Final Task of Zoogeografi Subject
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Transcript of Final Task of Zoogeografi Subject
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1. Factor that leading the appearance of new species!a. Natural selection, is the process by which certain heritable traitsthose that
make it more likely for an organism to survive and successfully reproduce
become more common in a population over successive generations. It is a key
mechanism of evolution. It is the differences in survival and reproduction
among individuals in a population because of their interaction with the
environment. In other words, some individuals possess alleles (genotypes) that
generate traits (phenotypes) that enable them to cope more successfully in
their environment than other individuals. The more successful individuals
produce more offspring. Superior traits are adaptations to the environment
and increase an individuals fitness, or relative ability to survive and leave
offspring. When the environment favors a trait, that is, when a trait increases
the survival of its bearer, selection is said to act for that trait. In contrast,
selection is said to act against unfavorable traits. Favorable traits are adaptive,
while unfavorable traits are maladaptive.
b.Polyploidy is the possession of more than the normal two sets ofchromosomes found in diploid (2n) cells. Polyploidy often occurs in plants
(and occasionally animals) where triploid (3n), tetraploid (4n), and higher
ploidy chromosome numbers are found.A quite different method by which
new species can appear is by polyploidy- the doubling of the whole set of
chromosomes in the nucleus of the developing egg or seed, so that each
chromosome automatically has an identical partner. Polyploidy species are
often larger than the original parent type, and also more hardy and vigorous;
many weeds are polyploidy. Only a few animals groups fall into this category
(e.g. turbellarians, lumbricid eartworms, and weevils). This new species,
formed by natural polyploidy, has been extremely successful and has spread
around the world, often creating problems for shipping by forming mats of
vegetation within which sediments are deposited, and so contributing to the
silting up to estuaries. Polyploidy occurs because of nondisjunction of all
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chromosomes during meiosis, producing two viable diploid gametes and two
sterile gametes with no chromosomes. A tetraploid zygote can be established
when a diploid sperm fertilizes a diploid egg. Since normal meiosis in the
tetraploid individual will continue to produce diploid gametes, reproductive
isolation with other individuals in the population (and thus speciation) occurs
immediately in a single generation.
c. Speciation (isolation), a species is usually defined as a group of individualscapable of interbreeding. Speciation, the formation of new species, occurs by
the following processes,
a) Allopatric speciation begins when a geographic barrier divides apopulation so that interbreeding between the two resulting populations is
prevented. Common barriers include mountain ranges or rivers, but any
region that excludes vital resources, such as a region devoid of water, a
burned area devoid of food, or an area covered with volcanic lava, can act
as a barrier because individuals cannot survive its crossing. Once
reproductively isolated by the barrier, gene frequencies in the two
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populations can diverge due to natural selection (the environments may be
slightly different), mutation, or genetic drift. If the gene pools sufficiently
diverge, then interbreeding between the populations will not occur if the
barrier is removed. As a result, new species have formed. For example, a
German shepherd dog and a greyhound that appear just as different from
one another but both are members of same species. The essential
difference is that under normal conditions in the wild, they will mate
together (due to artificial selection by human). Biologist commonly define
the species as consisting of a group of populations whose members can all
breed together to produce offspring that are fully fertile, but that in the
wild do not do so with other such groups.
b) Sympatric speciation is the formation of new species without the presence of a geographic barrier. For example, two different species of
lacewing insect are found in the cold temperate and boreal of North
America. Chrysopa carnea, is found in grasslands, meadows, and on
deciduous trees, but only rarely on conifer trees; it is light green in color.
C. downesi is found only on conifer trees, and is a very dark green. Apart
from their different color, the two species are nearly indistinguishable
morphologically and, although they will interbreed in the laboratory, they
do not in nature. It was originally suggested that the common ancestor of
these two living species lived in both habitats. A simple genetic mutation
could then have led to the appearance of the dark green coloration in some
individuals. This coloration advantage for individuals in the coniferous
habitats and disadvantage in the grassland/meadow/ deciduous tree
habitat.It has subsequently been shown that the two species also differ in
their complex mating calls, and it has been suggested that this, rather than
their coloration, was the original basis for the separation of the two species
c) Adaptive radiation (mutation) is the relatively rapid evolution of manyspecies from a single ancestor. It occurs when the ancestral species
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colonizes an area where diverse geographic or ecological conditions are
available for colonization. Variants of the ancestral species diverge as
populations specialize for each set of conditions.
2. Explain the evolution process without discuss about the fossil!Until recently, it was thought that evolution took place too slowly for it to
be detectable over the timescale of scientific studies of living organisms, so that
it could only be detected in the fossil record. However, it is now clear that this
perception was wrong. For many years, British workers Peter and Rosmary Grant
have studied the finches on the tiny Galapagos island called Daphne Major, only
34 ha (84 acres) in area and 8 km (5 miles) from the nearest large island. The
finches, which belong to the genus Geospiza, live in a variety of environments
and use a number of sources of food. Some, with narrow bills, live on the ground
and feed on insects, snails, etc. some, with strong, heavy bills, live in the cactus
thickets and feed on the cactus seeds, flowers and pollen. However, most live in
the bushes and trees and feed on the seeds, nuts and fruits of some dozen species
of plant.
G. magnirostris, have large bills; G. fuliginosa, have small bills; G. fortis,
have bills of intermediate size. When food is plentiful, all these three species
prefer to eat the softest types of seed. In drier seasons, there are fewer of these
favorite seeds, because smaller plants, which tend to wither and die during a
drought, produce them. As a result, each species has then to become more
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specialized, spending more of its time feeding on those seed to which its beak
size is be adapted. G. magnirostris whit their biggest and strongest beak (best for
cracking the hardest seeds) can gathering food faster than G. fortis for feed on
same seed, but not all ofG. fortis can eat that seed while, G. fuliginosa has to
feed on smaller, softer seeds. This provides a very precise example of the nature
and results of the correlation between morphology (beak size) and ecology
(availability of different types of food).
Two of the extreme years in the century- both the driest year, 1977, and
the wettest year, 1983 had immense effects on the finches of the island. The
drought of 1977, make the plants failed to flower and set seed so that it make
shortage for finches food. The first effected to the new generation of finches,
make fledglings of 1977 died before they were 3 months old. But the drought
also hit the adults and the death hit harder at the finches that cold only eat softer
seeds, number ofG. fortis dropped by 85% from 1200 birds down to 180 but
suffered worst to G. fuliginosa, it number from a dozen to only a single bird.
Finally, the drought had been kindest to birds with big beaks The average
size of the G. fortis birds that survived was 5.6% greater than the average size of
the 1976 population, and their beaks were correspondingly longer (and stronger)-
11,07 mm long and 9,96 mm deep, compared with 1976 averages of 10,68 and
9,42 mm.
In 1983 years, that was the years of the strongest 20th century El Nino
event; the rainfall was ten times the previously known maximum, and as result,
the plants grew rampantly. That time, small seeds predominated-they formed
up to 80% of the total mass of seeds. That was partly because the smaller plants
grew luxuriously so they can produced many more seeds, while the growth of
Tribulus had been hampered by smothering vines. The availability of seed (small
seed), make number ofG. fortis having increased by more than four times. Now
it was selecting smaller birds, with smaller beaks, more suited to eating the
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plentiful smaller seeds. The main information from that is the conditions and
selection, can oscillate violently.
A total reversal of the selection pressures within 6 years-a time span
that would have been totally invisible in the fossil record, had there been
one. Therefore, the fluctuation in relative numbers of finches that the Grants
have observed in response to the changes in the availability of food of different
types, have not been between different, independent species. This records show
precisely what Darwins explanation had predicted, and what biologist have long
accepted. Each population is adapted to its existing rnvironment, withing a
pattern of the availability of food and of competition with other populations. But
that adaptation is not immutable: it cannot afford to be, for the environment itself
is fluid and changeable. Therefore, as from year to year, the environment
makes new demands and provides new opportunities, and it will change in
harmony with those demands and opportunities. Adaptation by evolution is
never far behind our efforts to control the biological world.
Darwin yang berasumsi bahwa evolusi merambat selambat gletser dan
hanya dapat diamati dalam catatan fosil. Pada 1973, Peter dan Rosemary Grant,
sekarang di Princeton University, memulai pengamatan tahunan terhadap
populasi pipit di pulau kecil Daphne Major di Galpagos. Mereka segera
menemukan bahwa pada kenyataannya pipit berevolusi dari tahun ke
tahun, saat kondisi pulau itu beralih dari basah ke kering dan basah
kembali. Misalnya, Daphne Major awalnya mengamati hanya dua burung pipit
tanah yang berbiak secara teratur, salah satunya adalah burung pipit tanah sedang
(G. fortis) yang makan bebijian kecil. Ketika kemarau parah melanda pulau itu
pada 1977 dan bebijian kecil menjadi langka, burung pipit sedang terpaksa
beralih memakan bebijian yang lebih besar dan lebih keras. Burung yang
berparuh lebih besar lebih mampu mencari makan dan dapat bertahan hidup
untuk menurunkan sifat bawaan itu kepada anaknya. Perubahan lain terjadi
setelah pesaing tiba pada 1982: burung pipit tanah besar (G. magnirostris) yang
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juga makan bebijian besar yang keras. Selama bertahun-tahun, kedua spesies itu
hidup berdampingan dan pada 2002, kedua jenis menjadi sangat berlimpah. Lalu
kemarau tiba dan pada 2005 tinggal 13 pipit tanah besar dan 83 pipit tanah
sedang yang masih hidup. Yang luar biasa, alih-alih menyesuaikan diri
terhadap kemarau dengan makan bebijian yang lebih besar seperti yang
dilakukan 28 tahun sebelumnya, pipit sedang yang menyintas mengalami
pengecilan ukuran paruh yang mencolok karena saat bersaing dengan
sepupu mereka yang lebih besar, pipit sedang berjuang mencari ceruk
hidup dengan makan bebijian yang sangat kecil. Burung pipit yang berparuh
lebih kecil bukanlah spesies pipit yang baru, tetapi Peter Grant menduga hanya
diperlukan beberapa episode semacam itu sampai terbentuk spesies baru yang
tidak mau bereproduksi dengan spesies asalnya.
3. Why there are many variations in a species!In order for natural selection to operate, there must be variation among
individuals in a population. Indeed, considerable variation exists in nearly all
populations. The variation arises from or is maintained by the following
mechanisms:
1. Mutationsprovide the raw material for new variation. All other contributionsto variation, listed below, occur by rearranging existing alleles into new
combinations. Mutations, however, can invent alleles that never before existed
in the gene pool.
2. Sexual reproduction creates individuals with new combinations of alleles.These rearrangements, orgenetic recombination, originate from three events
during the sexual reproductive process, as follows.
y Crossing over, or exchanges of DNA between nonsister chromatids ofhomologous chromosomes, occurs during prophase I of meiosis.
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y Independent assortment of homologues during metaphase I createsdaughter cells with random combinations of maternal and paternal
chromosomes.
y Random joining of gametes during fertilization contributes to thediversity of gene combinations in the zygote.
3. Diploidy is the presence of two copies of each chromosome in a cell. In theheterozygous condition (when two different alleles for a single gene locus are
present), the recessive allele is hidden from natural selection. As a result,
more variation is maintained in the gene pool.
4. Outbreeding, or mating with unrelated partners, increases the possibility ofmixing different alleles and creating new allele combinations.
5. Balanced polymorphism is the maintenance of different phenotypes in a population. Often, a single phenotype provides the best adaptation, while
other phenotypes are less advantageous. In these cases, the alleles for the
advantageous trait increase in frequency, while the remaining alleles decrease.
However, examples of polymorphism (two or more different phenotypes) are
observed in many populations.4. Explain the controversy of evolutionary theory!
There is now a vast amount of evidence for Darwins explanation of
evolution by natural selection. Nevertheless, controversy still exists about some
details of the circumstances in which new species evolve or the rate at which this
happens.
1. Some biologists believe that evolutionary change normally takes place at asteady, gradual rate. Others instead believe that, even if genetic alterations
gradually accumulate within a population, this may not be reflected in
detectable morphological or physiological changes until they are so
numerous as to shift the balance ofthe whole genotype. Example: the finches
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were evolution from year to year, when the condition of Galapagos island
change from wet to dry and become wet again.
2. It is also difficult to isolate such underlying patterns from the more directeffects of the environment. For example: a study of the fossil shells of
gastropod mollusks that lived in northern Kenya over the last few million
years shows long periods during which their structure and size remain
unchanged, interrupted by shorter periods (5000-50.000 years) during which
they changed rapidly. This was interpreted as an example of punctuated
equilibrium.
3. Darwins suggests that it is the fittest that survive because those are the onethat survives, of course (the catchphrase survive of the fittest was not
Darwins but by Herbert Spencer).
Teori evolusi menyatakan bahwa makhluk hidup di muka bumi tercipta sebagai
akibat dari peristiwa kebetulan dan muncul dengan sendirinya dari kondisi alamiah.
Berbagai cabang ilmu pengetahuan seperti paleontologi (cabang geologi yang mengkaji
kehidupan pra-sejarah melalui fosil), genetika, biokimia dan biologi molekuler telah
membuktikan bahwa tak mungkin makhluk hidup tercipta akibat kebetulan atau muncul
dengan sendirinya dari kondisi alamiah. Sel adalah struktur paling kompleks yang
pernah ditemukan manusia. Ilmu pengetahuan modern mengungkapkan bahwa satu sel
hidup saja memiliki struktur dan berbagai sistem rumit dan saling terkait, yang jauh
lebih kompleks daripada sebuah kota besar. Struktur kompleks seperti ini hanya dapat
berfungsi apabila masing-masing bagian penyusunnya muncul secara bersamaan dan
dalam keadaan sudah berfungsi sepenuhnya. Jika tidak, struktur tersebut tidak akan
berguna, dan semakin lama akan rusak dan musnah. Tak mungkin semua bagian
penyusun sel itu berkembang secara kebetulan dalam jutaan tahun, seperti pernyataan
teori evolusi.