Red List of vascular plants of the Wadden Sea Area

25
HELGOLANDER MEERESUNTERSUCHUNGEN Helgol6nder Meeresunters, 50, Suppl., 43-67 (1996) zgl.: SCHRIFTENREIHE FOR LANDSCHAFTSPFLEGE UND NATURSCHUTZ, H. 47 (1996) IV. Red List of Vascular Plants of the Wadden Sea Area* CONTRIBUTORS: Denmark: P. Wind Germany: M. van der Ende, E. Garve, A. Schacherer The Netherlands: J. B. M. Thissen'" INTRODUCTION In the aquatic part of the Wadden Sea area, there are only very few vascular plants, mainly Zostera marina and Z. noltii, but floristically the Wadden Sea islands are among the most important areas of northwestern Europe, especially for plants of coastal dunes and salt marshes (Mennema & Weeda, 1983). However, according to Raabe (1981), the Schleswig-Holstein part is relatively poor in this respect. One has to bear in mind that the number of vascular plants of some Wadden Sea is- lands has increased a lot in this century: e.g. for Vlieland, Terschelling, Ameland and Schiermonnikoog, the flora list comprised 60-90% more species in 1986 than in 1870 (Westhoff & van Oosten, 1991). About one century ago, the dunes were in a very poor state because of intensive exploitation (Westhoff, 1985). Delimitation of the area and selection of species At the Trilateral Governmental Conference in Esbjerg, it was decided to work out a Red List of the species which can be found in and which are typical for typical coastal ha- bitats only. The vascular plants of the Wadden Sea itself (aquatic and intertidal part), the salt marshes and the dunes,excluding plantations, have been taken into consideration. So, for example, the flowering plants of arable land have been deliberately excluded from the Red List. (The weed flora of the Wadden Sea islands used to be very rich [Westhoff & van Oosten, 1991].) Further, the list has been restricted to indigenous or naturalized species. As a guide- line, the lists of Prins et al. (1983 a, b) have been used. The apomictic-species of the ge- nera Rubus and Taraxacum have not been assessed, because of lack of information on This list forms part of the Report on the RED LISTS OF BIOTOPES, FLORA AND FAUNA OF THE TRILATERAL WADDEN SEA AREA. For basic information concerning, for example, func- tion of these lists, species taken into account, structure of the lists and abbreviations used, see also the general introduction to the Red Lists. " " Author of the introductory text. Biologische Anstalt Helgoland, Hamburg und Bundesamt ffir Naturschutz, Bonn

Transcript of Red List of vascular plants of the Wadden Sea Area

Page 1: Red List of vascular plants of the Wadden Sea Area

HELGOLANDER MEERESUNTERSUCHUNGEN Helgol6nder Meeresunters, 50, Suppl., 43-67 (1996)

zgl.: SCHRIFTENREIHE FOR LANDSCHAFTSPFLEGE UND NATURSCHUTZ, H. 47 (1996)

IV. Red List of Vascu lar Plants

of the W a d d e n Sea Area*

C O N T R I B U T O R S :

D e n m a r k : P. W i n d

G e r m a n y : M. v a n d e r E n d e , E. G a r v e , A. S c h a c h e r e r

T h e N e t h e r l a n d s : J. B. M. T h i s s e n ' "

INTRODUCTION

In the aquatic part of the Wadden Sea area, there are only very few vascular plants, mainly Zostera marina and Z. noltii, but floristically the Wadden Sea islands are among

the most important areas of nor thwestern Europe, especial ly for plants of coastal dunes and salt marshes (Mennema & Weeda, 1983). However , according to Raabe (1981), the

Schleswig-Hols te in part is relatively poor in this respect. One has to bear in mind that the number of vascular plants of some Wadden Sea is-

lands has increased a lot in this century: e.g. for Vlieland, Terschelling, A m e l a n d and Schiermonnikoog, the flora list comprised 60-90% more species in 1986 than in 1870

(Westhoff & van Oosten, 1991). About one century ago, the dunes were in a very poor state because of in tensive exploitation (Westhoff, 1985).

D e l i m i t a t i o n of t h e a r e a a n d s e l e c t i o n of s p e c i e s

At the Trilateral Governmenta l Conference in Esbjerg, it was decided to work out a

Red List of the species which can be found in and which are typical for typical coastal ha- bitats only. The vascular plants of the Wadden Sea itself (aquatic and intert idal part), the salt marshes and the dunes ,exc luding plantations, have been taken into consideration.

So, for example, the f lowering plants of arable land have b e e n del iberate ly exc luded from

the Red List. (The w e e d flora of the Wadden Sea islands used to be very rich [Westhoff & van Oosten, 1991].)

Further, the list has been restricted to indigenous or natural ized species. As a guide- line, the lists of Prins et al. (1983 a, b) have b e e n used. The apomict ic-species of the ge-

nera Rubus and Taraxacum have not b e e n assessed, because of lack of informat ion on

�9 This list forms part of the Report on the RED LISTS OF BIOTOPES, FLORA AND FAUNA OF THE TRILATERAL WADDEN SEA AREA. For basic information concerning, for example, func- tion of these lists, species taken into account, structure of the lists and abbreviations used, s e e

also the general introduction to the Red Lists. " " Author of the introductory text.

�9 Biologische Anstalt Helgoland, Hamburg und Bundesamt ffir Naturschutz, Bonn

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44 Vascular Plants of the Wadden Sea Area

their distr ibution and status. Subspecies are only ment ioned if this adds any value, e.g. if the specific subspecies is threatened, whereas the species as such is not.

The taxonomy and the scientific names follow the standard list for plants in Germany (Zentralstelle ffir die floristische Kartierung, 1993).

Da ta sou rce

Denmark: As in the case of the other species groups, the Danish Wadden Sea Red List of vascular plants is an upda ted version of the Danish nat ional Red List (Asbirk & So-

gaard, 1991). Germany: The status of the vascular plants of Niedersachsen has b e e n based, to a

large extent, on the Red List of that region (Garve, 1993). For Schleswig-Holstein, howe- ver, the status categories of the Red List (Mierwald & Beller, 1990) had to be adapted to

the situation in the Wadden Sea part. The Netherlands: The status of the species has been determined with the aid of the

atlases by M e n n e m a et al. (1980, 1985) and van der Meijden et al. (1989), which give the distribution in the two periods before and after 1950. This information was upda ted from some new publications, mainly Westhoff & van Oosten (1991) and Weeda et al. (1985- 1994).

T h r e a t s

In 1956, Westhoff pointed out that the loss of plant species in the coastal dunes has in general been relatively slight. At that time, the main threats were extraction of drin- king water, dra inage of adjacent agricultural fields, which cause the desiccation of hu- mid and wet dune slacks, and afforestation, which causes desiccation too and even di- rectly the destruction of natural habitats. Nowadays, one has to take more threats into ac- count, namely atmospheric deposition of ni t rogen (Bellemakers et al., 1993; Veer et al., 1993), recreation (van Zoest, 1994), regulat ion of moving sand and coastal defense (Jans- sen, 1994).

Desiccation and atmospheric deposition of ni t rogen cause the dominance of a few grasses and of the invasive neophyte moss Campylopus introflexus. Recreation concen- trates on the beach and disturbs typical beach plants, like the three Atriplex species of the Red List. Pioneer plants are threa tened by fixation of moving sand. The flora of the outer dunes is threa tened by regression of the coastline, but also to some extent by the measures which are taken to counteract this process. A specific effect of coastal defense is the loss of brackish waters. Several plants of gradients be tween fresh and salt water are disappearing. A lot of salt marshes and their characteristic plants d i sappeared locally because of e m b a n k i n g and reclamation.

In 1932 the sublittoral Zostera marina populat ions perished because of the ,,wasting disease". After 1965, also Zostera noltii began to decline in the Dutch part because of in- creasing pollut ion (den Hartog & Polderman, 1975). The relevant effects of eutrophica- tion are turbidi ty and high densit ies of microalgae and lugworms (Arenicola marina L.) (Philippart & Dijkema, 1995). Since about 1972 Zostera noltii has b e e n recover ing in the Dutch part. In the German Wadden Sea mass deve lopment of macroa lgae probably cau- sed a recent decline of seagrass (Reise et al., 1994). Seagrasses are also th rea tened by fis- hery of shellfish (de Jonge & de Jong, 1992).

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Vascu la r P lan ts of t he W a d d e n Sea Area 45

R e g u l a t i o n of h u m a n d i s t u r b a n c e is no t a l w a y s bene f i c i a l to the flora. S o m e rare s p e -

cies b e n e f i t f rom a cer ta in , s l ight , a m o u n t of e u t r o p h i c a t i o n or t r e ad ing , e.g. t h e R e d List

s p e c i e s Alyssum alyssoides, Anacamptis pyramidalis, Asparagus officinalis ssp. prostra- tus a n d Dactylorhiza maculata. O n s o m e salt m a r s h e s , e spec i a l ly in S c h l e s w i g - H o l s t e i n ,

g r a z i n g p r e s s u r e res t r ic t s p l a n t d ive r s i ty (Stock, 1993). Low in t ens i ty g raz ing , h o w e v e r ,

can be bene f i c i a l to t he flora of salt m a r s h e s (Scher fose , 1993) a n d d u n e s (Westhoff , 1985).

S u m m a r y

In the W a d d e n Sea area, a total of 248 ( sub ) spec i e s of v a s cu l a r p lan t s are t h r e a t e n e d

in at l eas t one s u b r e g i o n . Of t hese , 216 ( s u b ) s p e c i e s are t h r e a t e n e d in the en t i r e a r e a a n d

are t h e r e f o r e p l a c e d on the t r i la tera l Red List. 17 ( sub ) spec i e s of the l i s ted v a s c u l a r p l an t s

are (probably) ex t inc t in t he en t i r e W a d d e n Sea area. The s ta tus of 47 ( sub ) spec i e s of vas -

cular p l an t s is (probably) critical; 61 ( s u b ) s p e c i e s are (probably) e n d a n g e r e d ; t he s t a tus

of 65 ( sub ) spec i e s is (probably) v u l n e r a b l e a n d tha t of 26 ( sub) spec ies s u s c e p t i b l e .

R E F E R E N C E S

Asbirk, S. & S. Sogaard (Eds), 1991. ,,Rodliste 90". - S~rligt beskyttelseskr~evende planter og dyr i Danmark. (Red Data book of threatened plants and animals in Denmark - with English summary) - Milj~ministeriet, Skov- og Naturstyrelsen, Kobenhavn, 222 pp.

Bellemakers, M. J. S., Maessen, M., Cals, M. J. R. & J. G. M. Roelofs, 1993. Effectgerichte maatre- gelen tegen verzuring en eutrofiering van oppervlaktewateren. Katholieke Universiteit, Nijme- gen, 147 pp.

Garve, E., 1993. Rote Liste der gef~ihrdeten Farn- und Blfitenpflanzen in Niedersachsen und Bre- men. 4. Fassung vom 1.1.1993. - Infn. Naturschutz Nieders. 13, 1-37.

Gravesen, P, 1983. Forelobig oversigt over botaniske lokaliteter. 4. S~nderjyllands Amt. - Miljomi- nisteriet, Fredningsstyrelsen, Kobenhavn, 158 pp.

Hansen, K. (red.), 1981. Dansk feltfIora.- Gyldendal, Kobenhavn, 757 pp. EIartog, C. den & P. J. G. Polderman, 1975. Changes in the seagrass populations of the Dutch Wad-

denzee. - Aquat. Bot. i, 141-147. Janssen, M., 1994. Dynamisch kustbeheer. - Duin 17(2), 17-19. Jonge, V. N. de & D. J. de Jong, 1992. Role of tide, light and fisheries in the decline of Zostera ma-

rina L. in the Dutch Wadden Sea. - Netherl. Inst. Sea Res. Publ. Ser. 20, 161-176. Meijden, R. van der, Plate, C. L. & E. J. Weeda, 1989. Atlas van de Nederlandse flora. 3: Minder

zeldzame en algemene soorten. Rijksherbarium, Leiden, 264 pp, Mennema, J., Quene-Boterenbrood, A. J. & C. L. Plate (Eds), 1980. Atlas van de Nederlandse flora.

1: Uitgestorven en zeer zeldzame planten. Kosmos, Amsterdam, 226 pp. Mennema, J., Qu~ne-Boterenbrood, A. J. & C. L. Plate (Eds), 1985. Atlas van de Nederlandse flora.

2: Zeldzame en vrij zeldzame planten. Bohn, Scheltema & Holkema, Utrecht, 349 pp. Mennema, J. & E. J. Weeda, 1983. Vascular plants. In: Ecology of the Wadden Sea. Ed. by W. J. Wolff.

Balkema, Rotterdam, 3, 9/38-9/50. Mierwald, U. & J. Belier, 1990. Rote Liste der Farn- und Blfitenpflanzen Schleswig-Holsteins. 3. Fas-

sung. Landesamt ffir Naturschutz and Landschaftspflege, Kiel, 64 pp. Pedersen, A., 1983. The vegetation of the Wadden Sea islands of Denmark and Schleswig-Holstein.

In: Ecology of the Wadden Sea. Ed. by W. J. Wolff. Balkema, Rotterdam, 3, 9/137-9/165. Philippart, C. J. M. & K. S. Dijkema, 1995. Wax and wane of Zostera noltii L. in the Dutch Wadden

Sea. - Aquat. Bot. 49, 255-268. Prins, D., Kuhbier, H., Pedersen, A., Mennema, J. & A. M. Kooijman, 1983 a. Main list with indige-

nous or naturalized phanerogams and pteridophytes of the Wadden Sea area. In: Ecology of the Wadden Sea. Ed. by W. J. Wolff, Balkema, Rotterdam, 3, 9/323-9/344.

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46 V a s c u l a r P l a n t s of t he W a d d e n S e a A r e a

Prins, D., Kuhbier, H., Pedersen, A., Mennema , J. & A. M. Kooijman, 1983 b. Separa te list with indi- genous or natural ized plants of smaller islands and ,, h igh-sands ". In: Ecology of the Wadden Sea. Ed. by W. J. Wolff. Balkema, Rotterdam, 3, 9/345-9/346.

Raabe, E.W., 1981. Uber das Vorland der 6stlichen Nordseek/iste. - Mitt. ArbGem. Geobotanik in Schleswig-Holstein und Hamburg 31, 1-118.

Reise, K., Kolbe, K. & V. de Jong, 1994. Makroa lgen und Seegrasbest~inde im Wattenmeer . In: Warn- signale aus dem Wattenmeer. Ed. by J. L. Lozan, E. Rachor, K. Reise, H. yon W e s t e r n h a g e n & W. Lenz. Blackwell, Berlin, 90-100.

Scherfose, V., 1993. Zum EinfluB der Beweidung auf das Gef~l~pflanzen-Artengeffige von Salz- und Brackmarschen. - Zeitschr. Okol. Naturschutz 2, 201-211.

Stock, M., 1993. Salt marshes in Schleswig-Holstein: from a green towards a na tura l succession. - Wadden Sea Newslet ter 1993-1, 11-14.

Veer, M., Meulen, F. van der, Heil, G. W. & A. M. Kooijman, 1993. Ecotoopbeheer in droge duinen: s turende processen en l imiterende factoren. In: Effectgerichte maa t rege len t e ge n verzuring en eutrofi~ring in natuur terre inen. Ed. by M. Cals, Graaf, M. de & J. Roelofs. Katholieke Universi- teit, Nijmegen, 147-170.

Westhoff, V., 1956. De verarming van flora en vegetatie. In: Vijftig jaar na tuu rbe sc he rming in Ne- derland. Na tuu rmonumen ten , Amsterdam, 151-186.

Weeda, E. J., Westra, R., Westra, C. & T. Westra, 1985. Neder landse oecologische flora 1. IVN, Am- sterdam, 304 pp.

Weeda, E. J., Westra, R., Westra, C. & T. Westra, 1987. Neder landse oecologische flora 2. IVN, Am- sterdam, 304 pp.

Weeda, E. J., Westra, R., Westra, C. & T. Westra, 1988. Neder landse oecologische flora 3. IVN, Am- sterdam, 302 pp.

Weeda, E. J., Westra, R., Westra, C. & T. Westra, 1991. Neder landse oecologische flora 4. IVN, Am- sterdam, 317 pp.

Weeda, E. J., Westra, R., Westra, C. & T. Westra, 1994. Neder landse oecologische flora 5. IVN, Am- sterdam, 400 pp.

Westhoff, V., 1985. Nature m a n a g e m e n t in coastal areas of Western Europe. - Vegetat ion 62, 523- 532.

Westhoff, V. & M.F. van Oosten, 1991. Plantengroei van de Waddenei landen. KNNV, Utrecht, 417 pp.

Wind, P., 1994 a. Oversigt over botaniske lokaliteter. Bind 10. Ribe Amt.- Miljoministeriet, Fred- ningsstyrelsen, Skov- og Naturstyrelsen, Kobenhavn, 176 pp.

Wind, P., 1994 b. Oversigt over botaniske lokaliteter. Status og forvaltningsbehov.- Miljoministeriet, Fredningsstyrelsen, Skov- og Naturstyrelsen, Kobenhavn, Vol. I, 68 pp; Vol. 2, 94 pp.

Zentralstelle ffir die floristische Kart ierung der Bundesrepubl ik Deutsch land-Nord (Hrsg.), 1993. Standardliste der Farn- und Blii tenpflanzen der Bundesrepublik. - Floristische Rundbr. Beih. 3, 48O pp.

Zoest, J. van, 1994. Effecten van openluchtrecreat ie op na tuu rwaa rden van de Waddene i landen . Landelijke Vereniging tot Behoud van de Waddenzee , Harlingen, 74 pp.

RED LIST O F V A S C U L A R P L A N T S O F T H E W A D D E N SEA A R E A "

E X - E x t i n c t :

Ajuga pyramidalis (?) Alyssum alyssoides Ana camp tis pyramidalis Carex dioica Carex hostiana Chenopodium botryodes

Erica cinerea (?) Pi]ago ]utescens Genista gennamca PIieracium vulga turn (?) PIypericum humifusum []lecebrum vertici]]a turn (?) A4arrubium vulgate

Quest ion-marks indicate that in one of the subregions the status of threat is uncertain.

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Vascular Plants of the Wadden Sea Area 47

(?) Ranunculus hederaceus Ranunculus tripartitus Subularia aquatica Zannichellia palustris ssp. polycarpa

C R - C r i t i c a l :

Arnica montana Artemisia campestris Baldellia ran unculoides Bassia hirsuta Blysm us compressus Ca]litriche brutia Calystegia soldanella Carex caryophyllea Carex ericetorum Carex p u]icaris Carex pun cta ta Catabrosa aquatica Cicendia filiformis Cochlearia o[ficinalis Cram be maritima Cuscuta epithymum Deschampsia setacea Dianthus carthusianorum Dianthus deltoides Eleocharis acicularis Equisetum variegatum Eriophorum vaginatum Euphorbia paralias Filago vulgaris Gentianella campestris ssp. baltica Gnapha]ium luteo-album Hammarbya paludosa Hermmium monorchis Horde um marin um Hottonia pa]ustrfs Huperzia se]ago Juncus capitatus Juniperus communis Orchis morio Pilulalffa globulifera Pinguicula vulgaris (?) Rhynchospora alba Rhynchospora fusca Sagina subulata Schoenoplectus pungens

Silene nutans Sparganium natans Stellaria aquatica (?) Thalictrum flavum Tlffchophorum cespitosum Tuberaria guttata Utricularia minor

E N - E n d a n g e r e d :

Acinos arvensis =Satureja acinos Alopecurus bulbosus Anagallis minima =Centunculus minimus Antennaria dioica Apium gra veolens Apium inunda turn A triplex gla briuscula A triplex la cinia ta Botrychium lunaria Bupleurum tenuissimum Carex trinervis Carex viridula

=C.oederi =C.serotina ssp. serotina

Carlina vulgaris Cirsium dissectum Cotula coronopifoh'a Dactylorhiza incarnata Drosera in termedia Epipactis palustris Erica scoparia Euphrasia micran tha Galium sterneri Gentiana pneumonanthe Gentianella uliginosa Gymnadenia conopsea Halimione peduncula ta Helictotrichon pubescens Hyperfcum pulchrum Hypochoeris glabra Isolepis fluitans Juncus pygmaeus La thyrus linifolius Limosella aquatica Liparis loeselii Listera cordata Littorella uniflora

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48 Vascular Plants of the Wadden Sea Area

(?) Lysimachia thyrsiflora (?) Myosurus minimus Myriophyllum alterniflorum Narthecium ossifragum Odontites litoralis Oenanthe lachenalii Pedicularis palustris Pedicularis sylvatica Platanthera bifolia Polygala serpyllifolia Polygonum oxyspermum ssp. rail Potamogeton gramineus Potamogeton perfolia tus Pyrola rotundifolia Ruppia cirrhosa Ruppia maritima Schoenoplectus triqueter Schoenoplectus X carinatus Schoen us nigricans Scorzonera humilis Scutellaria minor Senecio erucifolius (?) Stachys arvensis Ulex europaeus Vaccinium vitis-idaea Vulpia bromoides

V U - V u l n e r a b l e :

Agrostis vinealis =A.stricta ssp.stricta Anagallis teneHa (?) Anthyllis vulneraria ssp. maritima (?) Artemisia absmthium Asparagus officinalis ssp. prostratus (?) Atriplex Iongipes Blysm us rufus Bnza media Caltha palustris Carex distans Carex extensa Carex vesicaria Carex viridula van pulchella

=C.oederi ssp. oederi =C.oederi ssp. pulchella =C.scandinavica =C.serotina spp, pulchella

Centaurium erythraea

Crepis tectorum Dactylorhiza maculata Dacty]orhiza maja]is Dactylorhiza praetermissa E]eocharis m ultica ufis E]eocharis quinqueflora Eryngium maritim um Euphrasia stricta Fi]ago minima (?) Galeopsis speciosa Ha]imione portulacoides = Atrip]ex portu- lacoides Hierochloe odorata Inu]a britannica Juncus anceps Juncus balticus Juncus inflexus Lathyrus maritimus =L. japonicus Lepidium campestre Linum catharticum Lycopodie]la inundata Lycopodium davatum Menyanthes trifoliata Ophioglossum vulgatum Osmunda regalis Parapholis strigosa Parnassia palustris Peplis portula Polygala vulgaris Potamogeton polygonifolius Potamogeton trichoides Potentilla argentea Pyrola minor Radiola linoides Ranunculus baudotii Rhinanthus angustifolius (?) Rhinanthus angustifolius ssp. halophi-

]us Rhinanthus minor Rosa pimpinellifolia Sagina nodosa Samolus valerandi Sderan th us perennis Silene otites Taraxacum celticum agg. Taraxacum palustre agg.

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Vascular Plants of the W a d d e n Sea Area 49

Thalictrum minus Torilis nodosa Triglochm palustre Utricularia a ustralis Vaccinium myrtillus Zostera marina Zostera noltii

SU - S u s c e p t i b l e :

Ajuga reptans Alisma ]anceola turn Apium nodiflorum Arabis hirsuta Berberis vulgaris Carex hartmanii Carex lasiocarpa Carex remo ta (?) Cerastium pumilum

(?) Cicuta virosa Crithmum maritimum Deschampsia wibeliana Desmazeria marina Galeopsis segetum (?) duncus bulbosus ssp. kochii Lactuca tatarica Lithospermum officinale Polygonatum odoratum Potamogeton coloratus Ranunculus auricomus Sedum album Smyrnium olusatrum Spergula morisonii Thym us pulegioides Trifolium striatum Xanthium albinum ssp. albinum

LIST OF T H R E A T E N E D V A S C U L A R P L A N T S OF T H E W A D D E N SEA A R E A

Acinos arvensis (Lam.)Dandy

=Satureja acinos (L.)Scheele

Agrostis vineafis Schreb. =A.stricta J.F.Grnelin ssp.stricta

Ajuga pyramidalis L.

Ajuga reptans L.

Alisma lanceolatum With.

Alopecurus bulbosus Gouan

Alyssum alyssoides (L.) L. 1

Kleine steentijm Gemeiner Steinquendel Voldtimian

Zandstruisgras Sand-Straul~gras Sand-Hvene

Piramidezenegroen Pyramiden-G0nsel Pyramide-La~beles

Kruipend zenegroen Kriech-G0nsel Krybende Laebeles

Slanke waterweegbree Lanzett-FroschlOffel Lancet-Skeblad

Knolvossestaart Knollen-Fuchsschwanz Knold-Raevehale

Bleak schildzaad �9 Kelch-Steinkraut Gr&dodder

Red List (trilateral) Threats

EN HAB, AGR

VU AGR, HAB

EX

SU

SU

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

SU

EN

* * CR

EX

SU

EN WAT EN EN

- E X ? ? EX EX (*)

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50 Vascular Plants of the Wadden Sea Area

Hondskruid Anacamptis pyramidalis Pyramiden-Spitzorchis ( L . ) R i ch .

Horndrager i i

Red List (trilateral) Threats

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

Anagaflis minima (L.) E.H.L.Krause

,i ( =Centunculus minimus L_)

Dwergbloem Acker-Kleinling Knudearve

~ X AGR EX -

Teer guichelheil Anagallis tenella (L.) L. Zarter Gauchheil

Spaed arve

Rozenkransje Antennaria dioica (L.)Gaertn. Gemeines KatzenpfOtchen

Almindelig Kattefod

Wondklaver Anthyllis vulneraria ssp. Gemeiner Wundklee maritima (Schweigg.) Co rb . Rundbaelg

Selderij Apium graveolens L. Echter Sellerie

Vild Selteri

Apium inundatum (L.) Rchb.fiL

i ....

Apium nodiflorum (L.) Lag.

j ,

Arabis hirsuta (L.) Stop.

Armeria maritima (Mill.) WiUd.

Arnica montana L.

Artemisia absinthium L. 1

Artemisia campestris L.

Ondergedoken moeras- scherm Flutender Sellerie Svemmende Sumpskaerm

Groot moerasscherm KnotenbliJtiger Scheiberich Vand-Selleri

Ruige scheefkelk Rauhaarige G~nsekresse Stivh~ret Kalkkarse

Engels gras Gemeine Grasnelke Engelskgraes

Wolverlei Arnika ~lm. Guldblomme

Absintalsem Wermut Have-Malurt

Averuit Feld-Beifu& Mark-Bynke

I ,

Liggende asperge Asparagus officinalis L. ssp. Spargel (ssp. prostratus) prostratus (Dum.) Corb. Asparges (ssp. prostratus)

EN AGR, WAT VU EN CR VU

VU WAT VU

EN EUT VU EX CR *

VU? WAT * VU ?

EN WAT, AGR

EN HAB, EUT, AGR, WAT

SU

SU

* AGR, WAT

EUT, AGR CR DIS

VU? ?

CR AGR

VU AGR

EN EN CR

VU EX EN SU

SU

SU

VU * * *

EX EX EN SU

CR * (*) (*)

SU EX SU *

EN *

Page 9: Red List of vascular plants of the Wadden Sea Area

V a s c u l a r P l a n t s of t he W a d d e n Se a A r e a

R e d L i s t

( t r i l a t e r a l )

Gefleugeld sterrekroos Teich-Wasserstern Storfrugtet Vandstjerne

Threats

Kustmelde Atriplex glabriuscula Edmon- Kahle Melde EN DIS ston

Tykbladet Maelde

Gelobde melde Atriplex laciniata L. Gelappte Melde EN DIS

Selv-Maelde

Spiesmelde (var. Ionglpes) Atriplex Iongipes Drejer Gestielte Spie~melde VU? DIS

Stilk-Maelde

Stijve moerasweegbree Baldellia ranunculoides Igelschlauch CR WAT, EUT (L.) Parl. Aim. S~pryd

Ruig zoutkruid Bassia hirsuta (L.)Asch. Rauhhaarige Dornmelde CR DIS, WAT

Tangurt

Zuurbes Gemeine Berberitze, SU

Berberis vulgans L. Sauerdorn

iAIm. Berberis

Platte bies Blysmus compressus Flaches Quellried CR WAT (L) Link Fladtrykt Kogleaks

Rode bies AGR, HAB, Blysmus rufus (Huds.) L i n k Rotbraunes Quellried MU WAT

Redbrun Kogleaks

Getobde maanvaren Botrychium lunana (L.) Swartz Echte Mondraute EN EUT

~AIm. M~nerude

Bevertjes Briza media L. Gemeines Zittergras MU AGR

Aim. Hjertegraes

Zwenkdravik Bromus tectorum L. 2 Dach-Trespe *

Tag-Hejre

Fijn goudscherm Bupleurum tenuissimum L. Salz-Hasenohr EN AGR

Smalbladet Hareere

Callitriche brutia Petagna Stielfr(Jchtiger Wasserstern CR EUT Stilkfrugtet Vandstjerne

Callitriche stagnalis stop. *

51

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK i i

EN SU CR *

r l

�9 EX CR *

i i

? * CR *

EN CR EX

EX CR VU

SU

EN

VU

VU

SU

i i

EX CR *

I I

EN CR *

i i

CR CR

I I

!

. ( * ) *

II

CR CR *

I I

CR

SU *

Page 10: Red List of vascular plants of the Wadden Sea Area

52 Vascular Plants of the Wadden Sea Area

Caltha palustris L.

Calystegia soldanella (L.) Roem. & Schuft.

Carex canescens L.

Carex caryophyllea Lat.

Carex dioica L.

Carex distans L.

Dotterbloem Sumpf-Dotterblume Eng-Kabbeleje

Zeewinde Strand-Winde Strand-Snerle

Zompzegge !Grau-Segge Gr& Star

'Voorjaarszegge Fr0hlings-Segge V&r-Star

Tweehuizige zegge Zweih~lusige Segge Tvebo Star

Zilte zegge Entferntahrige Segge Fjernakset Star

Red List (trilateral) Threats

MU W A T

CR DIS

CR AGR

EX EUT, WAT, AGR

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

Carex ericetorum Po//ich Heidezegge Heide-Segge Lyng-Star

Carex extensa Good. Kwelderzegge Strand-Segge Udspilet Star

VU VU VU *

SU CR CR SU

SU * * *

EX EN

EX EX EX

VU AGR * VU EN *

Carex flacca Schreb. Zeegroene zegge Blaugr0ne Segge Bl~lgren Star

CR AGR, EUT

Carex hartmanii Caj. Kleine Knotszegge Hartmans Segge Hartmans Star

Carex hostiana DC. Blonde zegge Saum-Segge Skede-Star

VU AGR, HAB

* AGR, WAT

Draadzegge Carex lasiocarpa Ehrh . Faden-Segge

Tr&d-Star

Vlozegge Carex pulicaris L. Floh-Segge

Loppe-Star

Stippeizegge Carex punctata Gaudin Punktierte Segge

Prikket Star

SU

EX WAT, AGR

WAT, AGR, EUT

SU

CR

CR I

WAT

CR

* SU EN *

* VU * *

SU

EX EX EX

SU

EN EX EX *

SU CR

Page 11: Red List of vascular plants of the Wadden Sea Area

V a s c u l a r P l a n t s of t h e W a d d e n Se a A r e a

Carex remota L.

II

Carex trinervis Degl.

Carex vesicaria L.

Carex viridula # Michx. = C. oeder i = C.serotina Merat ssp. seroti-

na

Carex viridula ssp. oedocar-

0 a # ( A n d e r s s . ) S c h m i d

= C.demissa Hornem = C.tumidicarpa Anderss. = C o e d e r i ssp. oedocarpa

A n d e r s s .

Carex viridula var. pulchel la # ~Lonnr . ) S c h m i d

= C.oederi ssp. oederi = C.oederi ssp. pulchella = C.scandinavica Davies = C.serotina Merat spp. pul-

chella (LOnnr.)

Carlina vulgaris L. s.str.

II

Carum carvi L.

Catabrosa aquatica (L.) P.Beauv.

Centaur ium erythraea Rafn

IJle zegge WinkeI-Segge Akselblomstret Star

Drienervige zegge Dreinervige Segge Klit-Star

Blaaszegge Blasen-Segge Blaere-Star

Geelgroene zegge / Dwergzegge Kleine Gelb-Segge /Erte Star

Geelgroene zegge Grene Segge Gren Star

Dwergzegge Spate Segge Host-Star

Driedistel Golddistel Bakketidsel

Echte karwij Wiesen-K0mmel Kommen

Watergras Quellgras Taeppegraes

Echt duizendguldenkruid Echtes Tausendgt~lden- kraut Mark-Tusindgylden

53

,=

Status of threat in the subre- Red L is t Threats gions of the Wadden Sea Area

( t r i la tera l ) NL Nds SH DK

u

S U SU

II

EN * EN EX VU

II

VU WAT SU VU *

II

EN WAT VU VU CR

I

* SU *

VU WAT VU VU CR *

EN HAB,EUT VU CR

* * V U *

C R W A T C R E X EN

VU WAT VU

II

SU

II

SU

# Obviously there is a problem concerning the taxonomy of Carex viridula which could not be solved on the trilateral level.

Page 12: Red List of vascular plants of the Wadden Sea Area

54 Vascular Plants of the Wadden Sea Area

i

i I

,J Cerastium diffusum Pers.

i i

Cerastium pumilum Curtis

Chenopodium botryodes Sm.

Scheve hoombloem Viermanniges Hornkraut Firehannet Hensetarm

Steenhoornbloem Dunkles Zwerg-Hornkraut Liden Hensetarm

Beursjesganzevoet Dickbl.~ttriger G~tnsefu& Drue-G&sefod

Draadgentiaan Heide Zindelkraut Bitterblad

Red List (trilateral)

Spaanse ruiter Englische Kratzdistel Engelsk Tidsel

SU?

EX

Threats

DIS

Cicendia filiformis (L.)Detarbre CR WAT, EUT

Waterscheerling Cicuta virosa L. Wasserschierling SU?

Aim. Gifttyde

Cirsium dissectum (L.) Hill EN WAT, AGR

Engelss lepelblad Englisches Ltfffelkraut Engelsk Kokleare

Goudknopje Krahenfu&-Laugenblume Firkl~ft

AGR, DIS, WAT Cochleana anglica L. *

Echt lepelblad Cochlearia officinalis L. Gebrauchliches Ltfffelkraut CR WAT

Laege Kokleare

Cotula coronopifolia L. EN AGR, WAT

Zeekool Meerkohl Strandk~l

Smal streepzaad Dach-Pippau, Mauer-Pip- Dau Tag-H~geskaeg

Zeevenkel

Stranddild

Crambe maritima L.

Crepis tectorum L.

i II

Crithmum maritimum L.

I I

Cuscuta epithymum (L) L

I I

CR

VU

SU

AGR

CR AGR

Dactylorhiza incarnata (L.)See i i

Klein warkruid QuendeI-Seide Lyng-Silke

Vleeskleurige orchis Steifbl~ttriges Knabenkraut Kedfarvet Gogeurt

EN

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK i :

* SU SU VU

i |

SU ?

H

EX

EN

SU

EN

SU

SU

SU

i i

EX

n

(*)

I I

I I

* VU

I I

CR CR *

I I

EN EN

I'

EX

EN

SU

EX EN EN SU

WAT, EUT VU CR *

Page 13: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the Wadden Sea Area

! R e d L is t ( t r i la tera l )

Eleocharis quinqueflora (F .X .Har tmann) O. S c h w a r z

Armbloemige waterbies Wenigbl0tige Sumpfbinse F~blomstret Kogleaks

Threats

Gevlekte orchis Dactylorhlza maculata (L.) Soo Geflecktes Knabenkraut VU HAB, AGR L Plettet G~geurt

Breedbladige orchis EUT, WAT, Dactylorhiza majalis (Rchb.) Breitblattriges Knabenkraut VU Hunt & S u m m e r h . AGR

Maj-G~geurt

Rietorchis Dactylorhiza praetermissa 0bersehenes Knabenkraut VU EUT, WAT, (Druce) Soo AGR

~ Priklaebet G~geurt

Moerassmele Deschampsia setacea (Huds.) Borst-Schmiele CR WAT, EUT Hacke l

Fin Bunke

Deschampsia wibeliana (Son- Schlamm-Schmiele SU HAB, WAT der) Part .

Laksteeltje Desmazeria marina (L.) Druce S U

Karthuizer anjer Dianthus carthusianorum L. Karthauser-Nelke CR HAB, EUT

Karteuser-Nellike

Steenanjer Dianthus deltoides L Heide-Nelke CR HAB, EUT

Bakke-Nellike

Kleine zandkool Diplotaxis muralis (L.) DC 1 Mauer-Doppelsame *

Mursennep

Kleine zonnedauw Drosera intermedia Hayne Mittlerer Sonnentau EN WAT

Liden Soldug

Ronde zonnedauw Drosera rotundifolia L. Rundbl~ittriger Sonnentau * WAT, AGR

Rundbladet Soldug

Naaldwaterbies Eleocharis acicularis (L.) Roem. NadeI-Sumpfsimse CR WAT, EUT & Schul t . N&le-Sumpstr&

Veelstengelige waterbies WAT, EUT, Eleocharis multicaulis (sin.) Vielstengelige Sumpfsimse VU Sm. Mangestaenglet Sumpstr~ AGR, HAB

VU WAT

55

Status of threat in the subre- j' 4

gions of the Wadden Sea AreaL, /

NL Ntis SH DK ii

VU VU VU *

ii

VU EN EN *

ii

VU SU

ii

CR CR SU

SU SU

SU

CR

SU CR

s u * (*)

CR VU

* V U *

SU EX EN

CR

VU EN EN

ii

(*)

ii

n

SU [' ] r

I'

i

Page 14: Red List of vascular plants of the Wadden Sea Area

56 Vascular Plants of the Wadden Sea Area

Epipactis palustris (L.) Crantz .

Equisetum variegatum Sch le icher ex. Web. & M o h r

ii

Moeraswespenorchis Echter Sumpfwurz Sump-Hullaebe

Bonte paardestaart Bunter Schachtelhalm Liden Padderokke

Rode dophei Erica cinema L. Grau-Heide

Gr~lyng ii

Erica scoparia L. Bezemdopheide

Red List Threats (trilateral)

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

EN W A T VU EN CR *

CR WAT CR CR

EX ? EX

EN ? EN

Eenarig wollegras Eriophorum vaginatum L. Scheidiges Wollgras CR WAT

Tue-Kaerutd i

! Blauwe zeedistel iEryngium maritimum L. Stranddistel VU DIS, WAT

Strand-Mandstro

Zeewolfsmelk Euphorbia paralias L. CR DIS

Kilt-Vertemaelk

Slanke ogentroost AGR, HAB, Schlanker Augentrost EN Lyng-Ojentr~st EUT, AFF

Euphrasia micrantha Rchb.

Stijve ogentroost AGR, HAB, Euphrasia stricta Owoiff ex. Steifer Augentrost VU J .F .Lehm. EUT

Spids Ojentr~st

Geel viltkruid Filago lutescens Jord. 1 Gelbliches Filzkraut EX?

Gulgr& Museurt

Dwergviltkruid HAB, EUT, Zwerg-Filzkraut VU AFF Liden Museurt

Filago minima (sin.)Pets.

HAB, EUT, AFF

Duits viltkruid Filago vulgaris Lain Deutsches Filzkraut CR

Kugle-Museurt

Bleekgele hennepnetel Galeopsis segetum Necker Saat-Hohlzahn SU

Gul Hanekro ii

Dauwnetel Galeopsis speciosa Mill. Bunter Hohlzahn VU?

Hamp-Hanekro

EX EX

VU EN VU

CR

EN CR *

* V U VU *

EX (*)

EN VU * *

CR CR *

SU

su v u (*)

Page 15: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the Wadden Sea Area

R e d L i s t

( t r i l a t e r a l )

Gentianella campestris ssp.baltica (Murb.) A. & D.Love

Threats

Galium sterneri Ehrendf. Sterners Labkraut EN EUT Liden Snerre

Duitse brem Genista germanica L. Deutscher Ginster EX

Tysk Visse

Verfbrem Genista tinctoria L. F~rber-Ginster * AGR, EUT

Farve-Visse

Klokjesgentiaan Gentiana pneumonanthe L. Lungen-Enzian EN WAT, AGR

Klokke-Ensian

CR WAT Veldgentiaan (ssp. baltica) Baltischer Enzian Baltisk Ensian

Slanke gentiaan Sumpf-Enzian Eng-Ensian

Gentianella uliginosa (Willd) EN WAT Borner

Bleekgele droogbloem Gnaphalium luteo-album L. Gelbwei6es Ruhrkraut CR WAT

Gulhvid Evighedsblomst

Grote muggenorchis Gymnadenia conopsea (L . ) Gro6e-Handelwur-z EN WAT R.Br. Langakset Tr~dspore

I Gesteelde zoutmelde i Halimione pedunculata (L . ) Gestielte Keilmelde EN DIS, AGR Aellen

Stilket Kilebaeger

Halimione portulacoides (L.) Gewone zoutmelde Aeilen Strand-Salzmelde VU DIS, AGR,

HAB (= Atriplex portulacoides) Stilkles Kilebaeger

Veenmosorchis Hammarbya paludosa Sumpf-Weichwurz CR WAT, EUT (L.) O. Kuntze

Femkantet Hjertelaebe

Zachte haver Helictotrichon pubescens Flaumiger Wiesenhafer EN AGR, EUT (Huds.) Pilger Ounet Havre

Honingorchis Herminium monorchis ( L . ) Einknollige Honigorchis, CR WAT R.Br. Einknolle

Pukkellaebe

Breukkruid Hemiaria glabra L. Bruchkraut *

Brudurt

57

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

CR *

EX EX

V U * *

VU EX CR *

CR CR EX VU

EN CR VU

CR EX

EN CR

* E N C R *

* * E N *

EX EX CR SU

SU EN

CR

SU *

Page 16: Red List of vascular plants of the Wadden Sea Area

58 Vascular Plants of the Wadden Sea Area

R e d L i s t Threats

( t r i l a t e r a l )

Hieracium vulgatum Fries

Hierochloe odorata (L.)P.Be- auv.

Hordeum marinum Huds.

Hordeum secalinum Schreb.

Hottonia palustris L.

Huperzia selago (L.) Schrank & Mart.

Hypericum humifusum L.

Hypericum maculatum Crantz

Hypericum pulchrum L.

Hypochoeris glabra L.

Illecebrum verticillatum L.

Inula britannica L.

1Solepis fluitans (L.)R.Br.

Juncus anceps Laharpe

Dicht havikskruid �9 Gemeines Habichtskraut EX Almindelig Hegeurt

Veenreukgras Duft-Mariengras VU Vellugtende Festgr~s

Zeegerst Strand-Gerste CR WAT, AGR Strand-Byg

Veldgerst Roggen-Gerste * Eng-Byg

Waterviolier Wasserfeder CR WAT Vandrellike

Dennewolfsklauw CR EUT

Otteradet Ulvefod

Liggend hertshooi Liegendes Hartheu EX? WAT, AGR Dv~rg-Perikon

Kantig hertshooi Kanten-Hartheu Kantet Perikon

Fraai hertshooi iSch0nes Johanniskraut Smuk Perikon

E N

Glad biggekruid HAB, EUT, Kahles Ferkelkraut EN AFF Glat Kongepen

Grondster QuirUge Knorpelmiere EX WAT Bruskbaeger

Engelse alant Wiesen-Alant MU WAT Sol~je-Alant

Vlotende bies WAT, Flutende Schuppensimse EN AGR, EUT, Flydende Kogleaks HAB

DuJnrus Zweischneidige Binse VU WAT Sand-Siv

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

EX

* E N *

EN EX EX

* V U * *

EX *

SU CR

I

EX EX (*) EX?

SU * SU *

SU VU CR -

CR EN VU SU

EX

VU VU *

VU CR SU

�9 * E N *

Page 17: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the W a d d e n Sea Area

F ,

Juncus balticus Willd.

I L

Noordse rus Baitische Binse Klit-Siv

Kleine knolrus ZwiebeI-Binse Opret Siv

R e d List Threats

( t r i la tera l )

V U

S U ?

Zeegroene rus Blaugr(Jne Binse Bl&gr& Siv

Juncus bulbosus ssp. kochfi ( F . W , S c h u l t z ) R e i c h g .

IL

Koprus WAT, Juncus capitatus weigel Kopf-Binse CR

Fin Siv AGR, EUT i

Juncus inflexus L. VU WAT

Zeerus Juncus maritimus L a m . Strand-Binse

Strand-Siv

Dwergrus Juncus pygmaeus L. Zwerg-Binse

Dvaerg-Siv

* AGR

EN WAT, AGR, EUT

Lamsoor GewOhnlicher Strandflieder Taetblomstret Hindebaeger

Jeneverbes Juniperus communis L Gemeiner Wacholder CR AGR

Aim Ene

Smal fakkelgras Koeleria arenaria (Dum.)Conert Sand-Schillergras *

Klit-Kambunke

Strandsla Lactuca tatarica ( L . ) Tataren-Lattich SU C A M e y e r Strand-Salat

Knollathyrus Lathyrus linifolius (Reichard) Berg-Platterbse EN B a s s l e r

Krat-Fladbaelg

Zeelathyrus Lathyrus maritimus ( L . ) Strand-Platterbse VU WAT, DIS Bigel. (=L. japonicus Willd) Strandaert

Veldkruidkers Lepidium campestre (L.) R.Br.. 1 Feld-Kresse VU

Salomons Lysestage

Limonium vulgare Mill. *

Limosella aquatica L. Slijkgroen

Schlammkraut Dyndurt

E N

59

Status of threat in the subre- gions of the Wadden Sea Area!

NL Nds SH DK

* C R *

L, 1

s u - ? I

EX EX CR VU

VU

* S U *

EN CR *

CR CR *

* V U *

SU

SU

SU

EN

i i

EN * *

i i

v u * (*)

i i

VU * *

i i

WAT CR VU CR

Page 18: Red List of vascular plants of the Wadden Sea Area

60 Vascular Plants of the Wadden Sea Area

R e d List Threats (trilateral)

Geelhartje . Linum catharticum L. Purgier-Lein VU AGR, WAT

Vild Her i i

Groenknolorchis Liparis Ioeselii (L.) L.CMRichard Sumpf-Glanzkraut EN WAT

Mygblomst i ]

Kleoine keverorchis Listera cordata (L) RBr. Kleines Zweiblatt EN

Hjertebladet Fliglaebe i i

Glad parelzaad Lithospermum officinale L. Echter Steinsame SU

Leege-Stenfre i i

Oeverkruid Littorella uniflora (L.)Asch. See-Strandling EN WAT, EUT

Strandbo I i

~Smalle rolklaver Lotus glaber MiU. ,Salz-Hornklee *

Smalbladet Kaellingetand i i

Moeraswolfsklauw Lycopodiella inundata Gemeiner Moorb~rlapp VU WAT ( L . ) H o l u b Liden Ulvefod

iJ

Grote wolfsklauw Lycopodium clavatum L. Keulen-B~rlapp VU AGR, HAB,

Aim. Ulvefod EUT, AFF i i

Moeraswederik Lysimachia thyrsiflora L. Straul~-Gilbweiderich EN? WAT

Dusk-Fredles i i

Malrove Marrubium vulgare L.. 1 Gemeiner Andorn EX? AGR

Kransburre

Waterdrieblad Menyanthes trifoliata L. Dreibl~ttriger Fieberklee VU WAT

Bukkeblad

Muizestaart Myosurus minimus L.. 1 M~luseschw~nzchen EN? AGR

Musehale

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

* VU VU *

EN CR

* E X

SU

EN EN CR

* V U *

* VU EN *

SU VU EN SU

EX VU (*) *

EX - ?

VU EN VU *

CR VU (*) (*)

Teer vederkruid Myriophyllum altemiflorum Wechselbl0tiges Tausend- EN WAT, EUT VU oc. blatt

H~r-Tusindblad i1

Beenbreek Narthecium ossifragum (L.) Beinbrech EN WAT H u d s .

Benbr~ek

E X - *

CR *

Page 19: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the Wadden Sea Area 61

k ~

R e d List ( t r i l a t e r a l ) Threats

Vroege ogentroost Odontites litoralis (Fries) Fries Salz-Zahntrost EN WAT

Strand-R~ttop i1_

i Zilt torkruid i Oenanthe lachenalii Wiesen-Pferdesaat EN I C.C.Gmelin.

Eng-Kfaseskaerm IL

Oenothera ammophila Focke Zandteunisbloem =O.oakesiana (A.Gray) Denen-Nachtkerze *

R o b b i n s e x . S . W a t s o n & C o u l t Klit-Natlys L ,

Addertong Ophioglossum vulgatum L Gew0hnliche Natternzunge VU AGR, WAT

!Aim. Slangetunge L

Status of threat in the subre- I giens of the Wadden Sea Area

NL Nds SH DK

* C R *

* E N C R

* * * V U

* E N E N *

I Harlekijn Kleines Knabenkraut/Sa- Orchis morio L. CR WAT, AGR VU EX lep Knabenkraut Salep-Gegeurt

Koningsvaren Osmunda regalis L. K0nigsfarn VU WAT

Kongebregne

Dunstaart Parapholis strigosa (Dumort.) Gekr0mrnter DQnnschwanz VU AGR H u b b .

Spidshale

Parnassia Pamassia palustris L Sumpf-Herzblatt VU WAT

Aim. Leverurt

Moeraskartelblad Pedicularis palustris L. Sumpf-Lt~usekraut EN WAT

Eng-Troldurt

Heidekartelblad Pedicutaris sylvatica L. Wald-L~tusekraut EN AGR, EUT,

Mose-Troldurt HAB, WAT

Waterpostelein Peplis portula L. Sumpfquendel MU WAT, HAB

Vandportulak

Melkeppe Sumpf-Haarstrang, Olse- nich Kaer-Svovlrod

Peucedanum palustre (L.) Moench

Phleum arenarium L. Zanddoddegras Sand-Lieschgras Sand.Rottehale

Pilvaren Pillenfarn PiUedrager

* V U V U S U

* VU VU *

VU CR *

VU CR CR *

VU EX VU *

* VU EN SU

SU *

* V U *

CR EX CR VU HAB, WAT, Pilularia globulifera L. CR AGR, EUT

Page 20: Red List of vascular plants of the Wadden Sea Area

62 Vascular Plants of the Wadden Sea Area

i i . . . . . . . . . . i .......

Red List (trilateral)

I I I

Pimpinella saxifraga L.

II

Pinguicula vulgaris L.

II

Platanthera bifolia (L.) L.C.M.Richard

Poa palustris L. 3

i i

Polygala serpyllifolia Hose

ii

Polygala vulgaris L.

i i

Polygonatum odoratum (Mill) D r u c e

Kleine bevernel Kleine Pimpinelle Aim. Pimpinelle

Threats

Vetblad Echtes Fettkraut I Aim. Vibefedt

I

CR

Welriekende nachtorchis Zweiblattrige Waldhyazint- EN he Bakke-G~gelilje

i

Moerasbeemdgras Sumpf-Rispe Stortoppet R~pgraes

Liggende vleugeltjesbloem QuendeI-Kreuzbl0mchen EN Spaed Maelkeurt

i

Gewone vleugeltjesbloem GewOhnliche Kreuzblume VU Aim. Maelkeurt

i

Welriekende salomonsze- gel Duftende WeiSwurz, Sa- lomonssiegel Kantet Konval

SU

EUT, AGR, HAB

i i i

Kleine duizendknoop Polygonum minus Huds. Kleiner KnOterich *

Liden Pileurt i i i

Polygonum oxyspermum Mey- Zandduizenknoop e r & B u n g e s s p . rai i ( B a b . ) Sand-KnOterich EN W e b b & C h a t e r Sand-Pileurt

i

Weegbreefonteinkruid Potamogeton coloratus Gef~rbtes Laichkraut SU Hornem.

Vejbred-Vandaks ii i

Ongelijkbladig fonteinkruid Potamogeton gramineus L. Gras-Laichkraut EN WAT, EUT

Graesbladet Vandaks

Doorgroeid fonteinkruid Durchwachsenes Laich-

= EN WAT Potamogeton perfo/iatus L. kraut

Hjertebladet Vandaks = =

Duizendknoopfonteinkruid WAT, EUT, Potamogeton po/ygonifo/ius KnOterich-Laichkraut VU P o u r r . AGR

Aflangbladet Vandaks

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

SU * * *

CR EX SU

VU EN CR *

S U * ( * ) *

EX *

EUT VU VU * *

SU

s u * ( * ) *

CR VU

SU

EN EN SU *

EX - SU *

* VU VU *

Page 21: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the Wadden Sea Area 63

Red List

( t r i la teral ) Threats

Haarfonteinkruid Potamogeton trichoides Haarbl~ttriges Laichkraut VU WAT Chain. & Schlechtend.

HArfin Vandaks

Viltganzerik Potentilla argentea L. Silber-Fingerkraut VII ?

S~alv-Potentit

Bleek kweldergras Puccinellia capi l larJs Haar-Salzschwaden * (Liljeblad) Jansen

Slap Annelgraes

Klein wintergreen Pyrola minor L. Kleines WintergrOn VU ?

Liden Vintergr~n

Rend wintergreen Pyrola rotundifolia L Rundblattriges Wintergr0n EN WAT

Klit-Vintergron

Dwergvlas Radiola linoides R e t h Zwerg-Lein VU WAT

Tusindfr~

Gulden boterbtoem Ranunculus auricomus L Gotd-Hahnenful~ SU

Nyrebladet Ranunkel

Zilte waterranonkel Brackwasser-Wasser-

Ranunculus baudotii Godr. hahnenfur~ VU EUT

Strand-Vandranunkel

Klimopwaterranonkel Ranunculus hederaceus L. Efeubl~ttriger Hahnenfui~ EX? WAT

Vedbend-Vandranunkel

Driedelige waterranonkel Dreiteiliger Wasserhahnen- FX WAT

Ranunculus tripartitus o c fui5

Grote ratelaar Rhinanthus angustifolius Gro&er Klappertopf VU WAT, EUT C.CGmelin (Coll.) Stor Skjaller

Rhinanthus angustifolius ssp. VII? halophilus (U.Schneider) See Strand-Skjaller

Kleine ratelaar Rhinanthus minor L. Kleiner Klappertopf VII AGR

Liden Skjaller

Witte snavelbies Rhynchospora alba Wei~es Schnabelried CR? WAT (L.) Vahl Hvid Naebfro

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

VU *

EX * * *

SU SU * *

VU VU VU

VU EN CR *

VU EN EN *

SU

* EN V U *

EX (*)

EX

VU VU VU *

(*) v u

* V U V U *

EX (*) *

Page 22: Red List of vascular plants of the Wadden Sea Area

64 Vascular Plants of the Wadden Sea Area

Red List (trilateral) Threats

I

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH

Bruine snavelbies Rhynchospora fusca Braunes Schnabelried (3R WAT EX EX (L.) Ait. ill.

Brun N~ebfre I I I'

Duinroosje Rosa pimpinellifolia L. D0nen-Rose MU * EN

Kilt-Rose II I

Spiraalruppia Ruppia cirrhosa (Petagna) Schraubige Salde EN EUT, WATi VU SU

l Grande Langstilket Havgraes I

Snavelruppia Ruppia maritima L. Meeres-Salde EN EUT, HAB, VU EN

WAT Aim. Havgraes I

Sierlijke vetmuur Sagina nodosa (L.) F e n z l Knotiges Mastkraut MU WAT * EN

Knude-Firling I

Priemvetmuur Sagina subulata (Swartz) Pfriemen-Mastkraut CR WAT - C . P r e s l .

SyI-Firling I

Pijlkruid Sagittaria sagittifolia L. Pfeilkraut * SU

Aim. Pilblad i

Salix hastata L. SpieE,-Weide * WAT Spyd-Pil

I

Waterpunge Samolus valerandi L Salz-Bunge VU WAT *

Samel i

Stekende bies Schoenoplectus pungens Amerikanische Teichsimse CR WAT CR ( V a h l ) P a l l a

i

Driekantige bies Schoenoplectus triqueter !Dreikantige Teichsimse EN HAB, WAT - (L.) Palla

Bastaardbies Schoenoplectus X carinatus Tide-Simse EN Smith

I I

Knopbies Schoenus nigricans L. Schwarzes Kopfried EN i WAT VU

Sort Skaene I

EN

CR

EN

EN

EN

VU Overblijvende hardbloem

Scleranthus perennis L. Ausdauernder Kn~uel VU Flerw Knavel

VU

CR

CR

VU

CR

SU

EX

CR

SU

EX

VU

DK

II

II

I I

I I

I I

I !

i1

, i

i i

i i

i t

Page 23: Red List of vascular plants of the Wadden Sea Area

Vascular Plants of the W a d d e n Sea Area 65

Red List (trilateral) Threats

Kleine schorseneer Scor'zonera humilis L. Niedrige Schwarzwurzel EN EUT, AGR

Lay Skorsoner

Klein glidkruid Scutellaria minor Huds. Kleines Helmkraut EN WAT, EUT

:Liden Skjolddrager

Wit vetkruid Sedum album L Wei6e Fetthenne SU

Hvid Stenurt

Viltig kruiskruid

Senecio erucifolius L Raukenblattriges Greis- EN WAT kraut Smalfliget Brandbaeger

Nachtsilene Silene nutans L. Nickendes Leimkraut CR HAB, AGR,

EUT Nikkende Limurt

Oorsilene Silene otites (L . )Wibel OhrlOffeI-Leimkraut VU HAB, AGR

Klit-Limurt

Zwartmoeskervel Smymium olusatrum L. SU

Makedonisk Persille

Kleinste egelskop Sparganium natans L. CR WAT

Spaed Pindsvineknop

Heidespurrie Spergula morisonii Boreau FrQhlings-Spergel SU

V&r-Spergel

Akkerandoorn Stachys arvensis (L) L. Acker-Ziest FN? AGR

Ager-Galtetand

Waterrnuur Stellaria aquatica (L)scop. Wasserdarm CR

Kl~vkrone

Priemkruid Subularia aquatica L. ~friemenkresse EX

Sylblad

Schraallandpaardebloem Taraxacum celticum agg. MU AGR, WAT

Eng-Malkebette

Moeraspaardebloem Taraxacum palustre agg . ,Sumpf-Kuhblume VU WAT, AGR

Smalbladet Malkebotte

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK Jl

EN

i i

EN

u

SU

i i

EX - *

CR

VU VU CR SU

SU

EX *

SU

CR EN (*)

SU EX

EX

VU

VU VU ?

Page 24: Red List of vascular plants of the Wadden Sea Area

66

Thalictrum flavum L.

Thalictrum minus L.

Thymus pulegioides L.

Torilis nodosa (L.)Gaertn.

Trichophorum cespitosum ( L ) H a r t m a n

Trientalis europaeus L.

Trifofium omithopodioides L

Trifolium striatum L

Triglochin palustre L

Tuberaria guttata (L.) Fourr.

1 Ulex europaeus L..

Utricularia australis R.Br,

Utricularia minor L.

Vaccinium macrocarpon Ait.

Vascular Plants of the Wadden Sea Area

Poelruit Gelbe Wiesenraute GuI-Frostjerne

R e d L i s t Threats

( t r i l a t e r a l )

~ : R ?

Kleine ruit Kleine Wiesenraute VU Sand-Frostjerne

Grote tijm Gemeiner Thymian SU Bredbladet Timian

Knopig doornzaad KnaueI-Kiettenkerbel MU AGR Knudret Randfro

Veenbies I Rasige Haarsimse CR WAT Tue Kogleaks

I Zevenster I Europaischer Siebenstern * Skovstjeme

Vogelpootklaver Vogetfur~artiger Klee *

Gestreepte klaver Streifen-Klee SU Stribet Klover

Moeraszoutgras Sumpf-Dreizack VU AGR Kaer-Trehage

Gevlekt zonneroosje Geflecktes Sandr(~schen CR ? En&rig Stensoloje

Gaspeldoorn Stechginster EN HAB Tornblad

Loos blaasjeskruid S0dlicher Wasserschlauch VU WAT, EUT Slank Blaererod

Klein blaasjeskruid Kleiner Wasserschlauch CR WAT Liden Bl~rerod

Grote veenbes Groi%fr0chtige Moosbeere * Storfrugtet Tranebaer

...... t . . . . . . .

Status of threat in the subre- gions of the Wadden Sea Area

NL Nds SH DK

(*) CR *

* V U E X *

SU

VU EN VU

EX EN *

* SU

* SU

SU SU

* EN VU *

CR CR

VU EN EN (*)

VU VU *

EX - *

* S U

Page 25: Red List of vascular plants of the Wadden Sea Area

V a s c u l a r P l a n t s o f t h e W a d d e n S e a A r e a 6 7

Status of threat in the subre- Red List

; (tr i lateral) Threats gions of the Wadden Sea Area

NL Nds SH DK

Blauwe bosbes Vaccinium myrtillus L. Heidelbeere VU SU * CR

Bl&baer

Rode bosbes 'Vaccinium vitis-idaea L. Preiselbeere EN * * CR

Aim Tyttebaer

Duinviooltje Viola tricolor ssp curtisii D~Jnen-Stiefm0tterchen * DIS, HAB * * VU * (Forst.) S y m e Klit-Stedmoderblornst

Eekhoorngras Vulpia bromoides (L.) SFGray Trespen-Federschwingel EN AGR, EUT VU CR CR

Langstakket Vaeselhale

Oeverstekelnoot Xanthium albinum ssp. albi-

i EIb-Spitzklette SU SU Rum (Widder) H. Scholz sstr

Zannichellia palustris L. ssp. Brede zannichellia polycarpa (Rchb) K Richter Sumpf-Teichfaden EX WAT EX

Stor Vandkraus

Groot zeegras EXL, EUT, . Zostera marina L. GewOhnliches Seegras VU PAR EN EN VU

Aim. Baendeltang

Klein zeegras Zostera noltii Hornem :Zwerg-Seegras VU EXL, EUT SU EN VU *

I Dvaerg-Baendeltang

I n t r o d u c e d in D e n m a r k . A n t h r o p o c h o r o u s in D e n m a r k . A n t h r o p o c h o r o u s in t h e W a d d e n S e a a r e a .

S t a t u s o f t h r e a t : E X = E x t i n c t ; C R = C r i t i c a l ; E N = E n d a n g e r e d ; V U = V u l n e r a b l e ; S U = S u s c e p t i b l e ; " = n o t e n d a n - g e r e d ; ? = s t a t u s of e n d a n g e r m e n t is n o t e x a c t l y k n o w n ; - = s p e c i e s d o e s n o t o c c u r ; -I?) = s p e c i e s p r o b a b l y d o e s n o t o c c u r ; (*)= it is u n k n o w n if o r if n o t t h i s s p e c i e s o c c u r s . T h r e a t s : A F F = a f f o r e s t a t i o n ; A G R = a g r i c u l t u r e ; CLI = c l i m a t i c c h a n g e ; DIS = d i s t u r b a n c e ; E U T = e u t r o p h i - c a t i o n ; E X L = e x p l o i t a t i o n of l i v i n g r e s o u r c e s ; E X M = m i n e r a l e x p l o i t a t i o n ; F O R = f o r e s t r y ; H A B = loss of h a b i t a t ; P A R = p a r a s i t e s ; P O L = p o l l u t i o n ; W A T = w a t e r r e g u l a t i o n .

F o r m o r e d e t a i l e d d e s c r i p t i o n s s e e t h e g e n e r a l i n t r o d u c t i o n to t h e R e d Lists .