The Major Senses

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1 The Major Senses There are 6 major senses vision hearing touch taste pain smell The list can be extended with balance, joint senses and others Vision has been studied most extensively

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The Major Senses. There are 6 major senses vision hearing touch taste pain smell The list can be extended with balance, joint senses and others Vision has been studied most extensively. Vision. Purpose of the visual system transform light energy into an electro-chemical neural response - PowerPoint PPT Presentation

Transcript of The Major Senses

Page 1: The Major Senses

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The Major Senses There are 6 major senses

vision hearing touch taste pain smell

The list can be extended with balance, joint senses and others

Vision has been studied most extensively

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VisionPurpose of the visual system

transform light energy into an electro-chemical neural response

represent characteristics of objects in our environment such as size, color, shape, and location

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Light - The Visual Stimulus

Gammarays

X-raysUltra-violetrays

Infraredrays Radar

Broadcastbands

ACcircuits

Visible light

PrismWhitelight

400 500 600 700

10 -5 10 -3 10 -1 10 1 10 3 10 5 10 7 10 9 10 11 10 13 10 15 10 17

Wavelength in nanometers (billionths of a meter)

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Light - The Visual StimulusLight can be described as both a particle

and a waveWavelength of a light is the distance of

one complete cycle of the waveVisible light has wavelengths from about

400nm to 700nm Wavelength of light is related to its

perceived color

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Structure of the Eye

The eye works like a camera, using a lens to focus light onto a photo-sensitive surface at the back of a sealed structure.

Light rays

CorneaPupil

Blind spotOpticnerve

RetinaFovea (pointof central focus)

Lens

Iris

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Organization of Retina5 cell types

Photoreceptorsrods and cones

Horizontal Cell Bipolar Cell Amacrine Cell Ganglion Cell

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Organization of Retina

Toopticnerve

Ganglioncell

Amacrinecell

Bipolarcell

Horizontalcell

Cone Rod

Light

Light

Cross section of retina shown vastly magnifiedin the diagram to the right

Photochemical is located hereBack of the eye

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Function of PhotoreceptorsThe photoreceptors transduce the energy

in light into a neural responseThis occurs when light entering the eye is

absorbed by photopigment molecules inside the photoreceptors

When light interacts with the photopigment, it results in the photoreceptor becoming more negatively charged (hyperpolarization)

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Distribution of Rods and ConesCones - concentrated in center of

eye (fovea) approx. 6 million

Rods - concentrated in periphery approx. 120 million

Blind spot - region with no rods or cones

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Distribution of Rods and Cones

Thou

sand

s of

rods

per

sq

uare

mill

imet

er

Blind spotFovea

60 40 20 0 20 40 60

180

140

100

60

20

0

180

140

100

60

20

060 40 20 0 20 40 60

Distance on retina from fovea (degrees)

FoveaBlind spot

Thou

sand

s of

con

es p

er

squa

re m

illim

eter

Distance on retina from fovea (degrees)

Fovea

Blind spot

Rods Cones

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Differences Between Rods and ConesCones

allow us to see in bright light allow us to see fine spatial detail allow us to see different colors

Rods allow us to see in dim light can not see fine spatial detail can not see different colors

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Receptive Fields and Rod vs. Cone Visual Acuity

Light

Spots of light Spots of light

Receptivefields

Ganglioncells

Bipolarcells

Photo-receptors(cones)

Photo-receptors(rods)

Pigmentedepithelium

Light

(a) Fovea (b) Periphery of retina

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Receptive Fields and Rod vs. Cone Visual AcuityCones - in the fovea, one cone often

synapse onto only a single ganglion cellRods - the axons of many rods synapse

onto one ganglion cellThis allows rods to be more sensitive in

dim light, but it also reduces visual acuity

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Color VisionOur visual system interprets differences in

the wavelength of light as colorRods are color blind, but with the cones

we can see different colorsThis difference occurs because we have

only one type of rod but three types of cones

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Color MixingTwo basic types of color mixing

subtractive color mixtureexample: combining different color paints

additive color mixtureexample: combining different color lights

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Additive Color MixtureBy combining lights of

different wavelengths we can create the perception of new colors

Examples: red + green = yellow red + blue = purple green + blue = cyan

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Trichromatic Theroy of Color VisionResearchers found that by mixing only three

primary lights (usually red, green and blue), they could create the perceptual experience of all possible colors

This lead Young and Helmholtz to propose that we have three different types of photoreceptors, each most sensitive to a different range of wavelengths

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Sensitivity Curves for the Three Types of Cones

Physiological studies revealed that Young and Helmholtz were correct

We have three types of cones

Light of different wavelengths will stimulate these cone types by different amounts

0.00

0.50

1.00

1.50

400 450 500 550 600 650 700

“Blue”cones

“Green”cones

“Red”cones

Wavelength in nanometers(billionths of a meter)

Rel

ativ

e re

spon

sive

ness

of

con

es

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Trichromacy and TVAll color televisions are based on the fact

that normal human color vision is trichromatic

Although we perceive the whole range of colors from a TV screen, it only has three colored phosphors (red, green, and blue)

By varying the relative intensity of the three phosphors, we can fool the visual system into thinking it is seeing many different colors

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Opponent Process Theory of Color VisionSome aspects of our color perception

are difficult to explain by the trichromatic theory alone

Example: afterimages if we view colored stimuli for an

extended period of time, we will see an afterimage in a complementary color

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ComplementaryAfterimages

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Opponent-Process TheoryTo account for phenomena like

complementary afterimages, Herring proposed that we have two types of color opponent cells red-green opponent cells blue-yellow opponent cells

Our current view of color vision is that it is based on both the trichromatic and opponent process theory

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Visual Pathway

RetinaOptictract

Opticchiasm

Opticnerve

Visual areaof the thalamus

Visualcortex

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Visual PathwayAxons of the ganglion cells come

together to form the optic nerveHalf of optic nerve fibers cross into

opposite hemisphere and synapse onto LGN (lateral geniculate nucleus)

LGN neurons synapse onto primary visual cortex

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Overview of Visual System The eye is like a camera, but instead of using film

to catch the light we have rods and cones Cones allow us to see fine spatial detail and color,

but can not function well in dim light Rods enable us to see in dim light, but at the loss

of color and fine spatial detail Our color vision is based on the presence of 3

types of cones, each maximally sensitive to a different range of wavelengths