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CYNTHIA M. BEALL

CURRICULUM VITAE

BUSINESS ADDRESS: Department of Anthropology

Case Western Reserve University

238 Mather Memorial

Cleveland, Ohio 44106

Telephone (216) 368-2264; fax (216) 368-5334

Internet cmb2@case.edu

EDUCATION: 1970 University of Pennsylvania B.A., Biology

1972 The Pennsylvania State University M.A., Anthropology

1976 The Pennsylvania State University Ph.D., Anthropology

FIELDS OF SPECIALIZATION:

Human Biology/Human Adaptability: high altitude adaptation; growth, development and aging; human

variability; genetic, environmental, and socio-cultural influences on human biology; evolutionary medicine;

global health; Andean Plateau (Bolivia, Peru), Tibetan Plateau (Nepal, Tibet), Simien Plateau (Ethiopia),

Mongolia.

ACADEMIC APPOINTMENTS:

2010- Distinguished University Professor, Case Western Reserve University

2005- Adjunct Staff, Department of Pathobiology, Lerner Research Institute, The Cleveland Clinic Foundation

(secondary appointment)

2004- Research Associate in Physical Anthropology, Cleveland Museum of Natural History

(secondary Appointment)

2003- Professor of Global Health and Diseases, School of Medicine, Case Western Reserve University

(secondary appointment)

1995- Professor of Anatomy, School of Medicine, Case Western Reserve University

(secondary appointment)

1994- S. Idell Pyle Professor of Anthropology, Case Western Reserve University.

1987-94 Professor, Department of Anthropology, Case Western Reserve University.

1982-87 Associate Professor (with tenure), Department of Anthropology, Case Western Reserve

University.

1976-82 Assistant Professor, Department of Anthropology, Case Western Reserve University.

HONORS:

2011 John Simon Guggenheim Fellow

2009 Recipient, Franz Boas Distinguished Achievement Award, Human Biology Association

2001 Member, American Philosophical Society

1997 Fellow, American Association for the Advancement of Science

1996 Member, National Academy of Sciences

CURRENT EDITORIAL BOARDS:

American Journal of Human Biology, 2002 -

Journal of High Altitude Medicine and Biology, 1999 -

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CURRENT PROFESSIONAL SERVICE:

2005 – Member, Steering Committee, Section H Anthropology, American Association for the Advancement of

Science (AAAS)

2007 - Member, Temporary Nomination Group for Foreign Associates from underrepresented countries,

Class V (Social Sciences), National Academy of Sciences.

2010 – 2012 Member, proposal reviewing panel, Wenner-Gren Foundation for Anthropological Research

2010 – Chair-elect, Chair, Retiring Chair, Section H Anthropology, American Association for the Advancement

of Science (AAAS)

2011 - Chair, Nominating Committee, National Academy of Sciences

2011 -Member of Advisory Council, National Academies’ Division of Behavioral and Social Sciences and

Education (DBASSE), three year term begins 5/2011

2011 – Member, NEScent (National Evolutionary Synthesis Center) Working Group: Infusing Medical

Education with EvolutionaryThinking, chaired by Mark D Schwartz, MD and Peter Ellison, PhD, Co-

Directors

MEMBERSHIP IN PROFESSIONAL SOCIETIES

American Anthropological Association, General and Biological Anthropology sections

American Association for the Advancement of Science

American Association of Physical Anthropologists

Association for Anthropology and Gerontology

Human Biology Association (formerly Human Biology Council)

International Mountain Medicine Society

Sigma Xi, The Scientific Research Council

Society for the Study of Human Biology (U.K.)

SELECTED RECENT INVITED PAPERS/PRESENTATIONS AT CONFERENCES and NAMED

LECTURESHIPS

2012 Pearl Memorial Lecturer, Human Biology Association Annual Meeting, April 10, Portland, ME

2012 Invited speaker, Genetics and adaptation to high altitude. Keystone Symposium, Advances in Hypoxic

Signaling: From Bench to Bedside, Feb 14, 2012, Banff, Alberta

2011 Thurston Visiting Professor and Lecturer, Washington University School of Medicine, Department of

Pediatrics, May 12-13, 2011

2011 Invited presentation, Gordon Research Conference Nitric Oxide. Recent human evolution and

adaptation to oxygen shortage. Ventura, CA February 14, 2011.

2010 Plenary Lecture, International Association of Physiological Anthropology, September 15, 2010,

Freemantle, Australia.

2010 Invited presentation, VIII International Congress on Medicine and Physiology of High Altitude,

Arequipa, Peru, August 9, 2010.

2010 Landmark Lecture. Human Evolution at High-altitude – an overview. Invited presentation at the 4th

European Hypoxia Society Meeting, January 29, 2010.

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2009 Candidate Gene Associations with Oxygen Delivery Traits at High Altitude. Invited presentation at

16th International Hypoxia Symposium, March 12, 2009.

2008 Tibetan, Andean and Ethiopian Patterns of Adaptation to High Altitude, UT Southwestern School of

Medicine, Center for Human Genetics, McDermott Center for Human Growth and Development, September 16.

2008 Tibetan and Andean Patterns of Adaptation to High Altitude, Class of 1960 lecture at Williams

College, March 7.

2008 Long-term studies of Tibetan Nomads and Economic Development (with M. C. Goldstein), AAAS

Annual Meeting, Invited paper in session From Global to Local: Impact of Field Research in

Biological Anthropology, organized by Karen Strier, February 17.

2008 Contrasting Adaptations to Hypoxia by Andean, Tibetan, and East Africa Populations. Invited

plenary talk at Keystone Symposium on Molecular, Cellular, Physiological, and Pathogenic

Responses to Hypoxia. January 16.

2007 Adaptive Tissue Blood Flow and Circulating Nitric Oxide Products in Tibetans at High Altitude.

Invited plenary talk at Second International Meeting on the role of Nitrice in Physiology,

Pathophysiology, and Therapeutics, NIH. September 7.

PAST PROFESSIONAL SERVICE:

2011 Chair of the organizing committee for convocation “Thinking Evolutionarily: Evolution Education

Across the Life Sciences; Organized by: Board on Life Sciences, National Research Council, National Academy

of Sciences, October 25-26, 2011

2010 – 2011 Organizer, Symposium on Anthropology and Global Health: integrating genes, biology, and

culture, for the annual meetings of the AAAS, February 2011

2008-09 Co-Organizer, with Peter Robbins, National Evolutionary Synthesis Center, international catalygis

meeting on Human evolution and adaptation to High-altitude Hypoxia, March 19-21, 2009, Durham, N Ca

2008-09 Organizer, Plenary Session for Annual Meetings of the Human Biology Association, The Past 10,000

years of human evolution. Chicago, IL, April 2009,

2003 - 2008 Chair, National Academy of Sciences Board on International Scientific Organizations (BISO),

member 2001 – 2003. The Board on International Scientific Organizations (BISO) oversees many groups whose

effort is to strengthen U.S. participation in international scientific, engineering, and medical organizations.

2005 - 2008 – Executive Board, International Council on Science (ICSU).

ICSU is a non-governmental organization with a global membership that is frequently called upon to speak on

behalf of the global scientific community and to act as an advisor in matters ranging from the environment to

conduct in science.

2002-2007 - Member, Advisory Committee for the Social, Behavioral and Economic Sciences (SBE) Directorate

of the National Science Foundation,

2002-2005 - Councilor, National Academy of Sciences,

2002-2005 - Chair, Section 51 Anthropology, National Academy of Sciences,.

2001-2003 - Chair, U. S. National Committee for the International Union of Biological Sciences, member, 1998-

2003.

1995 – Chair, Electorate Nominating Committee, Section on Anthropology, American Association for the

Advancement of Science (AAAS), member 1993 – 1995.

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1991 – 1995 - President-elect, President, and Past-President. Human Biology Council (now Human Biology

Association). The mission of the Human Biology Association is to advance the understanding human biological

variation.

1989 – 1991 Executive Committee, Human Biology Council (now Human Biology Association)

1984-1986 Nominations Committee, Human Biology Council (now Human Biology Association)

1989 –1992 Executive Committee, American Association of Physical Anthropology (AAPA)

1983-1986 Executive Committee Member, American Anthropological Association (AAA), Biological

Anthropology Unit

1981-1987 Sub-Editor for Bio-Cultural and Physical Aging, Association for Anthropology and Gerontology

(AAGE) Newsletter

1983 Corresponding secretary, Association for Anthropology and Gerontology (AAGE)

Sub-Editor for Bio-Cultural and Physical Aging

PAST EDITORIAL BOARDS:

Annual Review of Anthropology: 2004 – 2009

Yearbook of Physical Anthropology, 1983-1997

Journal of Cross-Cultural Gerontology: founding Co-Editor, with M.C. Goldstein and C. Ikels, 1986-1995

Human Biology 1985-1987, 1988-91, 1997-2008

Journal of Wilderness Medicine, 1992-2002

PUBLICATIONS

THESES:

M.A. Respiratory diseases in Migrant Populations living in a Peruvian Costal Valley, 1972.

Ph.D. The Effects of High Altitude on Growth, Morbidity and Mortality of Peruvian Infants, 1976.

MONOGRAPHS:

1993 Goldstein, M.C. and C.M. Beall. The Changing World of Mongolia’s Nomads. University of California

Press, Berkeley.

1990 Goldstein, M.C. and C.M. Beall. Nomads of Western Tibet: The Survival of a Way of Life. The

Guidebook Company, Ltd., Hong Kong. Co-Published by University of California Press, Berkeley. ISBN 0-520-

07210-3 (German edition 1991 Erst Auflage).

1982 Beall C.M. (editor) Cross-Cultural Studies of Biological Aging. Plenum Press, Oxford. Reprinted from

special issue of Social Science and Medicine, volume 16(2). ISBN 0 08 028946 0.

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PUBLICATIONS IN REFEREED JOURNALS:

2012 Beall, C.M., Laskowski, D., Erzurum, S.C. Nitric oxide in altitude adaptation. Free Radical

Biology and Medicine 52(7):1123-34. Epub 2012 Jan 20.

2011 Janocha, A., Koch, C., Tiso, M., Ponchia, A., Doctor, A., Gibbons, L., ,…, Beall, C.M. Erzurum, S. Nitric oxide during altitude acclimatization. New England journal of medicine, 365(20), 1942-4. doi:10.1056/NEJMc1107887. PubMed ID: 22087700

2011 Beall CM. Genetic changes in Tibet. High Alt Med Biol. 2011 Summer;12(2):101-2. PubMed ID:

21718152

2011 Howes RE, Patil AP, Piel FB, Nyangiri OA, Kabaria CW, Gething PW, Zimmerman PA, Barnadas C,

Beall CM, Gebremedhin A and others. 2011. The global distribution of the Duffy blood group. Nat

Commun 2:266, PubMed ID: 21468018

DOI: 10.1038/ncomms1265

2011 Hancock AM, Witonsky DB, Alkorta-Aranburu G, Beall CM, Gebremedhin A, Sukernik R, Utermann G,

Pritchard JK, Coop G, Di Rienzo A.. Adaptations to climate-mediated selective pressures in humans.

PLoS Genetics 7(4):e1001375, PubMed ID: 21533023

2011 Hoit BD, Dalton ND, Gebremedhin A, Janocha A, Zimmerman PA, Zimmerman AM, Strohl KP,

Erzurum SC, Beall CM. Elevated pulmonary artery pressure among Amhara highlanders in Ethiopia.

Am J Hum Biol. 2011 Mar-Apr;23(2):168-76. doi: 10.1002/ajhb.21130. Epub 2010 Dec 22, Pub MedID:

21319245

2010 Beall CM, Cavalleri GL, Deng L, Elston RC, Gao Y, Knight J, Li C, Li JC, Liang

Y, McCormack M, Montgomery HE, Pan H, Robbins PA, Shianna KV, Tam SC, Tsering N,

Veeramah KR, Wang W, Wangdui P, Weale ME, Xu Y, Xu Z, Yang L, Zaman MJ, Zeng C,

Zhang L, Zhang X, Zhaxi P, Zheng YT. Natural selection on EPAS1 (HIF2alpha)

associated with low hemoglobin concentration in Tibetan highlanders. Proc Natl

Acad Sci U S A. 107(25):11459-64. Epub 2010 Jun 7. PubMed PMID:

20534544; PubMed Central PMCID: PMC2895075.

2010 Hancock AM, Witonsky DB, Ehler E, Alkorta-Aranburu G, Beall C, Gebremedhin A,

Sukernik R, Utermann G, Pritchard J, Coop G, Di Rienzo A. Colloquium paper: human

adaptations to diet, subsistence, and ecoregion are due to subtle shifts in

allele frequency. Proc Natl Acad Sci U S A. 2010 May 11;107 Suppl 2:8924-30. Epub

May 5. PubMed PMID: 20445095.

2010 Beall CM, Strohl KP, Laskowski D, Hutte R, Erzurum SC . Response to “Lowered exhaled

nitric oxide in acute hypobaric than in normobaric hypoxia by T. Hemmingsson and D.

Linnarsson. Respir. Physiol. Neurobiol 169: 74-77”. Respir Physiol Neurobiol. 2010 Feb 1.

[Epub ahead of print], Pub MedID: 20123043

2009 Laskowski D, Beall CM, Dweik R, Strohl KP, Hutte R, Erzurum SC. Response to “Measuring

exhaled nitric oxide at high altitude. Respir. Physiol Neurobiol 167: 292-298”. Respir Physiol

Neurobiol. 2010 Jan 31;170(1):1-2 [Epub 2009 Nov 14.], Pub MedID: 19922816

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2009 Vitzthum VJ, Worthman CM, Beall CM, Thornburg J, Vargas E, Villena M, Soria R,

Caceres E, Spielvogel H. Seasonal and circadian variation in salivary testosterone in rural Bolivian men. Am J

Hum Biol. 21(6):762-8.Apr 14. [Epub ahead of print], Pub MedID: 19367574

2007 Erzurum, S.C., Ghosh, S., Janocha, A.J., Xu, W., Bauer, S., Bryan, N.S., Tejero, J., Hemann, C., Hille,

R., Stuehr, D.J., Feelisch, M., Beall, C.M Higher blood flow and circulating NO products offset high-altitude

hypoxia among Tibetans. Proceedings of the National Academy of Sciences 104, 17593, Pub MedID: 17971439

2007 Beall, C.M. Detecting natural selection in high-altitude human populations.

Respir Physiol Neurobiol. 158: 161-171. [2007 Jun 8; Epub ahead of print], Pub MedID: 17644049

2007 Beall, C.M. Two routes to functional adaptation: Tibetan and Andean high-altitude natives. Proc Natl

Acad Sci U S A. 2007 May 15;104 Suppl 1:8655-60. Epub 2007 May 9, Pub MedID: 17494744

2006 Brown, D. E., Beall, C. M., Strohl, K. P., and Mills, P. S. Exhaled Nitric Oxide Decreases upon Acute

Exposure to High-Altitude Hypoxia. American Journal of Human Biology 18(2): 196-202, Pub MedID:

16493632

2006 Beall, C. M.. Andean, Tibet, and Ethiopian Patterns of Adaptation to High-Altitude Hypoxia. Journal of

Integrative and Comparative Biology [Advance Access, January 6, 2006] Integrative and Comparative Biology

2006 46(1):18-24; doi:10.1093/icb/icj004

2005 Geoff Childs, M.C. Goldstein, Ben Jiao, C.M. Beall. Tibetan Fertility Transitions in China and south

Asia. Population and Development Review (31):2: 337-351.

2005 Hoit BD, Dalton ND, Erzurum SC, Laskowski D, Strohl KP, Beall CM. Jul 14; Nitric Oxide and Cardio-

Pulmonary Hemodynamics in Tibetan Highlanders. J. Appl Physiol. 99:1796-1801. First published July 14,

2005; doi: 10.1152/japplphysiol.00205.2005, Pub MedID: 16024527

2004 Beall, C. M., Song, K., Elston, R. C., Goldstein, M. C. Higher offspring survival among Tibetan women

with high oxygen saturation genotypes residing at 4000m. Proc. Nat. Acad. Sci. 101(39):14300-4 , Pub MedID:

15353580

2003 Goldstein, M. C., Jiao, B., Beall, C.M., and Tsering, P. 2003 Development and Change in rural Tibet.

Problems and Adaptations. Asian Survey 43:5: 758-779.

2002 Beall CM, Decker MJ, Brittenham GM, Kushner I, Gebremedhin A and Strohl KP. An Ethiopian pattern

of human adaptation to high-altitude hypoxia. Proc Natl Acad Sci U S A 99: 17215-17218, 2002, Pub MedID:

12471159

2002 Goldstein, Melvyn C., Jiao Ben (Benjor), Beall, C. M., and Tsering P. Fertility and Family Planning in

rural Tibet. The China Journal 47: 19-39.

2001 Beall, C., Laskowski, D., Strohl, K. P., Soria R., Villena, M., Vargas, E., Alarcon, A. M., Gonzales, C.

and Erzurum, S. C. Pulmonary nitric oxide in mountain dwellers. Nature 414: 411-412, Pub MedID: 11719794

2001 Beall, C. Adaptations to Altitude: A Current Assessment. Annual Review of Anthropology 16: 423-

455.

2000 Beall, C. Tibetan and Andean patterns of adaptation to high-altitude hypoxia. Human Biology 72(1):

201- 228, Pub MedID: 10721618

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2000 Beall, C. Oxygen saturation increases during childhood and decreases during adulthood among high

altitude native Tibetans residing at 3800-4200m High Altitude Medicine and Biology 1(1): 25-32, Pub MedID:

11258584

1999 Beall, C. M., Almasy, L. A., Blangero, J., Williams-Blangero, S., Brittenham, G. M., Strohl, K. P.,

Decker, M. J., argas, E., Villena, M., Soria, R., Alarcon, A. M., and Gonzales, C. Percent of oxygen saturation of

arterial hemoglobin among Bolivian Aymara at 3900-400m. American Journal of Physical Anthropology.

108(1): 41-52, Pub MedID: 9915300

1998 Beall, C. M., Brittenham, G. M., Strohl, K. P., Blangero, J. Williams-Blangero, S., Goldstein, M. C.,

Decker, M. J., Vargas, E., Villena, M., Soria, R., Alarcon, A. M., Gonzales, C. Hemoglobin concentration of

high-altitude Tibetans and Bolivian Aymara. American Journal of Physical Anthropology. 106: 385-400, Pub

MedID: 9696153

1997 Beall, C. M., K. P. Strohl, J. Blangero, S. Williams-Blangero, L. A. Almasy, M. J. Decker, C. M.

Worthman, M. C. Goldstein, E. Vargas, M. Villena, R. Soria, A. M. Alarcon, C. Gonzales. Ventilation and

hypoxic ventilatory response of Tibetan and Aymara high altitude natives. American Journal of Physical

Anthropology 104: 427-447, Pub MedID: 9453694

1997 Beall, C. M., K. P. Strohl, J. Blangero, S. Williams-Blangero, G. M. Brittenham, and M. C. Goldstein

Quantitative genetic analysis of arterial oxygen saturation in Tibetan Highlanders.

Human Biology 69(5): 597-604, Pub MedID: 9299881

1997 Beall, C. M., A. Gebremedhin, G. M. Brittenham, and M. Shamebo. Blood pressure variation among

Ethiopians on the Simien Plateau. Annals of Human Biology 24(4): 333-342, Pub MedID: 9239439

1996 Beall, C.M., J. Henry, C. Worthman, M.C. Goldstein. Basal Metabolic Rate and Dietary Seasonality

Among Tibetan Nomads. American Journal of Human Biology 8: 361-370.

1994 Beall, C.M., J. Blangero, S. Williams-Blangero and M.C. Goldstein. A major gene for %O2 saturation of

arterial hemoglobin in Tibetan highlanders. American Journal of Physical Anthropology 95(3): 271-276, Pub

MedID: 7856765

1993 Williams-Blangero, S., J. Blangero, and C. M. Beall. Genetic Analysis of Physical Chest Dimensions in a

High Altitude Tibetan Population from Upper Chumik, Nepal. American Journal of Human Biology. 5(6): 719-

724.

1993 Beall, C.M. and M.C. Goldstein. Dietary seasonality among Tibetan Nomads. Research and

Exploration 9(4): 477-479.

1992 Beall, C.M. and M.C. Goldstein. High prevalence of excess fat and central fat patterning among

Mongolian pastoral nomads. American Journal of Human Biology 4(6): 747-756.

1992 Beall, C.M., K.P. Strohl, B. Gothe, G.M. Brittenham, M. Barragan, E. Vargas. Respiratory

and hematological adaptations of young and older Aymara men native to 3600m. American Journal of

Human Biology 4(1): 17-26.

1992 Beall, C.M., C.M. Worthman, J. Stallings, K.P. Strohl, G.M. Brittenham and M. Barragan Salivary

testosterone concentration of Aymara men native to 3600m. Annals of Human Biology 19(1): 67-78, Pub

MedID: 1734824

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1992 Beall, C.M., C. Ikels, and M.C. Goldstein. Introducing the “Aging Trends” series. Journal of Cross-

Cultural Gerontology 7(1): 1-2

1991 Goldstein, M.C. and C.M. Beall. China's Birth Control Policy in Tibet. Asian Survey 31(3): 285-303.

1991 Goldstein, M.C. and C.M. Beall. Nomadic Pastoralism on the Western Tibetan Plateau. Nomadic

Peoples 28: 105-122.

1991 Cincotta, R.P., P.M.Van Soest, J.B. Robertson, C.M. Beall and M.C. Goldstein. Foraging ecology of

livestock on the Tibetan changtang: a comparison of three adjacent grazing areas. Arctic and Alpine Research

23(2): 149-161.

1990 Beall, C.M., G.M. Brittenham, F. Macuaga, M. Barragan.. Variation in hemoglobin

concentration among samples of high altitude natives in the Andes and the Himalayas. American Journal

of Human Biology. 2(6): 639-662.

1990 Goldstein, M.C., C.M. Beall, and R.P. Cincotta. Indigenous nomadic pastoralism and

ecological conservation on Tibet's "Northern Plateau" National Geographic Research 6(2): 139-156.

1989 Goldstein, M.C. and C.M. Beall. The Impact of China's Cultural and Economic Reform Policy on

Nomadic Pastoralism in Western Tibet. Asian Survey. 29(6): 619-641.

1987 Beall, C.M., M.C. Goldstein and the Tibetan Academy of Social Sciences. Hemoglobin Concentration of

Tibetan Nomads Permanently Resident at 4,850-5,450m. American Journal of Physical Anthropology. 73(4):

433-438, Pub MedID: 3661681

1986 Goldstein, M.C. and C.M. Beall. Family change, Caste and the Elderly in a Rural Locale in Nepal.

Journal of Cross-Cultural Gerontology. 1(3):305-316.

1986 Beall, C.M., J.K. Eckert and M.C. Goldstein. Editorial: On Cross-Cultural Gerontology. Journal of

Cross-Cultural Gerontology 1(1):1-3.

1986 Beall, C.M. and M.C. Goldstein. Age differences in sensory and cognitive function in an elderly non-

Western population. Journal of Gerontology 41(3):387-389, Pub MedID: 3486204

1985 Beall, C.M., M.C. Goldstein, E.S. Feldman. Social Structure and Intracohort Variation in Physical

Fitness Among Elderly Males in a Traditional Third World Society. Journal of American Geriatrics Society

33(6):406-412. (Abstracted in 1986, Geriatrics Digest, 3:14-15), Pub MedID: 3998350

1985 Beall, C.M., M.C. Goldstein, E.S. Feldman. The Physical Fitness of Elderly Nepalese Farmers Residing

in Rugged Mountain and Flat Terrain. Journal of Gerontology 40(5):529-535, Pub MedID: 4031400

1984 Goldstein, M.C., C.M. Beall and P. Tsarong. On Studying Fertility at High Altitude: A rejoinder to

Hoff's reply. American Anthropologist 86(2):419-422

1984 Beall, C.M. and A.B. Reichsman. Hemoglobin Levels in a Himalayan High Altitude Population.

American Journal of Physical Anthropology 63(3):301-306, Pub MedID: 6731601

1984 Goldstein, M.C., C.M. Beall and P. Tsarong. Response to Abelson's Comment on Goldstein et al.

American Anthropologist 86(3):703-705.

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1984 Goldstein, M.C., C.M. Beall and P. Tsarong. Response to Basu and Gupta's comments on High Altitude

Hypoxia, Culture and Human Fecundity/Fertility. American Anthropologist 86 (4):996-997.

1983 Goldstein, M.C., P. Tsarong, and C.M. Beall. High Altitude Hypoxia, Culture and Human

Fecundity/Fertility: a Comparative study. American Anthropologist 85(1):28-49

1983 Beall, C.M., Age at Menopause and Menarche in a High Altitude Himalayan Population. Annals of

Human Biology 10(4):365-370.

1983 Beall, C.M., K.P. Strohl and G. Brittenham. A Reappraisal of Andean High Altitude Erythrocytosis from

a Himalayan Perspective. Seminars in Respiratory Medicine 5(2):195-201.

1982 Beall, C.M. Introduction to special issue of Social Science and Medicine devoted to Biocultural Studies

of Aging. Social Science and Medicine 16(2):131-134.

1982 Beall, C.M. and M.C. Goldstein. Work, aging and dependency in a Sherpa population in Nepal. Social

Science and Medicine 16(2):141-148, Pub MedID: 7100965

1982 Beall, C.M. and M.C. Goldstein. Biological function, activity and dependency among elderly Sherpa in

the Nepal Himalayas. Social Science and Medicine 16(2):135-140, Pub MedID: 7100964

1982 Goldstein, M.C. and C.M. Beall. Indirect Modernization and the Status of the Elderly in Rural Third

World Countries. Journal of Gerontology 37(6):743-748, Pub MedID: 7130649

1982 Goldstein, M.C. and C.M. Beall. Tibetan Fraternal Polyandry and Sociobiology: A rejoinder to

Abernethy and Fernandez. American Anthropologist 84(4):898-901.

1981 Beall, C.M. Growth in a Population of Tibetan Origin at High Altitude. Annals of Human Biology

8(1):31-38, Pub MedID: 7224586

1981 Beall, C.M. and M.C. Goldstein. Tibetan Fraternal Polyandry: A Test of Sociobiological Theory.

American Anthropologist 83(1):5-12.

1981 Goldstein, M.C. and C.M. Beall. Modernization and Aging: views from the rural, pre-industrial

hinterland in Nepal. Human Organization 40(1):48-55.

1981 Beall, C.M. Optimal Birthweights in Peruvian Populations at High and Low Altitudes. American

Journal of Physical Anthropology 56(3):209-216, Pub MedID: 7325220

1981 Beall, C.M. A Comparison of Chest Morphology in High Altitude Asian and Andean Populations.

Human Biology 54(1):145-163.

1977 Beall, C.M., P.T. Baker, T.S. Baker and J.D. Haas. The Effects of High Altitude on Adolescent Growth

in Southern Peruvian Amerindians. Human Biology 49(2):109-124, Pub MedID: 889591

OTHER PUBLICATIONS: INVITED CONTRIBUTIONS:

2012 (expected) Beall, C. M. Human Evolution at High Altitude, chapter 19 in Erik R. Swenson and

Peter Bartsch, editors, Hypoxia: Human adaptation to high altitude, second edition, New York,

N.Y.: Marcel Dekker, Inc.

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2012 (expected) Beall, C. M., Jablonski, N and A. T. Steegmann, Jr. Human Adaptation to Climate:

Temperature, Ultraviolet Radiation,and Altitude. In Human Biology. An Evolutionary and Biocultural

Perspective, 2nd

edition, S. Stinson, B. Bogin, R. Huss-Ashmore, and E. O’Rourke (eds.), Wiley-Liss,

Inc., New York

2011 Beall CM Guest Editorial. Genetic changes in Tibet. In press High altitude medicine & biology. 12 (2):

101-102. DOI: 10.1089/ham.2011.1007

2010 (expected) Beall, C. M., Jablonski, N and A. T. Steegmann, Jr. Human Adaptation to Climate:

Temperature, Ultraviolet Radiation,and Altitude. In Human Biology. An Evolutionary and Biocultural

Perspective, 2nd

edition, S. Stinson, B. Bogin, R. Huss-Ashmore, and E. O’Rourke (eds.), Wiley-Liss, Inc., New

York

2008 Beall, C. M. Why Are We Vulnerable to Acute Mountain Sickness? Pp. 259-276 in Trevathan, W.,

McKenna, J. M. and E. O. Smith (eds.), Evolutionary Medicine and Health. New Perspectives, Oxford University

Press, New York.

2005 Goldstein, M. C., Jiao, B., Beall, C. M. and Tsering, P. Development and Change in Rural Tibet:

Problems and Adaptations. Pp 193-213 in Sautman, B. and J. T. Dreyer (eds.), Contemporary Tibet. Politics,

Developoment, and Society in a Disputed Region. M. E. Sharpe Pubs., Armonk, NY.

2003 Beall, C.M. High-altitude adaptations. The Lancet 362(1): S14-S15, Pub MedID: 14698112

2002 Beall, C. M. Biodiversity of Human Populations in Mountain Environments, pp. 199-210 in Mountain

Biodiversity. A Global Assessment, Ch. Koerner and E. M. Spehn (eds.), The Parthenon Publishing Group,

Boca Raton, FL.

2002 Goldstein, M. C. and Beall, C.M. Changing pattern of Tibetan nomadic pastoralism., pp. 131-150 in

Human Population Biology of Pastoral Populations, W. R. Leonard and M. H. Crawford (editors), Cambridge

University Press, Cambridge.

2001 Little, M. A., Badgley, C., Beall, C., Balick, M. Munstermann, L. E., Weiss, K. M. Bert, Th. M., and

Chernoff, B. A Framework for a program in the human dimensions of biodiversity. Biology International 42: 3-

15.

2001 Ikels, C. and Beall, C. M.Age, Aging, and Anthropology, In Handbook of Aging and the Social Sciences,

R. H. Binstock and L. K. George (eds.).,Academic Press, San Diego, pp. 125-13

2000 Beall , C. M. Tibetan and Andean Contrasts in Adaptation to High-Altitude Hypoxia. In Oxygen Sensing.

Molecule to Man. S. Lahiri, N. R. Prabhakar, R. E. Forster (eds.), Advances in Experimental Medicine and

Biology, volume 475, pp. 63-74, Pub MedID: 10849649

1999 Beall, C. M. and A. T. Steegmann, Jr. Human Adaptation to Climate: Temperature, Ultraviolet

Radiation,and Altitude. In Human Biology. An Evolutionary and Biocultural Perspective. S. Stinson, B. Bogin,

R. Huss-Ashmore, and E. O’Rourke (eds.), Wiley-Liss, Inc., New York, pp. 163-224.

1997 Beall, C. M. Are Sherpas Different? In: Everest. Mountain Without Mercy. Broughton Coburn,

National Geographic Society, Washington, D. C., p. 126

1997 Strohl, K. P. and C. M. Beall Ventilatory Response to Experimental Hypoxia in Adult Male and Female

Natives of the Tibetan and Andean Plateaus. In: Hypoxia: Women at Altitude. C. S. Houston and G. Coates

(eds.), Queen City Printers, Burlington, VT., pp. 162-165.

11

1997 Goldstein, M. C. and C. M. Beall. Growing Old in Tibet—Tradition, Family and Change. In: Aging:

Asian Concepts and Experiences Past and Present. S. Formanek and S. Linhart (eds.) Verlag Der

Osterreichischen Akademie Wissenschaften, Wien, pp. 155-176.

1994 Beall, C.M. Aging and Adaptation to the Environment. In: Biological Anthropology and

Aging: Interdisciplinary, cross-cultural and comparative approaches. R.M. Garruto and D.J. Crews

(eds). Oxford University Press, pp. 339-372.

1994 Beall, C.M. High altitude human biology. In: N. Allen (ed). Illustrated Library of the Earth:

Mountains, Weldon Owen Pubs, Sydney, pp. 62-63.

1993 C.M. Beall and M.C. Goldstein. The Changing world of Mongolia's nomads. National

Geographic. 183(5):127-138.

1993 Beall, C.M. Genetic Bases of oxygen transport at high altitude, traditional and new approaches. In

Hipoxia: investigaciones basicas y clinicas. Homenaje a Carlos Monge Cassinell. Leon-Velarde, F. and Arregui,

A. (eds.). IFEA, Lima, pp. 111-124.

1992 Goldstein, M.C. and C.M. Beall. Nomads on the roof of the world, pp.420-423. In M.C. Howard and J.

Dunaif-Hattis, Anthropology Understanding Human Adaptation, Harper Collins Pub.

1990 Beall, C.M. and M.C. Goldstein. Hemoglobin Concentration, % oxygen saturation and arterial oxygen

content of Tibetan nomads at 4,850 m- 5450m. pp 59-65 in:Hypoxia. The Adaptations, J.R. Sutton, G.Coates

and J.E. Remmers (eds). B.C. Decker Inc., Toronto.

1989 Goldstein, M.C. and C.M. Beall. The secluded world of Tibet's nomads. National Geographic 175(6):

752-781.

1989 Beall, C.M. and C.A. Weitz. The Human Population Biology of Aging. pp.189-202 in: Human

Population Biology: a Transdisciplinary Science, M.A. Little and J.D. Haas (editors), Oxford University Press.

1989 Weitz, C.A, L.P. Greksa, R.B. Thomas, and C.M. Beall. An Anthropological Perspective on the Study

of Work Capacity. pp 132-151 in: Human Population Biology: a Transdisciplinary Science, M.A. Little and J.D.

Haas (editors), Oxford University Press.

1989 Greksa, L.P. and C.M. Beall. Development of Chest size and Lung Function at High Altitude. pp. 222-

238 in: Human Population Biology: a Transdisciplinary Science, M.A. Little and J.D. Haas (editors), Oxford

University Press.

1987 Beall, C.M. Studies of Longevity. pp. 73-93 in Modern Pioneers: an Interdisciplinary View of the Aged,

P. Silverman (editor), Indiana University Press.

1988 Beall, C.M. and M.C. Goldstein. Socio-cultural influences on the working capacity of elderly Nepali

men. pp. 215-226 in Capacity for work in the Tropics. K.J. Collins and D.F. Roberts (editors), Cambridge

University Press.

1987 Beall, C.M. Nutrition and Variation in Biological Aging. pp.197-221 in Nutrition in Anthropology, F.E.

Johnston (editor), Alan R. Liss, Inc. N.Y.

1986 Beall, C.M. and J.K. Eckert. Measuring Functional Status Cross-Culturally. pp. 21-56 in New Methods

for Old Age Research, second edition, C. Fry and J. Keith (editors), Bergin and Garvey, Pubs., Inc., MA

12

1984 Beall, C.M. Theoretical Dimensions of a Focus on Age in Physical Anthropology. pp. 82-95 in Age and

Anthropological Theory. D. Kertzer and J. Keith (editors). Cornell University Press.

1984 Beall, C.M. Aging and Growth at High Altitudes in the Himalayas. pp. 365-385 in The People of South

Asia J. Lukacs (editor), Plenum Press.

1982 Baker, P.T. and C.M. Beall. The Biology and Health of Andean Migrants: A Case Study in South

Coastal Peru. Mountain Research and Development 2(1):81-95, Special issue. Human Population and Biosphere

Interactions in the Central Andes, P.T. Baker (editor).

1982 Escobar, G. and C.M. Beall. Contemporary patterns of Migration in the Central Andes. Mountain

Research and Development 2(1):63-80, Special issue: Human Population and Biosphere Interactions in the

Central Andes, P.T. Baker (editor).

1982 Beall, C.M. A Historical and Analytical Review of the Study of Human Growth and Development in

Extreme Environment in Extreme by American Physical Anthropologists. Pp. 447-465 in A History of American

Physical Anthropology 1930-1980. F. Spencer (editor) Academic Press, pp. 447-465.

1981 Beall, C.M. Some Aspects of the Study of Physical Growth at High Altitude in Asia. pp. 49-54 in

Environmental and human Population Problems at High Altitude. Editions du Centre National de Recherche

Scientifique, Paris.

1980 Eckert, J.K. and C.M. Beall. Approaches to Measuring Functional Capacity Cross-Culturally. Pp. 27-56

in New Methods for Old Age Research. C. Fry and J. Keith (editors). Loyola University Press.

OTHER PUBLICATIONS: UNREFEREED JOURNALS, NEWSLETTERS, BOOK REVIEWS,

COMPILATIONS:

2006 Beall, C.M. Book Review of An Ecology of High Altitude Infancy. By Andrea S. Wiley, New York:

Cambridge University Press. 2004. 245 pp. American Journal of Physical Anthropology 130: 276-277.

2002 Beall, C. M. Book Review of High Altitude: An Exploration of Human Adaptation. Edited by

Thomas F. Hornbein and Robert B. Schoene. xxii + 982 pp. New York: Marcel Dekker, Inc. 2001. $235.00

(cloth). American Journal of Human Biology 14: 786-787.

2001 Little, M., Badgley, C.. Beall, C., Balick, M., Munstermann, L. E., Weiss, K. M., Bert, Th. M, and

Chernoff, B. A Framework for a Program in the Human Dimensions of Biodiversity. Biology International 42:

3-15.

2001 Beall, C.M. Turkana Herders of the Dry Savana: Ecology and Biobehavioural Response of Nomads

to an Uncertain Environment. M. A. Little and P. W. Leslie (eds.), Oxford University Press. American Journal

of Human Biology 13(1): 81.

1996 Beall, C. M. (compiler) Human Biology Association Guide to Graduate Programs and Graduate

Training in Human Biology. American Journal of Human Biology 8: 1-20.

1993 Beall, C.M. Book review of Flocks and Food by M.J. Casimir. Ecology of Food and Nutrition 30: 323-

4.

1993 Beall, C. M. Book review of Growth, Maturation and Physical Activity by R. M. Malina and C.

Bouchard. American Journal of Physical Anthropology. 91(1): 134-135.

13

1992 Beall, C.M. Book review of Longevity, Senescence and the Genome by C.E. Finch.,

Annals of Human Biology. 19(6): 618-620.

1990 Beall, C.M. Book review of Hypoxia: The Tolerable Limits by J.R. Sutton, C.S. Houston and G. Coates

(eds), American Journal of Human Biology 2(1): 91-92.

1989 Beall, C.M. Book Review of Patterns of Human Growth by B. Bogin. Annals of Human Biology. 16(5):

476-477.

1986 Goldstein, M.C. and C.M. Beall. Studying Nomads on the Tibetan Plateau. China Exchange News,

14(4):2-7.

1986 Beall, C.M. Book Review of Human Adaptation edited by A. Basu and K.C. Malhotra. American

Journal of Physical Anthropology 69(3):416-418.

1984 Beall, C.M. Book Review of Origins of the Study of Human Growth by Edith Boyd, edited by B.S.

Savara and J.F. Schilke. American Journal of Physical Anthropology 64(1):93-94.

1983 Beall, C.M. Book Review of Social and Biological Predictors of Nutritional Status, Physical Growth and

Neurological Development edited by L.S. Green and F.E. Johnston. Human Biology 54(4):843-844.

1982 Beall, C.M. Ages at menopause and menarche in a High Altitude Himalayan Population. Contributions

to Nepalese Studies 9(1-2):49-54.

1982 Beall, C.M. Himalayan-Andean differences in hemoglobin levels among high altitude natives:

implications of a study in upper Chumik, Mustang District, Nepal. Journal of the Institute of Medicine,

Kathmandu, Nepal. 4(1):31-39.

1982 Goldstein, M.C. and C.M. Beall. Brief Note on Demographic Aspects of Aging in the Lesser Developed

Countries. Newsletter of the Association for Anthropology and Gerontology 3(3):2-3.

1981 Beall, C.M. A Review of Morphological aspects of Growth at High Altitudes in Asia. Journal of the

Institute of Medicine, Kathmandu, Nepal. 3(1):71-84.

1980 Goldstein, M.C. and C.M. Beall. Growing old in Helambu: Aging, Migration and Family Structures

among Sherpas. Contributions to Nepalese Studies 8(1):41-56.

PUBLISHED ABSTRACTS:

2009 C. M. Beall, B. Hoit, N. Dalton, A. Gebremedhin, K. P. Strohl, S. C. Erzurum. Pulmonary artery

hemocynamics of high and low altitude native Ethiopian Amhara. American Journal of Physical Anthropology,

supplement 48: 87.

2008 C.M.Beall, S. Ghosh, A.J. Janocha, W. Xu, S. Bauer, N.S. Bryan, J. Tejero, C. Hemann, R. Hille, D. J.

Stuehr, M. Feelisch, S. C. Erzurum. Higher blood flow and circulating nitric oxide products among high-altitude

Tibetans. American Journal of Physical Anthropology, supplement 46: 34.

2007 C. M. Beall Origins: human adaptation to high-altitude hypoxia. American Journal of Physical

Anthropology, supplement 44: 70.

2002 C. Beall Overview of the International Union of Biological Sciences (IUBS), American

14

Journal of Physical Anthropology, supplement 34: 40.

2002 D. Brown, P. Mills, K. Strohl, C. Beall, Nitric oxide exhalation is elevated upon acute exposure to high

altitude hypoxia and is related to individual distress, American Journal of Physical Anthropology, supplement 34:

49.

2001 Beall, C. M., K. P. Strohl, S. C. Erzurum, D. Laskowski. Exhaled nitric oxide concentration of Andean

and Tibetan high-altitude natives. American Journal of Physical Anthropology. Supplement 32: 37.

2001 Olatunde, S. B., Kemp, B. F., Beall, C. M., and Merriwether, D. A. Mitochondrial Diversity of Amharic

Speakers in Ethiopia Compared with Twenty African Populations. (Abstract). American Journal of Human

Biology 13(1): 136

2001 Tanner, S. M., McDonough, J. E., Beall, C. M., and Merriwether, D. A. Mitochondrial DNA Sequence

Variation in the Bolivian Aymara. (Abstract) American Journal of Human Biology 13(1): 143

Beall, C. M. M. J. Decker, K. P. Strohl, I. Kushner, G. M. Brittenham, L. A. Almasy, J. Blangero,

A. Gebremedhin. Hemoglobin concentration of Ethiopians at 3530m. American Journal of Physical

Anthropology Supplement 29: 104.

2000 D. A. Merriwether, B. Kemp, and C. M. Beall Mitochondrial DNA Sequence Variation in a High-

Altitude Population in Tibet. American Journal of Human Biology 12: 2, 275.

2000 B. Kemp, D. A. Merriwether, and C. M. Beall. Mitochondrial DAN Sequence Variation in the

Simien Plateau in Ethiopia. American Journal of Human Biology 12: 2, 283.

1999 Beall, C. M., C. M. Worthman, J. Stallings. Oxygen saturation and the response to sub-maximal exercise

among Tibetans at 3900m. American Journal of Physical Anthropology, Supplement 28: 88

1997 Beall, C. M., G. M. Brittenham., K. P. Strohl, M. J. Decker, M. C. Goldstein, J. Blangero, S. Williams-

Blangero, L. Almasy. Ventilation and hypoxic ventilatory response of Tibetan and Aymara high altitude natives.

American Journal of Physical Anthropology, Supplement 24: 73.

Worthman, C. M., C. M. Beall, J. F. Stallings. Population differences in DHEAS across the lifespan:

implications for aging. American Journal of Human Biology 9(1): 149.

1997 Worthman, C. M., C. M. Beall, J. F. Stallings. Population variation in reproduction function of men.

American Journal of Physical Anthropology, Supplement 24: 246.

1996 Beall, C. M. The future of Human Population Biology. American Journal of Physical Anthropology,

Supplement 22: 66.

1995 J. Blangero, S. Williams-Blangero, M.J. Decker, K.P. Strohl, G. Brittlenham and C.M. Beall; Genetic

analysis of hypoxic ventilatory response in a high-altitude Tibetan population. American Journals of Physical

Anthropology, Supplement 19.

1995 Strohl, K. P., C. M. Beall, M. J. Decker, G. Brittenham, J. Blangero, S. Williams-Blangero. Quantitative

genetic analysis of hypoxic ventilatory response at 4000m. Respiratory and Critical Care Medicine v. 151 (4):

A634.

1994 J. Blangero, S. Williams-Blangero, and C.M. Beall. Genetic analysis of hemoglobin levels in two high-

altitude Tibetan populations. American Journal of Physical Anthropology. Supplement 18:56.

15

1993 Beall, C.M. The ecological approach in physical anthropology since the 1969 publication of Lasker’s

article “Human Biological Adaptability”. American Journal of Physical Anthropology, Supplement 16:53.

1993 M.C. Goldstein, J. Blangero, S. Williams-Blangero, C.M. Beall. Statistical genetic evidence that Tibetan

highlanders have major gene for percent oxygen saturation of arterial hemoglobin. American Journal of Physical

Anthropology, Supplement 16:96.

1993 S. Williams-Blangero, J. Blangero, C.M. Beall. Genetic analysis of chest dimensions in a high altitude

Tibetan population from Upper Chumik, Nepal. American Journal of Physical Anthropology, Supplement

16:209.

1992 Beall, C.M., and M.C. Goldstein. Excess fat, centripetally distributed, of Mongolian Pastoral Nomads.

American Journal of Physical Anthropology. Supplement 14, to vol. 87: 46.

1990 Beall, C.M., G.M. Brittenham, B.M. Gothe, M. Barragan, E. Vargas Aging and adaptation to high

altitude hypoxia. American Journal of Physical Anthropology 81 (2): 192 (Abstract)

1989 Beall, C.M., G.M. Brittenham, F. Macuaga, M. Barragan. Intra- and Interpopulation Variation in

Hemoglobin Concentration of Andean and Himalayan High Altitude Populations. American Journal of Physical

Anthropology 78(2):189-190. (Abstract)

1987 Beall, C.M., M.C. Goldstein and E.S. Feldman. Hemoglobin Concentration and Iron Status of Elderly

Nepali Males. American Journal of Physical Anthropology 72(2):176. (Abstract)

1986 Beall, C.M. Factors Associated with Menopausal Status in a High Altitude Tibetan Population.

American Journal of Physical Anthropology 69(2):174 (Abstract).

1985 Beall, C.M. and M.C. Goldstein. Caste and Terrain Influences on the Physical Fitness of Elderly

Nepalese. American Journal of Physical Anthropology 66(2):143 (Abstract).

1983 Beall C.M. Functional and Morphological Growth and Development of High Altitude Natives:

Comparison of Findings Among Himalayan and Andean Populations. American Journal of Physical

Anthropology 60(2):172 (Abstract).

1983 Beall, C.M. Tibetans Adapt to High Altitude Without High Hemoglobin Levels. In J.R. Sutton, C.S.

Houston, and N.L. Jones (eds.): Hypoxia. Exercise and Altitude. Alan R. Liss Inc. N.Y 450 (Abstract).

1981 Beall, C.M. An Historical Review of Studies Conducted by American Physical Anthropologists of

Human Growth and Development in Extreme Environments. American Journal of Physical Anthropology

54(2):199-200 (Abstract).

1980 Beall, C.M.and M.C. Goldstein. Biological Function, Activity and Dependency among Elderly Sherpas

in the Nepal Himalayas. American Journal of Physical Anthropology 52(2):203 (Abstract).

1979 Beall, C.M. and M.C. Goldstein. Multiple Brother Marriages and Parental Investment in the Limi

Population of Northwest Nepal. American Journal of Physical Anthropology 50(3):417 (Abstract).

1978 Beall, C.M. Allometric Growth Patterns of a Population Residing at High Altitude in Nepal. American

Journal of Physical Anthropology 48(3):380 (Abstract).

16

1977 Beall, C.M. Natural Selection for Optimum Birthrates in Peruvian Populations at High and Low

Altitudes. American Journal of Physical Anthropology 47(1):116 (Abstract).

1976 Beall, C.M. The Impact of High Altitude on Infant Mortality and Morbidity. American Journal of

Physical Anthropology 44(1):165 (Abstract).

1975 Beall, C.M. The Effects of Acute Illness on Early Growth at High Altitude. American Journal of

Physical Anthropology 42(2):290 (Abstract).

RESEARCH GRANT AWARDS: (Principal Investigator unless noted otherwise.)

2009 – 2011 National Science Foundation Grant No. BCS-0924726 “Nitric oxide and the microcirculation in

the Tibet Autonomous Region”

2005-2007 National Science Foundation Grant No. BCS-0452326 “Ethiopian adaptation to high-altitude

hypoxia”

2002-2005 National Science Foundation Grant No. BCS-021547 “Nitric oxide and oxygen delivery among

Tibetans”

2000 National Science Foundation Grant No. SES-00074567 “POWRE: Pulmonary Nitric Oxide and

adaptation to High Altitude Hypoxia”.

2000 National Science Foundation Grant No. BCS-00084355 “High Risk Exploratory Research:

Exhaled Nitric Oxide of Andean and Tibetan High-Altitude Natives”, with Kingman P. Strohl.

1998 Wenner-Gren Foundation for Anthropological Research Grant No. 6322 “Mitochondrial DNA analyses of

high-altitude Tibetan, Andean, and Simien Plateau samples”, with D. Andrew Merriwether (University of

Michigan)

1997-2000 National Science Foundation Grant No. SBR-9706980, “Fitness and function of oxygen

saturation genotypes in Tibet”, co-P. I. with Carol M. Worthman (Emory University) and John Blangero

(Southwest Foundation for Biomedical Research)

1997-2000 The Henry Luce Foundation ”Socioeconomic effects of China’s Socioeconomic reform policy in

the Tibet Autonomous Region, co-P.I. Melvyn C. Goldstein (Case Western Reserve University)

1996-97 Research Program Scholar, Committee on Scholarly Communication with China, "Ten Year

Longitudinal field study of the pastoral nomads of Phala, TAR"

1996-97 National Geographic Society Grant No. 5603-95, “Ten-year follow-up study of Tibetan Pastoral

nomads in Tibet”, co P. I. Melvyn C. Goldstein

1995 National Geographic Society Grant No. 5520-95 "Genetics of Ethiopian highlanders' hematologic

adaptation to high altitude hypoxia"

1993 National Science Foundation Grant No. DBS 9221724 “Genetics of oxygen transport at high altitude”

1993 National Geographic Society Grant No. 4976-93 “Follow up field study of Tibetan Pastoral Nomadism”

(Co-Principal Investigator Melvyn C. Goldstein)

1991 International Research and Exchange Board, "Field study of Mongolian Pastoral Nomadism".

17

1990 National Geographic Society, "High altitude hypoxia, testosterone levels and hemoglobin concentration

of pastoral nomads in the Tibet Autonomous Region of China". (Co-Principal Investigator: Melvyn C.

Goldstein).

1990 International Research and Exchange Board, "Field study of Mongolian Pastoral Nomadism".

1987 National Science Foundation Grant No. BNS-8704213, "Hypoxemia and Pastoral Nomads in the Tibetan

Autonomous Region of China." (Co-Principal Investigator: Melvyn C. Goldstein).

1986-87 Western Reserve College, Research Incentives Program grant award, "Hemoglobin

Concentration and Iron Status in Pucarani, Bolivia."

1986-87 American Federation for Aging Research (AFAR) grant award, "Aging and Adaptation to High

Altitude Hypoxia in Bolivia".

1986-87 National Geographic Society Grant No. 3322-86, "Pastoral Nomads in Western Tibet" (Co-

Principal Investigator: Melvyn C. Goldstein).

1986-87 National Academy of Sciences, National Program for Advanced Study and Research in China,

"Tibetan Pastoral Nomadism: Cultural and Ecological Perspectives".

1986 Western Reserve College, Research Incentives Program grant award, "Hypoxemia of Bolivian High

Altitude Natives".

1983 American Federation for Aging Research (AFAR) grant award, "Lifelong High Physical Activity Level

and the Aging Process".

1983 National Geographic Society Grant No. 2641-83, "Lifelong High Physical Activity Levels and Biological

Aging in Nepal".

1983 National Science Foundation Grant No. BNS82-19188, "The Effect of Lifelong High Levels of Physical

Activity on the Aging Process" (Co-Principal Investigator: M.C. Goldstein).

1982 Case Western Reserve University, Office of Research Administration, Research Initiation Grant, "High

Altitude Adaptation among Muslims of the Pakistani Karakorum Mountain Range".

1981 National Science Foundation Grant No. BNS-8014317, "Senescence and Growth at High Altitude".

1980 Case Western Reserve University, Office of Research Administration, Research Initiation Grant,

"Physical Activity and Aging in Rugged Mountain Environments".

1980 Case Western Reserve University, Biomedical Research Award, "Physical Activity and Aging in Rugged

Mountainous Environments: Differential Aging in Hindu Populations in Helambu, Nepal".

1979 Case Western Reserve University, Biomedical Research Award, "Biocultural Aspects of Aging among

Sherpas". (Co-Principal Investigators: M.C. Goldstein, G.M. Brittenham).

1976 Case Western Reserve University, Biomedical Research Award, "Growth of Tamang Children at High

Altitude in Nepal".

18

CONFERENCE GRANT AWARDS:

2008 National Center for Evolutionary Synthesis Catalysis Meeting Award, co-PI, “Human Evolution and

adaptation to high-altitude hypoxia”

1993 Wenner-Gren Foundation for Anthropological Research. Grant No. 48-111 to aid conference on

“Individual, Intra- and Inter-population variation in DNA” sponsored by Human Biology Council, in

collaboration with Michael H. Crawford, University of Kansas.

RESEARCH OUTSIDE THE UNITED STATES:

2011: August, follow-up demographic study of Tibetan nomads and villagers

2010: May through July, nitric oxide and the microcirculation among Sherpas in Nepal

2009: May – July, follow up study of Tibetan nomads and villagers

2007 & 2008: June-August, study of modernization and elderly Tibetan villagers

2007: December, Bale Mountains, Ethiopia: oxygen delivery of high-altitude Oromo

2005: June – August, Tibet Autonomous Region: follow-up study of Tibetan nomads and villagers.

2005: March – May, Semien Mountains, Ethiopia: oxygen delivery of high-altitude Amharas

2002 June-August, Tibet Autonomous Region: nitric oxide and oxygen delivery of high-altitude Tibetans

2000 June, Bolivia; July-August, Tibet Autonomous Region: exhaled nitric oxide of high-altitude natives

1997-98 December-March, Pembo, Tibet Autonomous Region: biological fitness and function of Tibetan

villagers

1997 May-August, Phala, Tibet Autonomous Region: 10 year follow-up study of Tibetan pastoral nomads

1995 December, Amba Ras, North Gondar, Ethiopia: hematological adaptation to high altitude

1994 May-August, Ventilla, Provincia Murillo, Bolivia: genetics of oxygen transport.

1993 May-November, Lhasa Municipality, Tibet Autonomous Region, Peoples Republic of

China: genetics of oxygen transport.

1993 July-August,December, Ningor Xiang, Namring Zone, Tibet Autonomous Region, Peoples Republic of

China: seasonal variation in basal metabolic rate.

1991 May-June, Moct Som, Hovd Aimak, Mongolian Peoples' Republic: ecology of Mongolian pastoral

nomadism.

1990 September-November, Moct Som, Hovd Aimak, Mongolian Peoples Republic: ecology of Mongolian

pastoral nomadism.

1990 June-August, Phala Xiang, Namring Zone, Tibet Autonomous Region, Peoples' Republic of China: high

altitude hypoxia, testosterone and hemoglobin concentrations.

19

1987-88 June-June, Phala Xiang, Namring Zone, Tibet Autonomous Region, Peoples' Republic of China:

ecology and high altitude adaptation, growth, health and nutritional status of Tibetan nomads.

1987 April, Pucarani, Bolivia: hematological survey of adults in a rural high altitude zone.

1987 January-March, La Paz, Bolivia: aging, sleep and hematological adaptation to high altitude.

1986 May-August, Phala Shan, Namring Zone, Tibetan Autonomous Region, Peoples' Republic of China: high

altitude adaptation of Tibetan nomads.

1983 May-September, Lamjung District, Nepal: physical fitness of elderly Nepalis in rugged and flat terrains.

1982 July-August, Kathmandu, Nepal: epidemiology of menopause among Nepalis and Newaris.

1981 May-October, Upper Chumik, Nepal: high altitude Tibetan growth and senescence.

1980-81 December-January, Palchok, Nepal: differential aging in Nepalese and Sherpas.

1979 July-August, Helambu, Nepal: biological aging in Sherpas.

1977 May-August, Mugu, Nepal: high altitude Tibetan growth.

1973-74 May-May, Puno and Tacna, Peru: altitude effects on fetal growth, infant mortality.

1971 May-August, Tambo Valley, Peru: high altitude migrants, health, growth, and development.

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