.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XI. Pandinurus kmoniceki sp. n.
(Scorpionidae) from Somaliland.
Euscorpius. 2017(243):1-14.
Pandinurus lorenzoi Rossi,
2015
Synonymization
Synonymized with Pandinus
viatoris (Pocock, 1890)
Prendini L. Redefinition and
systematic revision of the East
2020 © Jan Ove Rein, The Scorpion Files
New sp.
Tanzania
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus mazuchi
(Kovarik, 2011)
Pandinus mazuchi Kovarik,
2011
New comb.
New sp.
Ethiopia
Previously Pandinus mazuchi
Kovarik, 2011.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Kovarik F. A review of the
subgenus Pandinus Thorell, 1876
with descriptions of two new
species from Uganda and
2020 © Jan Ove Rein, The Scorpion Files
Ethiopia (Scorpiones:
Scorpionidae). Euscorpius.
2011(129):1-18.
Pandinurus oromo Kovarik,
Lowe, Soleglad & Pliskova,
2017
New sp. Ethiopia Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus pallidus
(Kraepelin, 1894)
New comb. Previous Pandinus kraepelin
(Pocock, 1894).
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus pantinii Rossi,
2015
Synonymization
Synonymized with Pandinus
viatoris (Pocock, 1890)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
2020 © Jan Ove Rein, The Scorpion Files
New sp. Malawi Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus phillipsii (Pocock,
1896)
New status Restored from synonymy. Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus prendinii Rossi,
2015
Synonymization
New sp.
South Africa
Synonymized with Pandinus
sudanicus (Hirst, 1911).
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
2020 © Jan Ove Rein, The Scorpion Files
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus pygmaeus Rossi,
2015
Synonymization
New sp.
Democratic
Republic of
Congo
Synonymized with Pandinus
viatoris (Pocock, 1890)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinurus sabbadinii Rossi,
2015
Synonymization
Synonymized with Pandiborellius
magrettii (Borelli, 1901)
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
2020 © Jan Ove Rein, The Scorpion Files
New sp.
Eritrea
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A. Due ulteriori nuove
specie del sottogenere
Pandiborellius Rossi, 2015 dal
Corno d'Africa (Scorpiones,
Scorpionidae). Arachnida -
Rivista Aracnologica Italiana.
2015;1(2):2-36.
Pandinurus smithi (Pocock,
1897)
New comb. Previously Pandinus smithi (Pocock,
1987).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Pandinurus sudanicus (Hirst,
1911)
New comb.
New status
Sudan
Previous Pandinus sudanicus (Hirst,
1911).
Previous status:
Pandinus exitialis sudanicus Hirst,
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
2020 © Jan Ove Rein, The Scorpion Files
1911 (in part)
Pandinus magretti Birula, 1928 (in
part).
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
Pandinurus trailini (Kovarik,
2013)
Pandinus trailini Kovarik,
2013
New comb.
New sp.
Ethiopia
Previously Pandinus trailini
Kovarik, 2013.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Kovarik F. Pandinus (Pandinus)
trailini sp. n. from Ethiopia
(Scorpiones: Scorpionidae), with
data on localities and life
strategy. Euscorpius. 2013
(163):1-14.
Pandinurus vachoni (Rossi,
2014)
Synonymization
Synonymized with P. sudanicus
(Hirst, 1911).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2020 © Jan Ove Rein, The Scorpion Files
New comb.
New sp.
Chad
Previous Pandinus vachoni (Rossi,
2014)
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Rossi A. The fragmented peri-
Saharan distribution of the
subgenus Pandinurus Fet, 1997
with the description of a new
species from Chad (Scorpiones,
Scorpionidae, Pandinus). Serket.
2014;14(1):6-14.
Pandinus bottegoi Rossi, 2015 Synonymization
New sp.
Ethiopia
Synonymized with P. platycheles
(Werner, 1916).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A. Una revisione
preliminare del genere Pandinus
2020 © Jan Ove Rein, The Scorpion Files
Thorell, 1876 con la descrizione
di un nuovo sottogenere e due
nuove specie dall'Etiopia
(Scorpiones, Scorpionisae).
Arachnida - Rivista
Aracnologica Italiana.
2015;1(3):37-52.
Pandinus camerounensis
Lourenco, 2014
Synonymization
New sp.
Cameroon
Synonymized with Pandinus
imperator (C. L. Koch, 1841)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Lourenco WR. Further
considerations on the identity
and distribution of Pandinus
imperator (C. L. Koch, 1841)
and description of a new species
from Cameroon (Scorpiones:
Scorpionidae). entomologische
Mitteilungen aus dem
Zoologischen Museum
Hamburg. 2014;17(192):139-51.
Pandinus platycheles
(Werner, 1916)
New comb.
See below for previous status.
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
2020 © Jan Ove Rein, The Scorpion Files
Pandinus platycheles
(Werner, 1916)
Pandinus platycheles
(Werner, 1916)
New comb.
New comb.
Previous Pandinoides platycheles
(Werner, 1916)
Previous Pandinus platycheles
(Werner, 1916).
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
Kovarik F, Lowe G, Pliskova J,
Stahlavsky F. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part VII.
Parabuthus Pocock, 1890
(Buthidae), with Description of
P. hamar sp. n. and P. kajibu sp.
n. from Ethiopia. Euscorpius.
2016(228):1-58.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Pandinus riccardoi Rossi,
2015
New status
Seems to be restored to species
status and moved to Pandinurus. See
p. 9.
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XIII. Review of Pandinops
hawkeri, P. peeli, P.platycheles,
and P. pugilator (Scorpionidae).
Euscorpius. 2017(254):1-20.
2020 © Jan Ove Rein, The Scorpion Files
Synonymization
New sp.
Ethiopia
Synonymized with P. platycheles
(Werner, 1916).
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A. Una revisione
preliminare del genere Pandinus
Thorell, 1876 con la descrizione
di un nuovo sottogenere e due
nuove specie dall'Etiopia
(Scorpiones, Scorpionisae).
Arachnida - Rivista
Aracnologica Italiana.
2015;1(3):37-52.
Pandinus roeseli (Simon,
1872)
Synonymization
Synonymized with Pandinus
imperator (C. L. Koch, 1841)
Prendini L. Redefinition and
systematic revision of the East
African scorpion genus
Pandinoides (Scorpiones:
Scorpionidae) with critique of
the taxonomy of Pandinus, sensu
lato. Bulletin of the American
Museum of Natural History.
2016(407):1-67.
2020 © Jan Ove Rein, The Scorpion Files
New status Guinea
Restored from synonymization with
Pandinus imperator (Koch, 1841)
Lourenco WR. Further
considerations on the identity
and distribution of Pandinus
imperator (C. L. Koch, 1841)
and description of a new species
from Cameroon (Scorpiones:
Scorpionidae). entomologische
Mitteilungen aus dem
Zoologischen Museum
Hamburg. 2014;17(192):139-51.
Pandinus ugandaensis
Kovarik, 2011
New sp. Uganda Kovarik F. A review of the
subgenus Pandinus Thorell, 1876
with descriptions of two new
species from Uganda and
Ethiopia (Scorpiones:
Scorpionidae). Euscorpius.
2011(129):1-18.
Pandinus ulderigoi Rossi,
2014
New sp. Central African
Republic
Rossi A. Notes on the
distribution of Pandinus
(Pandinus) Thorell, 1876 and
Pandinus (Pandinurus) Fet, 1997
with the descriptions of two new
species from Central African
Republic. Onychium. 2014;10
(2013):10-31.
Pandipalpus Rossi, 2015 New status Restored from synonymy. Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
2020 © Jan Ove Rein, The Scorpion Files
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Pandipalpus lowei Kovarik,
2012
Pandinus lowei Kovarik, 2012
Pandinurus lowei (Kovarik,
2012)
New status
Synonymization
New comb.
Democratic
Republic of
Congo
Transferred to Pandipalpus.
Restored to species status.
Syn. with Pandipalpus viatoris
(Pocock, 1890)
Previous Pandinus lowei (Kovarik,
2012).
Kovarik F, Lowe G, Elmi HSA.
Scorpions of the Horn of Africa
(Arachnida: Scorpiones). Part
XXV. Description of Pandinurus
awalei sp. n. and the male of
Pandiborellius somalilandus
(Kovařík, 2012), with remarks
on recent synonymies
(Scorpionidae: Pandininae).
Euscorpius. 2020(322):1-21.
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
2020 © Jan Ove Rein, The Scorpion Files
New comb.
New sp.
Previously Pandinurus lowei
(Kovarik, 2012)
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Kovarik F. Review of the
subgenus Pandinurus Fet, 1997
with descriptions of three new
species (Scorpiones,
Scorpionidae, Pandinus).
Euscorpius. 2012(141):1-22.
Pandipalpus viatoris (Pocock,
1890)
New comb.
Previous status Pandinus viatoris
(Pocock, 1890)
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
2020 © Jan Ove Rein, The Scorpion Files
Pandinus viatoris (Pocock,
1890)
Pandinurus viatoris (Pocock,
1890)
New comb.
New comb.
Previous Pandinus viatoris (Pocock,
1890).
Previous Pandinurus viatoris
(Pocock, 1890).
480.
Kovarik F, Lowe G, Soleglad
ME, Pliskova J. Scorpions of the
Horn of Africa (Arachnida:
Scorpiones). Part X.
Pandiborellius stat. n. and
Pandinurus (Scorpionidae) with
Description of Four New Species
from Eritrea and Ethiopia, and
Review of Pandinus Sensu Lato
Taxonomy. Euscorpius.
2017(238):1-103.
Rossi A: Sui sottogeneri di
Pandinus Thorell, 1876 con
revisione del genere Pandinurus
Fet, 1997 stat. n. e descrizione di
sette nuove specie e tre nuovi
sottogeneri (Scorpiones:
Scorpionidae). Onychium 2015;
11:10-66
Oiclus nanus Teruel &
Chazal, 2010
New Sp. Guadeloupe,
Lesser Antilles
Teruel R, Chazal L. A new
species of the genus Oiclus
Simon, 1880 (Scorpiones:
Scorpionidae: Diplocentrinae)
from Guadeloupe, Lesser
Antilles. Euscorpius. 2010
(92):1-9.
Oiclus questeli Teruel, 2008 New Sp. Lesser Antilles Teruel R. A new species of
Oiclus Simon 1880 (Scorpiones:
Scorpionidae: Diplocentrinae)
2020 © Jan Ove Rein, The Scorpion Files
from Saint-Barthelemy, Lesser
Antilles. Boletin Sociedad
Entomológica Aragonesa.
2008(43):95-9.
Sahyaddrimetrus Prendini &
Loria, 2020
Sahyadrimetrus mathewi
Prendini & Loria, 2020
Sahyadrimetrus rugosus
(Couzijn, 1981)
Sahyadrimetrus tikaderi
Prendini & Loria, 2020
Species transferred from
Heterometrus (New
combinations):
Sahyadrimetrus barberi
(Pocock, 1900)
Sahyadrimetrus kanarensis
(Pocock, 1900)
Sahyadrimetrus scaber
(Thorell, 1876)
New genus
New sp.
New status
New sp.
India
India
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
Scorpio birulai Fet, 1997 New status Morocco Previous subspecies status: Scorpio
maurus subtypicus Birula, 1910
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
2020 © Jan Ove Rein, The Scorpion Files
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio ennedi Lourenco,
Duhem & Cloudsley-
Thompson, 2012
New sp. Chad Lourenco WR, Duhem B,
Cloudsley Thompson JL.
Scorpions from Ennedi, Kapka
and Tibesti, the mountains of
Chad, with descriptions of nine
new species (Scorpiones:
Buthidae, Scorpionidae).
Arthropoda Selecta.
2012;21(4):307-38.
Scorpio fuliginosus Pallary,
1928
New status Morocco Previous status: Heterometrus
fuliginosus Pallary, 1928
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio fuscus (Ehrenberg,
1829)
New status Israel, Palestina Previous subspecies status: Scorpio
maurus fuscus Ehrenberg, 1829
Talal S, Tesler I, Sivan J, Ben-
Shlomo R, Muhammad Tahir H,
Prendini L, et al. Scorpion
speciation in the Holy Land:
Multilocus phylogeography
corroborates diagnostic
differences in morphology and
burrowing behavior among
Scorpio subspecies and justifies
recognition as phylogenetic,
ecological and biological
2020 © Jan Ove Rein, The Scorpion Files
species. Mol Phylogenet Evol.
2015 May 15.
Scorpio hesperus Birula, 1910 New status Morocco Previous subspecies status: Scorpio
maurus hesperus Birula, 1910
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio kruglovi (Ehrenberg,
1829)
New status Israel, Jordan,
Syria
Previous subspecies status: Scorpio
maurus kruglovi Ehrenberg, 1829
Talal S, Tesler I, Sivan J, Ben-
Shlomo R, Muhammad Tahir H,
Prendini L, et al. Scorpion
speciation in the Holy Land:
Multilocus phylogeography
corroborates diagnostic
differences in morphology and
burrowing behavior among
Scorpio subspecies and justifies
recognition as phylogenetic,
ecological and biological
species. Mol Phylogenet Evol.
2015 May 15.
Scorpio mogadorensis Birula,
1910
New status Morocco Previous subspecies status: Scorpio
maurus mogadorensis Birula, 1910
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio niger Lourenco & New sp. Niger Lourenco WR, Cloudsley
2020 © Jan Ove Rein, The Scorpion Files
Cloudsley-Thompson, 2012 Thompson JL. About the
enigmatic presence of the genus
Scorpio Linnaeus, 1758 in
Congo with the description of a
new species from Niger
(Scorpiones, Scorpionidae).
Serket. 2012;13(1/2):1-7.
Scorpio occidentalis Werner,
1936
New status Mali, Mauritania
& Senegal
Previous subspecies status: Scorpio
maurus occidentalis Werner, 1936
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio palmatus (Ehrenberg,
1829)
New status Israel, Palestina,
Egypt
Previous subspecies status: Scorpio
maurus palmatus Ehrenberg, 1829
Talal S, Tesler I, Sivan J, Ben-
Shlomo R, Muhammad Tahir H,
Prendini L, et al. Scorpion
speciation in the Holy Land:
Multilocus phylogeography
corroborates diagnostic
differences in morphology and
burrowing behavior among
Scorpio subspecies and justifies
recognition as phylogenetic,
ecological and biological
species. Mol Phylogenet Evol.
2015 May 15.
Scorpio propinquus (Simon,
1872)
New status Israel, Syria Previous subspecies status: Scorpio
maurus propinquus (Simon, 1872)
Talal S, Tesler I, Sivan J, Ben-
Shlomo R, Muhammad Tahir H,
Prendini L, et al. Scorpion
speciation in the Holy Land:
2020 © Jan Ove Rein, The Scorpion Files
Multilocus phylogeography
corroborates diagnostic
differences in morphology and
burrowing behavior among
Scorpio subspecies and justifies
recognition as phylogenetic,
ecological and biological
species. Mol Phylogenet Evol.
2015 May 15.
Scorpio punicus Fet, 2000 New status Algeria, Libya,
Morocco &
Tunisia
Previous subspecies status: Scorpio
maurus tunetanus Birula, 1910
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio savanicola Lourenco,
2009
New sp. Cameroon Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Scorpio sudanensis Lourenco
& Cludsley-Thompson, 2009
New sp. Sudan Lourenco WR, Cloudsley
Thompson JL. A new species of
the genus Scorpio Linnaeus 1758
from Sudan (Scorpiones,
Scorpionidae). Boletin Sociedad
Entomológica Aragonesa. 2009
(45):123-6.
2020 © Jan Ove Rein, The Scorpion Files
Scorpio tassili Lourenco &
Rossi, 2016
New sp. Algeria Lourenco WR, Rossi A.
Confirmation of a new species of
Scorpio Linnaeus, 1758 in the
Tassili N’Ajjer Mountains,
South Algeria (Scorpiones:
Scorpionidae). Onychium.
2016;12:11-8.
Scorpio weidholzi Werner,
1929
New status Morocco Previous subspecies status: Scorpio
maurus weidholzi Werner, 1929
Lourenco WR. Reanalysis of the
genus Scorpio Linnaeus 1758 in
sub-Saharan Africa and
description of one new species
from Cameroon (Scorpiones,
Scorpionidae). Entomol Mitt
Zool Mus Hamburg.
2009;15(181):99-113.
Srilankametrus Couzijn, 1981
Srilankametrus caesar (C.L.
Koch, 1841)
Srilankametrus couzijni
Prendini & Loria, 2020
Srilankametrus pococki
Prendini & Loria, 2020
Species transferred from
Heterometrus (New
New status
New status
New sp.
New sp.
India
Sri Lanka
Prendini L, Loria SF. Systematic
revision of the Asian Forest
Scorpions (Heterometrinae
simon, 1879), revised
suprageneric classification of
Scorpionidae latreille, 1802, and
revalidation of Rugodentidae
Bastawade et al., 2005. Bulletin
of the American Museum of
Natural History. 2020(442):1-
480.
2020 © Jan Ove Rein, The Scorpion Files
combinations):
Srilankametrus gravimanus
(Pocock, 1894)
Srilankametrus indus
(DeGeer, 1778)
Srilankametrus serratus
(Pocock, 1900)
Srilankametrus yaleensis
(Kovařik et al., 2019)
Urodacus butleri Volschenk,
Harvey & Prendini, 2012
New sp. Australia Volschenk ES, Harvey MS,
Prendini L. A new species of
Urodacus (Scorpiones:
Urodacidae) from Western
Australia. American Museum
Novitates. 2012(3748):1-18.
