Jim Luby, Breeding Team Leader, Univ. of Minnesota

RosBREED

Demonstration

Breeding Programs

Outline of Presentation

Intro to Demonstration Breeding Programs

and Traits for each RosBREED Crop Strawberry

Peach

Tart Cherry

Sweet Cherry

Apple

Reference Germplasm Sets

Tart and Sweet Cherries

• Tart Cherry - Amy Iezzoni- Michigan State University

• Sweet Cherry - Nnadozie Oraguzie- Washington State University

RosBREED Cherry Phenotyping

• Sweet Cherries for Fresh Market in Pacific

Northwest with firmness and a range of colors

• Tart Cherries for Processing in Michigan

– Fruit size genes to enable leaf spot introgression

from wild relative

– Freestone for better pitting

Strawberry

• Chad Finn - USDA-ARS, Corvallis, OR

• Jim Hancock - Michigan State University

• Tom Davis - University of New Hampshire

• Phil Stewart - Driscoll Strawberry Associates

RosBREED Strawberry Phenotyping

• Entire germplasm set evaluated at multiple

locations

• Allows for dissection of GxE effects in PBA

• Includes fresh market and processing fruit traits

• Plant architecture and remontancy (rebloom)

Peach

• Ksenija Gasic - Clemson University

• Tom Gradziel – UC-Davis

• John Clark - University of Arkansas

• Dave Byrne - Texas A&M University

RosBREED Peach Phenotyping

• Covers southern and western geography of US peach

production, range of chilling zones

• Several market types

– Fresh, Processing

– Nectarine, acidity/sugar, flesh color, shape

• Disease resistances (bacterial spot, brown rot)

Apple• Kate Evans - Washington State Univ.

• Susan Brown - Cornell University

• Jim Luby - University of Minnesota

RosBREED Apple Phenotyping

• Primary emphasis on fresh market traits

• Components of Flavor, Texture and Appearance

• Postharvest life and storage disorders

Traits and Standardized Phenotyping

Protocols

• Identify critical fruit quality traits and other

important traits

• Develop standardized phenotyping

protocols to enable data pooling across

locations/institutions

• Protocols available at www.RosBREED.org• “Fruit Evaluation”

Reference Germplasm SetsCrop Reference Set (CR Set)

Breeding Pedigree Set (BP Set)

Crop Reference Set (CR Set)• ~480 individuals (cultivars, ancestors, founders,

breeding lines, selections, and seedlings) that

are fruiting in 2010-2012

• Enable efficient validation and utility assessment

of Marker-Locus-Trait associations

• Genotyped genome-wide with SNP markers and

phenotyped for fruit quality traits and other high-

impact traits.

• Resource for common benefit

Breeding Pedigree Sets (BP Set)

• ~100-300 extra plants for each breeding program

that, together with the CR Set, allow full

representation of important parents in that

program

• Information may remain proprietary with each

breeding program; data-sharing improves power

Crop Reference Set

Visualizing Structure in a Breeding Program

with PediMapTM

Tart Cherry Crop Reference Set:Visualizing a Breeding Program

• Up to 5

generations

• Important

ancestors:

– English Morello

– Meteor

• Pedigree linkage

4 crosses

containing 201

individuals

provide

representation

of important

parents

39 individualsHalf are selections

20 Ancestors

unavailable

Tart Cherry Crop Reference Set:Visualizing a Breeding Program

Crop Reference Set

Allele Representation

Strategy for Constructing

Reference Germplasm Sets

Pedigree-linked reference germplasm sets represent alleles

across the genomes of Important Breeding Parents

Each relative represents an Important Parent; goal was

>12.5 representation units (e.g. 25 F1 s) for statistical power

• Offspring represent alleles of these Parents

• Other relatives can also represent Parent alleles through

shared ancestors (because a proportion of alleles will be Identical By

Descent)

F1 offspring, full-sibs, and parents provide best genome

coverage

Sweet Cherry Crop Reference Set

BlackRepublican

EmperorEugenie

EmperorFrancis

Napoleon

Windsor

Kordia

Bertoille

Corum

Eugenie

GilPeck

Katalin

Margit

NY54

Rube

Schmidt

Schneiders

Bing

Van

Lambert

JI

Sam

PMR-1

StMargaret

BlackEagle

Pop_KordiaxUNK

Rainier

Stella

Summit

Beaulieu

Regina

EarlyBurlat

Pop_BingxPMR1

Pop_PMR1xVan

P8-79

47-4

46-11

Benton

Glacier

Cashmere

Chelan

Newstar

GG

Pop_RainxPMR1

BB

DD

MIM17

Lapins

Brooks

Tieton

Ambrunes

Pop_RainxReg

Selah

7903-2

8007-2

Sweetheart

Pop_DDxMIM17

Pop_LapxBrooks

Pop_LapxTieton

Pop_RainxBent

Pop_LapxChel

Pop_LapxAmb

Pop_SelahxVan

Pop_7903-2xSumm

Pop_8007-2xChln

Pop_Swthtx8007

Pop_GGxSwtht

Pop_SwthtxBB

Pop_SwthtxReg

Lapins

• Up to 6 generations

• 49 individuals

– 38 cultivars

– 11 selections

• 18 crosses

containing 191

individuals (5 to 24

per cross)

• „Lapins‟ is an

Important Parent

No.

trees

14 13 8 101 1

1

Population 2 Population 3 Population 4

Lapins

Selection A Selection B Population 1

Direct descendants of Lapins each

provide 0.5 units allele representationPedigree diagrams are

outputs of Pedimap program

Representation by offspring in CRS

0.5 0.5 0.516 * 21 * 10 *

Lapins represents itself 1.0

BlackHeart

Lambert JI2420 EmpressEugenie

Van

Selection A Selection B Population 1 Population 2 Population 3 Population 4

Stella

0.5

Napoleon

0.5

0.5

cv Emperor Francis

represents Lapins at 0.125

units

(0.5 x 0.5 x 0.5 x 1

individual)

Emperor Francis

Immediate

ancestors of

Lapins

Representation by other relatives

Lapins

BlackHeart

Lambert JI2420 EmpressEugenie

Van

Selection A Selection B Population 1 Population 2 Population 3 Population 4

Stella

0.5

Napoleon

0.5

0.5

Population 5 represents

Lapins at 1.5 units

(0.5 x 0.5 x 0.5 x 0.5 x

24 offspring)

Population 5

Emperor Francis

24

0.5

Representation by other relatives

Lapins

Lapins

Stella and relatives

Direct descendants

(16 units)

27 units

12 units 11 units

7 units

Total representation of Lapins

in the sweet cherry CR Set

Van and relatives

Lapins is represented at 58

units in the sweet cherry Crop

Reference Set

Crop Reference Set

Integrating Multiple Breeding Programs Apple Peach Strawberry

Apple Crop Reference Set

• Need to integrate 3 breeding programs

• Much overlap of Important Parents among

3 breeding programs

• Maximum 6 generations

• Results in CR Set with much commonality

and high pedigree connectedness

• Allows transfer of information among

breeding programs

U of MN

35 Important

Parents

McIntosh

Delicious

NorthSpy

P14-126

EsopSpitz

Frostbite

GoldenDel

DuchOlden

Cox

LadyWill

YelTransp

Malinda

Wealthy

BenDavis

Montgomery

BlushiGol

Coop-17

Linda

NY247

NY66305-289

F2_26829-2-2

Williams

Raritan

Hudson

MN1802

Monark

PitmPine

Mflor821

MN1706

Winesap

WintBanana

YellNewtown

MN1760

GMAL4327

GMAL4328

CrimsCrisp

Dayton

PixieCrunch

Empire

NY55158-002

Jonathan

Fuji

RomBeauty

MN1628

Keepsake

Sweet16

Clivia

KidsOrRed

Braeburn

CrippsPink

Sundowner

PRI1661-1

PRI1661-2

Wagener

Mantet

Oriole

Fireside

Goodland

WorcPearm

JamesGr

Melba

Haralson

Cortland

Lodi

Delblush

Wijcik

Suncrisp

GingerGold

Beacon

Macoun

Spartan

Regent

MN1627

RedBaron

Goldrush

Prima

GMAL4332

IngMarie

Dolgo

Fantazja

NY76406-007

CoromandelRed

BeautBath

Cameo

Ambrosia

PRI14-126

Sundance

Tsugaru

NJ53

Redfree

Blackjon

RussianSeedling

Crandall

MN1708

Honeycrisp

Monroe

Melrose

Pinova

Gala

Splendour

Enterprise

Idared

StateFair

MN1691

Akane

Lambourne

Jonamac

VistaBella

Honeygold

Sunrise

EarlyCortland

NY06

NY92607-16

Hatsuaki

Jonagold

NY543

MN1702

MN1789

Liberty

MN1764

NY65707-19

Freedom

Jonafree

Snowsweet

Britemac

NJ90

Coop-15

GMAL4329

Fortune

MN1839

AE0213

CornellPop07

Elstar

Kerr

Sawa

NY09

Discovery

NY59333-006

669NJ5

NY88

AutumnCrisp

AurGoldGa

PacQueen

PacRose

Arlet

Nicola

WSU2

BC8S-27-43

BC-8S-27-2

Chinook

Zestar

Sansa

GranSmith

Rubin

Vanda

NY632

Empress

Burgundy

Silken

Sonya

Scifresh

WSU24

WSU37

NY04

NY05

NY07

CornellPop08

NY11

WSU46

Delorgue

Fiesta

MN1915

MN1942

MN1955

MN2034

MN1965

MN2062

MN1971

MN1980

MN2036

MN1984

MN2056

MN2057

MN2063

WSU34

WSU65

NY13

MN1988

MN2031

WSU38

AE_HCxJonf

AE0118

Pop_HCxCP

Pop_HCxEnt

MN1899

MN1964

MN1837

MN1888

MN1836

NY913

WSU7

WSU5

AE0210

NY70712-53

Coop-18

NY752

NY03

NY02

Pop_AurxCP

Pop_AurxEnt

Pop_AurxArl

Pop_AurxHC

WSU49

WSU50

WSU52

WSU57

AE0251

AE0206

WSU30

WSU17

WSU36

CornellPop13

Topaz

WSU35

WSU48

MN2016

Pop_EntxArl

Pop_HCxAr

MN2000

MN1937

MN1940

MN2042

AE0406

MN1914

MN1925

MN2033

MN2041

MN2044

MN2050

MN2046

NY10

NY08

NY241

NY01

NY14

NY15

CornellPop09

NY12

Apple Crop

Reference Set

McIntosh

Delicious

NorthSpy

P14-126

EsopSpitz

Frostbite

GoldenDel

DuchOlden

Cox

LadyWill

YelTransp

Malinda

Wealthy

BenDavis

Montgomery

BlushiGol

Coop-17

Linda

NY247

NY66305-289

F2_26829-2-2

Williams

Raritan

Hudson

MN1802

Monark

PitmPine

Mflor821

MN1706

Winesap

WintBanana

YellNewtown

MN1760

GMAL4327

GMAL4328

CrimsCrisp

Dayton

PixieCrunch

Empire

NY55158-002

Jonathan

Fuji

RomBeauty

MN1628

Keepsake

Sweet16

Clivia

KidsOrRed

Braeburn

CrippsPink

Sundowner

PRI1661-1

PRI1661-2

Wagener

Mantet

Oriole

Fireside

Goodland

WorcPearm

JamesGr

Melba

Haralson

Cortland

Lodi

Delblush

Wijcik

Suncrisp

GingerGold

Beacon

Macoun

Spartan

Regent

MN1627

RedBaron

Goldrush

Prima

GMAL4332

IngMarie

Dolgo

Fantazja

NY76406-007

CoromandelRed

BeautBath

Cameo

Ambrosia

PRI14-126

Sundance

Tsugaru

NJ53

Redfree

Blackjon

RussianSeedling

Crandall

MN1708

Honeycrisp

Monroe

Melrose

Pinova

Gala

Splendour

Enterprise

Idared

StateFair

MN1691

Akane

Lambourne

Jonamac

VistaBella

Honeygold

Sunrise

EarlyCortland

NY06

NY92607-16

Hatsuaki

Jonagold

NY543

MN1702

MN1789

Liberty

MN1764

NY65707-19

Freedom

Jonafree

Snowsweet

Britemac

NJ90

Coop-15

GMAL4329

Fortune

MN1839

AE0213

CornellPop07

Elstar

Kerr

Sawa

NY09

Discovery

NY59333-006

669NJ5

NY88

AutumnCrisp

AurGoldGa

PacQueen

PacRose

Arlet

Nicola

WSU2

BC8S-27-43

BC-8S-27-2

Chinook

Zestar

Sansa

GranSmith

Rubin

Vanda

NY632

Empress

Burgundy

Silken

Sonya

Scifresh

WSU24

WSU37

NY04

NY05

NY07

CornellPop08

NY11

WSU46

Delorgue

Fiesta

MN1915

MN1942

MN1955

MN2034

MN1965

MN2062

MN1971

MN1980

MN2036

MN1984

MN2056

MN2057

MN2063

WSU34

WSU65

NY13

MN1988

MN2031

WSU38

AE_HCxJonf

AE0118

Pop_HCxCP

Pop_HCxEnt

MN1899

MN1964

MN1837

MN1888

MN1836

NY913

WSU7

WSU5

AE0210

NY70712-53

Coop-18

NY752

NY03

NY02

Pop_AurxCP

Pop_AurxEnt

Pop_AurxArl

Pop_AurxHC

WSU49

WSU50

WSU52

WSU57

AE0251

AE0206

WSU30

WSU17

WSU36

CornellPop13

Topaz

WSU35

WSU48

MN2016

Pop_EntxArl

Pop_HCxAr

MN2000

MN1937

MN1940

MN2042

AE0406

MN1914

MN1925

MN2033

MN2041

MN2044

MN2050

MN2046

NY10

NY08

NY241

NY01

NY14

NY15

CornellPop09

NY12

Apple Crop

Reference Set

Wash. State

38 Important

Parents

Cornell

18 Important

Parents

McIntosh

Delicious

NorthSpy

P14-126

EsopSpitz

Frostbite

GoldenDel

DuchOlden

Cox

LadyWill

YelTransp

Malinda

Wealthy

BenDavis

Montgomery

BlushiGol

Coop-17

Linda

NY247

NY66305-289

F2_26829-2-2

Williams

Raritan

Hudson

MN1802

Monark

PitmPine

Mflor821

MN1706

Winesap

WintBanana

YellNewtown

MN1760

GMAL4327

GMAL4328

CrimsCrisp

Dayton

PixieCrunch

Empire

NY55158-002

Jonathan

Fuji

RomBeauty

MN1628

Keepsake

Sweet16

Clivia

KidsOrRed

Braeburn

CrippsPink

Sundowner

PRI1661-1

PRI1661-2

Wagener

Mantet

Oriole

Fireside

Goodland

WorcPearm

JamesGr

Melba

Haralson

Cortland

Lodi

Delblush

Wijcik

Suncrisp

GingerGold

Beacon

Macoun

Spartan

Regent

MN1627

RedBaron

Goldrush

Prima

GMAL4332

IngMarie

Dolgo

Fantazja

NY76406-007

CoromandelRed

BeautBath

Cameo

Ambrosia

PRI14-126

Sundance

Tsugaru

NJ53

Redfree

Blackjon

RussianSeedling

Crandall

MN1708

Honeycrisp

Monroe

Melrose

Pinova

Gala

Splendour

Enterprise

Idared

StateFair

MN1691

Akane

Lambourne

Jonamac

VistaBella

Honeygold

Sunrise

EarlyCortland

NY06

NY92607-16

Hatsuaki

Jonagold

NY543

MN1702

MN1789

Liberty

MN1764

NY65707-19

Freedom

Jonafree

Snowsweet

Britemac

NJ90

Coop-15

GMAL4329

Fortune

MN1839

AE0213

CornellPop07

Elstar

Kerr

Sawa

NY09

Discovery

NY59333-006

669NJ5

NY88

AutumnCrisp

AurGoldGa

PacQueen

PacRose

Arlet

Nicola

WSU2

BC8S-27-43

BC-8S-27-2

Chinook

Zestar

Sansa

GranSmith

Rubin

Vanda

NY632

Empress

Burgundy

Silken

Sonya

Scifresh

WSU24

WSU37

NY04

NY05

NY07

CornellPop08

NY11

WSU46

Delorgue

Fiesta

MN1915

MN1942

MN1955

MN2034

MN1965

MN2062

MN1971

MN1980

MN2036

MN1984

MN2056

MN2057

MN2063

WSU34

WSU65

NY13

MN1988

MN2031

WSU38

AE_HCxJonf

AE0118

Pop_HCxCP

Pop_HCxEnt

MN1899

MN1964

MN1837

MN1888

MN1836

NY913

WSU7

WSU5

AE0210

NY70712-53

Coop-18

NY752

NY03

NY02

Pop_AurxCP

Pop_AurxEnt

Pop_AurxArl

Pop_AurxHC

WSU49

WSU50

WSU52

WSU57

AE0251

AE0206

WSU30

WSU17

WSU36

CornellPop13

Topaz

WSU35

WSU48

MN2016

Pop_EntxArl

Pop_HCxAr

MN2000

MN1937

MN1940

MN2042

AE0406

MN1914

MN1925

MN2033

MN2041

MN2044

MN2050

MN2046

NY10

NY08

NY241

NY01

NY14

NY15

CornellPop09

NY12

Apple Crop

Reference Set

All Programs‟

72 Important

Parents

McIntosh

Delicious

NorthSpy

P14-126

EsopSpitz

Frostbite

GoldenDel

DuchOlden

Cox

LadyWill

YelTransp

Malinda

Wealthy

BenDavis

Montgomery

BlushiGol

Coop-17

Linda

NY247

NY66305-289

F2_26829-2-2

Williams

Raritan

Hudson

MN1802

Monark

PitmPine

Mflor821

MN1706

Winesap

WintBanana

YellNewtown

MN1760

GMAL4327

GMAL4328

CrimsCrisp

Dayton

PixieCrunch

Empire

NY55158-002

Jonathan

Fuji

RomBeauty

MN1628

Keepsake

Sweet16

Clivia

KidsOrRed

Braeburn

CrippsPink

Sundowner

PRI1661-1

PRI1661-2

Wagener

Mantet

Oriole

Fireside

Goodland

WorcPearm

JamesGr

Melba

Haralson

Cortland

Lodi

Delblush

Wijcik

Suncrisp

GingerGold

Beacon

Macoun

Spartan

Regent

MN1627

RedBaron

Goldrush

Prima

GMAL4332

IngMarie

Dolgo

Fantazja

NY76406-007

CoromandelRed

BeautBath

Cameo

Ambrosia

PRI14-126

Sundance

Tsugaru

NJ53

Redfree

Blackjon

RussianSeedling

Crandall

MN1708

Honeycrisp

Monroe

Melrose

Pinova

Gala

Splendour

Enterprise

Idared

StateFair

MN1691

Akane

Lambourne

Jonamac

VistaBella

Honeygold

Sunrise

EarlyCortland

NY06

NY92607-16

Hatsuaki

Jonagold

NY543

MN1702

MN1789

Liberty

MN1764

NY65707-19

Freedom

Jonafree

Snowsweet

Britemac

NJ90

Coop-15

GMAL4329

Fortune

MN1839

AE0213

CornellPop07

Elstar

Kerr

Sawa

NY09

Discovery

NY59333-006

669NJ5

NY88

AutumnCrisp

AurGoldGa

PacQueen

PacRose

Arlet

Nicola

WSU2

BC8S-27-43

BC-8S-27-2

Chinook

Zestar

Sansa

GranSmith

Rubin

Vanda

NY632

Empress

Burgundy

Silken

Sonya

Scifresh

WSU24

WSU37

NY04

NY05

NY07

CornellPop08

NY11

WSU46

Delorgue

Fiesta

MN1915

MN1942

MN1955

MN2034

MN1965

MN2062

MN1971

MN1980

MN2036

MN1984

MN2056

MN2057

MN2063

WSU34

WSU65

NY13

MN1988

MN2031

WSU38

AE_HCxJonf

AE0118

Pop_HCxCP

Pop_HCxEnt

MN1899

MN1964

MN1837

MN1888

MN1836

NY913

WSU7

WSU5

AE0210

NY70712-53

Coop-18

NY752

NY03

NY02

Pop_AurxCP

Pop_AurxEnt

Pop_AurxArl

Pop_AurxHC

WSU49

WSU50

WSU52

WSU57

AE0251

AE0206

WSU30

WSU17

WSU36

CornellPop13

Topaz

WSU35

WSU48

MN2016

Pop_EntxArl

Pop_HCxAr

MN2000

MN1937

MN1940

MN2042

AE0406

MN1914

MN1925

MN2033

MN2041

MN2044

MN2050

MN2046

NY10

NY08

NY241

NY01

NY14

NY15

CornellPop09

NY12

Apple Crop

Reference Set

Peach Crop Reference Set

• 4 Breeding programs• Limited overlap of germplasm

between market types

• 118 cultivars and selections

• 373 progeny in 23 crosses

• Up to 11 generations

JH_Hale

O'Henry

Flordaprince

Frank

OP11

Pallas

Bolinha

P97-14

CAF4

Nemaguard

ChineseCling

Greensboro

Everts

OP1

Andross

P.argentea

Hakuho

UFGold

White_County

ArkC

Blazeprince

Orange_Cling

Lola

ZinDai

Springtime

Mayfire

91,17-262

F8,5-166

Dr._Davis

BY00P6346-t

Jefferson

Peento

18,8-11

18,8-23

Lovell

Nonpareil

OP12

Diamante

Early_Crawford

Hiley

Mayflower

RioOsoGem

S-37

Admiral_Dewey

Okinawa

OP10

OP13

OP14

Panamint

Slappey

Kakamas

OldmixonFree

Babcock

LeGrand

F10C,12-28

Sunfre

Fairhaven

Halehaven

TropicBeauty

P34-106

Saturn

Redskin

CAF2

S-37OPxNemaguard

Georgia_Belle

Rizzi

2001,7-180

BY01P6160

ArkB

SC-POP22

Dixon1

St_John

Marigold

SC-POP_Blush

Springold

Springbrite

Riegels

Crimsonbaby

CA-Pop1

CA-Pop3

SC-POP23

Redhaven

Loring

A34-160

Winblo

TXW1293-1

Guardian

Cumberland

CA-Pop2

SC-Pop3

AR-Pop1

Candor

Goldprince

Y142-75

Contender

TX2B136

SC-POP_G

Summercrest

Clayton

TX2293-3

CAF3

BY01P6245

China-Pearl

TX-Pop3

TX-Pop1

BY03P22388

TX-Pop2

SC-Pop2

SC-POP09

SC-Pop1

Peach Crop Reference SetComplete Pedigree

NJ5107397

Panamint

Jewel

FLA2-13

Palomar

FLA68-50

Okinawa

Early_Crawford

ChineseCling

OP43

OP36

NJ53939

OP19

FLA16-61

FLAR9T10

Earligold

Carman

Slappey

dummy2

OP29

Luken's_Honey

July_Elberta

Strawberry

Peento

Mayflower

FLA5-58

OP10

Sunred

FLA28-48

Admiral_Dewey

OP35

OP13

OP4

Lola

OP44

Hiley

Quetta

Lippiatt

Levi

OP38

OP24

OP23

NJN14

Stock3-17

Pallas

OP21

DukeofGA

K173-9

OP2

OP28

OP11

OP22

OP26

OP12

OP32

OP41

Lemon_Free

PI35201

OP48

OP37

OP46

PI91459

PI101681

Honeydew_Hale

OP40

Splendid

Bolivian

OP45

OP39

OP31

Greensboro

DukeofYork

Reina_Elena

5105965

Hann_Almond

OP47

Mexican_Honey

OP25

Lovell

AustralianMuir

Orange_Cling

Transvaal-Cling

GoodmansChoice

Alameda

RoundTuscan

Nonpareil

Jordanolo

GoldenGlory

Bonanza

OP17

E7-35

dummy1

Hakuto

Denjuro

OP7

Diamante

OP15

Yellow_Alberge

Erly-Red-Fre

OP16

OP30

PI,292557

18,8-11

OP5

R3-53

Ross

R1-1

Tuscan

Leader

OP27

ZinDai

Fla.79-20C

OP8

OP9

PI134401

Bolinha

dummy93

dummy94

dummy92

H_523

Jefferson

Ark_154N

OP3

RedGold

2W_68W

RR_53-194

Jing

OP14

OP6

D62-193

OP1

Halford

Ogawa

19,1-40

StHelena

dummy98

P.mira#19

Klamt

F8,1-71

P.scoparia

P.argentea

18,8-23

TX86606

P55-454

S-37

dummy89

Nemaguard

dummy90

dummy91

D6-15W

CarolynG

Klampt

9,20-C

persicaXdavidiana

Yumyeong

OldmixonCling

dummy97

LateCrawford

dummy99

F8,5-104-146

Merrill_Prince

OP20

Dix22A-5

dummy96

OP18

PP904

dummy95

OP33

OP34

OP42

VP

VP

VP

PanaxJewel

FLA4-65

FLA8B27

EarlyCrawford

FLA3-56

NJ40CS

St_John

Luken'shoneyxJElberta

StrawberryxPeento

Maravilha

Goldfinch

Meredith

Nectared4

Nectared7

Early_Hale

125936

5109028

Georgia_Belle

562158

Loadel

Paloro

Dixon1

Dixon2

Dix,58-6

Dix,16-3

NonpareilBF

TardyNonpareil

54P455

TachibanaWasa

FLA84-18C

OrangeClingSdlg

E22-59N

87,13-13

90,9-116

90,9-134

Lilliland

Leader_sdlg

Lilleland

Fla.86-28C

Oro_A

HoneydewHale

A419-P

Carson

D,62-193

LateRoss

Bolinha3

Bolinha6

Bolinha7

Kakamas

2000,16-133

2000,15-116

2000,15-119

2000,15-122

2000,15-126

2000,15-127

2000,15-73

2000,15-88

S-37OP

2000,3-217

OldmixonFree

RioOsoGem

99,19-23

Nickels

Brights

Hansen1

Hansen2

F8,5-55

Ross*R,1-1

FLA4695

VP

FLAQ3033

Kaygold

Elberta

FLA35-30

Southhaven

Babcock

NJN55

R1T6

561483

Cumberland

113115

Dix,22A-5

Dix,8

TardyxJordan

Hakuho

Dix,6-6

E22-59

TuscanxPaloro

(TxP)

FLA92-2C

NJ257

A130P

89,10-44

89,10-53

89,10-55

89,10-59

89,10-60

89,10-61

Wolvamade

F8,76-45

Walgant

88,2-75

Dixon

88,2-77

S-37OPxNemaguard

2005,10-10-90

2000,2-8

2000,2-9

91,9-161

93,1-75

CA-Pop-7-4-176

CA-Pop-5-9-248

CA-Pop-5-9-76

CA-Pop-9-1-198

CA-Pop-7-3-204

CA-Pop-7-13-72

CA-Pop-7-11-108

CA-Pop-7-11-24

CA-Pop-7-5-207

CA-Pop-7-8-68

CA-Pop-8-15-77

CA-Pop-7-13-63

CA-Pop-5-17-192

CA-Pop-8-9-165

CA-Pop-9-4-49

CA-Pop-9-7-244

2001,1-185

2001,2-4

2001,2-84

2001,2-93

2002,9-76

2002,9-8

89,15-309

92,7-25

CA-Pop-5-1-223

CA-Pop-5-17-147

CA-Pop-8-3-157

CA-Pop-9-1-150

CA-Pop-7-12-126

VP

VP

FLA15-85W

JH_Hale

FLA10-48

FLA2-7

Robin

K6E121

Arp_Beauty

RR38-4-662001

Frank

Valiant

RR105-10--

Jungerman

Dix,5A-1

F8,72-33

Da30-13

7-5

Fortuna

Ark_392

A270N

Woltemade

2000,12-110

2000,12-56

96,9-292

F10C,20-51

92,10-119

93,5-220

93,5-24

93,5-26

93,5-50

92,10-125

93,5-80

96,9-8

94,4-223

96,7-165

Padre_al

Guardian

CA-Pop-9-14-48

CA-Pop-9-20-174

CA-Pop-9-31-182

CA-Pop-9-31-301

CA-Pop-9-32-253

93,5-19

CA-Pop-10-2-8

CA-Pop-5-18-181

CA-Pop-7-10-3

CA-Pop-7-10-198

2004,2-142

2004,2-148

2004,2-150

2000,6-38

VP

Candoka

Sunhigh

Fairhaven

Halehaven

Springtime

FLA3-2

Flordaprince

104325

Marigold

JHHxQuetta

JHHxLippiatt

JHHxLevi

NJ66325

76925

C1-19-122--

58127

Summercrest

Raritan_Rose

94727

38026

6128

Everts

F10C,12-28

Kalhaven

Redskin

D25-9E

GH2-48

Andross

ArkC

90,1-4

96,4-49

96,4-51

96,4-55

96,5-11

CA-Pop-7-18-192

CA-Pop-6-5-95

CA-Pop-9-1-265

CA-Pop-9-20-1

CA-Pop-9-5-82

CA-Pop-9-7-274

CA-Pop-9-7-91

CA-Pop-9-14-25

Da,30-13

2001,3-303

2000,15-241

2000,B-202

2000,11-177

2001,7-256

2000,8-64

F8,72-22

SC-POP_G

VP

NJN21

Southland

TropicBeauty

Sunrise

PalxSprtime

Golden_Globe

Fireglow

LeGrand

Kim

Merrill.Bon

Loring

Jerseyland

Prarie_Dawn

NJ196

Cherryred

13232

NJ156

98838

NJLA3

Riegels

Vilmos

Redhaven

NC7130

G40-5E

Winblo

Norman

ArkB

Rizzi

H,6-55

F8,1-42

F8,1-96

E,1-16

F10E,1-16

89,6-103

91,12-4

91,13-26

91,13-7

96,1-148

96,1-149

2000,3-205

2001,7-180

CA-Pop-5-9-231

CA-Pop-9-17-197

18,9-8

91,12-54

2002,3-236

2002,3-124

2002,3-128

2002,1-50

Sunlite

FV131-48

TXW1293-1

G69-83N

Armking

FV15-48

KimxJuly_Elberta

602903

O'Henry

Armqueen

NJ230

Babygold7

5110301

5102896

13232xCherryred

5110007

5106543

Hesse

KimxLeGrand

Candor

FLA8-11

Dr._Davis

18,6-33

NC7366

Pekin

RR122-15

18,9-41

89,9-82

90,10-120

97,2-19

92,14-73

97,5-270

2001,7-126

2000,10-223

2000,1-215

2000,1-222

2000,1-228

2000,1-234

2000,B-217

2000,B-218

2000,B-219

2000,3-159

CA-Pop-5-19-168

CA-Pop-5-19-79

CA-Pop-7-1-1

CA-Pop-5-20-139

CA-Pop-7-13-245

CA-Pop2

CA-Pop-5-18-246

CA-Pop-5-18-11

97,5-152

98,4-24

CA-Pop-8-47-55

CA-Pop-8-52-11

CA-Pop-9-23-55

2000,11-58

99,15-99

98,2-132

CA-Pop-5-16-209

CA-Pop-5-9-33

VP

VP

Columbina

Springbrite

K61-48

FV89-14

SunGrand

Saturn

K519-47

Mayfire

554039

555036

554367

562025

582990

90,10-91

Red_Grand

FLAK5W35

NCA2219

DrDavis

91,17-195

NCA2679

BY81P2840

Ark_145N

87,12-31

F8,1-121

Clayton

2000,8-44

2000,16-92

2002,1-79

CA-Pop-8-5-100

CA-Pop-9-22-229

CA-Pop-9-5-273

CA-Pop-7-13-130

CA-Pop-7-26-67

CA-Pop-6-4-51

CA-Pop-7-19-164

CA-Pop-7-19-20

CA-Pop-7-27-1

CA-Pop-7-2-73

CA-Pop-8-33-7

CA-Pop-8-4-99

CA-Pop-7-2-209

CA-Pop-7-16-218

CA-Pop-7-17-1

KimxJelberta

CA-Pop-5-16-146

CA-Pop3

VP

VP

FLA3-4N

FLA5-5

P83-48

Springcrest

Springold

Sungrand

A34-160

FV317-97

FV9-46

P51-138

Crimsonbaby

C1-25-177--

RR51-32--

RR71-322--

90,10-92

Early_Sun_Grand

FLNCseedling

91,18-6

Goodwin

NC64

Blazeprince

Ark_224

91,17-262

93,3-108

93,3-22

93,3-225

93,3-247

93,3-159

93,4-166

93,4-185

93,3-175

93,3-189

99,12-155

CA-Pop-5-22-204

2005,22-166

2005,23-6

97,9-200

97,9-273

98,8-188

99,16-131

BY03P22388

2003,1-329

2005,10-223

2005,10-228

2005,10-236

2005,11-56

CA-Pop-5-10-61

CA-Pop-5-7-168

CA-Pop-5-8-229

2001,9-130

2001,9-215

2001,9-65

CA-Pop-10-1-3

CA-Pop-8-28-264

CA-Pop-9-24-286

Hermosillo

P97-14

P60-38

FV3-257

P34-106

NJ271

D2-46-143--

F8,5-166

Red_Diamond

FLA9-20C

F8,5-147

F8,5-156

F8,5-161

F8,5-162

F8,5-164

F8,5-165

F8,5-171

F8,5-172

F8,5-179

F8,5-182

2001,18-215

Contender

BY96P3240

SC-POP22

Ark_433N

96,1-171

99,4-123

98,9-7

98,4-177

96,9-171

CA-Pop-5-1-78

CA-Pop-5-4-228

CA-Pop-5-5-248

CA-Pop-10-1-5

CA-Pop-10-3-5

CA-Pop-8-61-26

2002,15-255

2001,19-6

SC-Pop1

CA-Pop-5-17-24

CA-Pop-5-2-200

CA-Pop-9-9-250

CA-Pop-8-13-154

CA-Pop-8-4-112

CA-Pop-8-18-11

CA-Pop-8-42-67

CA-Pop-10-1-7

CA-Pop-10-1-9

CA-Pop-10-1-11

CA-Pop-10-2-6

2000,11-157

2001,9-275

2001,9-305

2001,9-6

99,15-145

VP

VP

TX2B136

P42-81

P42-91

Goldprince

Y142-75

P87-79

P37-79

NJ291

NJF8

2000,2-18

Crimson_Lady

UFGold

F85-147

2000,A-235

2000,3-157

2000,8-138

2000,8-149

2000,8-150

2000,8-153

2000,8-154

2000,8-157

2000,8-158

2000,8-164

2000,A-241

2000,16-93

2000,16-125

2000,16-110

2000,16-177

2000,2-15

BY01P6245

China-Pearl

BY00P6346-t

White_County

CA-Pop-9-34-69

2000,16-199

2000,16-202

2000,8-37

2000,8-69

2000,8-71

2000,A-151

2000,A-219

2000,A-220

2000,16-207

CA-Pop-9-34-301

2000,16-159

2000,16-209

F85-166

CA-Pop1

CA-Pop-5-21-145

CA-Pop-6-4-234

CA-Pop-9-34-227

CA-Pop-9-34-253

CA-Pop-10-2-10

CA-Pop-9-12-102

CA-Pop-9-23-152

CA-Pop-9-23-25

CA-Pop-9-28-284

CA-Pop-9-30-233

CA-Pop-9-31-99

CA-Pop-9-33-66

CA-Pop-9-34-105

CA-Pop-5-12-199

2005,12-231

2005,12-256

CA-Pop-9-22-282

CA-Pop-9-36-29

CA-Pop-5-11-227

2005,12-51

CA-Pop-9-20-126

CA-Pop-9-30-298

CA-Pop-5-12-88

CA-Pop-5-14-124

CA-Pop-9-20-301

2002,13-152

VP

Sunfre

TX2293-3

P87-147

D33-1

CrimsonLady

BY01P6160

CA-Pop-9-33-341

CA-Pop-7-25-178

CA-Pop-6-1-3

CA-Pop-6-1-33

CA-Pop-7-26-80

CA-Pop-7-10-235

CA-Pop-7-11-23

CA-Pop-8-35-81

CA-Pop-8-42-50

CA-Pop-8-55-50

CA-Pop-7-3-47

CA-Pop-7-9-28

CA-Pop-7-2-167

CA-Pop-8-17-71

CA-Pop-8-19-90

CA-Pop-8-25-99

CA-Pop-8-27-128

CA-Pop-8-30-54

CA-Pop-7-3-157

CA-Pop-7-7-146

CA-Pop-8-33-208

CA-Pop-7-6-82

CA-Pop-5-5-86

CA-Pop-7-17-214

CA-Pop-8-41-92

CA-Pop-7-17-29

CA-Pop-7-23-78

CA-Pop-7-9-218

CA-Pop-8-38-106

CA-Pop-7-13-254

CA-Pop-8-20-64

SC-POP_Blush

SC-Pop2

SC-POP09

SC-POP23

AR-Pop1

SprGoldxSunfre

P107-78

P83-62

P79-56

Y142-194

P7-34

2002,6-283

SC-Pop3

P57-56

P48-57

Y140-77

P78-49

CA-Pop-8-27-83

P30-122

P91-23

CAF4

Y150-13

CAF3

TX-Pop1

CAF2

TX-Pop2

TX-Pop3

Crop Reference and

Breeding Pedigree Sets

Allele Representation of Important Parents in

Multiple Breeding Programs

Allele Representation of 72 Important

Parents in Apple Germplasm Set

6

1013

6

CRS +UMN BPS35 Parents

10

8

CRS + Cornell BPS18 Parents

Deficient (<12.5 units)

Adequate (12.5-20 units)

Very good (21-50 units)

Excellent (>50 units)

6

17

15

CRS +WSU BPS38 Parents

Strawberry

Large Integrated Crop Reference and

Breeding Pedigree Set

Strawberry Crop Reference Set

• 26 generations

• Covers several germplasm groups- CA, PNW, Eastern US, Europe, Wild

• 209 cultivars and selections

• 78 wild strawberries - F. virginiana, F. chiloensis or F. virginiana x chiloensis

• 618 seedlings in 44 crosses

• Results in CRS with much commonality

and high pedigree connectedness

Strawberry Crop Reference SetHudson_Bay

dummy13

White_Carolina

dummy15

dummy14

dummy10

dummy7

dummy17

F.virg'Native_Iowa'

dummy6

dummy5

dummy11

King_of_the_Earlies

N.J.Scarlet(F.virg)

Belmont

Goliath

Jucunda

dummy2

OP

F.virginiana-M_Marshall

F_Marshal

Miner

Likely_Lake_Stevens

Likely_Chesapeake

Missionary

William_Belt

dummy1

Chesapeake

Jersey_Queen

dummy9

TD8

Markee

UnserFritz

F_Konig.von.Sachsen

dummy16

Bubach

Captain_Jack

Scot_BK-46

Cal_39.82-19

F_Fendall

Middlefield

dummy8

US_3328

Cal_BH_14_(Mixed_Cross)

WA_80053-17

WA_7531-22

Cal_43.1-36

Cal_85.92-602

Rockhill_Derivative

dummy4

F1_hybrid

Streamliner

Early_Jersey_Giant

Montana_Missionary

S0_20-2

MN_1181

Joe

Glendale

MDUS_5132

NY_113

FL_83-418

FL_84-1389

FL_84-3163

FL_83-1452

Harvester

Formans_Excelsior

Pocomoke

Early_Ozark

Cal_75.121-101

MN_2374

NJ_8713-8

LA_59-2012

LA_59-138

FL_80-456

F.moschata

Princesse_Dagmar

T._de_Leopold

Reiko

Kurume103

NJ_821-5

NJ_459

Hummi_Gento

Uberreich

Sel.Salvi_N.15

US_542

Sitkahybrid

46-4J

Ottawa485

J.23

Beaver

Berri_Supreme

F.chiloensis-M_Sitka

F_Sitka

SYS_3672-179

Red_Giant

F_JH_101-1

J._Riffaud

Madame_Lebreton

Miyazaki

The_Sun

F_ORUS_1407-76

B6.117

D5.23

G7

G8

L1

BC6

H4

Palmyre

Chilean

British_Queen

Cal_87.112-6

Cal_88.270-1

Cal_94.16-1

FL_82-1556P

LA_2556

LA_883

General_de_Castelnau

St._Joseph

Lucie_Boiselet

Everglade

J92D12

BCPink

British_Sovereign

Cal_45.5-1

dummy12

dummy18

dummy3

Frel

FRENCHBIN

Hawaii4

Heidi

Hummi_Ferma

Lake_Stevens

Liberation_d'Orleans

Marajox

Ooishi-shikinari2

SARGENTDIPLOIDBIN

St._Claude

UNK

Methven_Scarlet

Keens_Imperial

Downer's_Prolific

F.chiloensis

F.vesca

Marshall

Aberdeen

Portia

Parry

Konig.von.Sachsen

Fendall

Nich_Ohmer

F.virg.glauca.Wasatch

Cal_60.19-2

F.virginiana

Sarian

Bemidji_Chief

Gandy

B753

B754

FL_87-418

FL_87-200

Cassandra

Vanguard

Tangi

Florida_90

Capron

F.cuneifolius

F.virginiana_ovalis

(Sitkahybridx46-4J)

Sitka

General_Chanzy

(Miyazaki_x_The_Sun)

Crown

Joe_Reiter

LB48

Duc_de_Malakoff

Diamante

Aromas

LA_8311

Madame_Raymond_Poincare

Evie-2

Keens_Seedling

Charles_Downing

F.chiloensis(Chile)

F.chiloensis(Ecuador)

F.chiloensis(PatPoint)

F.chiloensis(Peru)

F.chiloensis(Rockaway#3)

F.chiloensis(ScottsCreek)

F.chiloensis10-B

FRA_1075

FRA_1088

FRA_1092

FRA_1100

FRA_1104

FRA_1108

FRA_1267

FRA_1691

FRA_1692

FRA_24

FRA_34

FRA_357

FRA_368

FRA_372

FRA_42

FRA_48

FRA_688

FRA_743

FRA_796

OSC_812

Yaquina_B

Del_Norte

F.chil.Cape.Mendicino

Pawtuckaway

Yellow_Wonder

Alpine(F.vesca)

Auchincruive_Climax

Kaiser_Samling

F.virginiana(AL)

F.virginiana(MS)

F.virginiana(NC)

F.virginiana(SC)

FRA_110

FRA_1435

FRA_1455

FRA_1466

FRA_1694

FRA_1695

FRA_1696

FRA_1697

FRA_1698

FRA_1699

FRA_1700

FRA_1701

FRA_1702

FRA_1703

FRA_338

FRA_58

FRA_982

Lion's_Head_3

MS_34-12

MS_34-4

Sable_Beach_8

y32b

Lupton

Rosa_Linda

Ville_de_Paris

Cal_134.31

((Sitkahybridx46-4J)xOttawa485)

Washington

JH_101-1

Fukuba

Commander

LB48xH4

Docteur_Morere

Albion

Pelican

Sans_Rivale

Hovey

Ross_Phoenix

Sharpless

Ettersburg_121

Huachi(Ambato_NAH)

NJ_selection

Souvenir_de_Charles_Machiroux

MAR1A

ORUS_2161-1

BRA1A

Fred9xLH50-4

RH30xMR10

RH30xLH50-4

SCxFRA24

MR10xRH23

MR10xN8688

Hogyoku

Benizuru

ORUS_1407-76

ORUS_2361-1

Yellow_WonderXPawtuckaway

FRA_1557

FRA_1408

FRA_1414

FRA_1620

FRA_1580

Wilson

Crescent

Green_Prolific

Jessie

OSC_154

(SharplessxParry)F3

Fendalcino

Claribel

Laxtons_Noble

ORUS_2075-1

Oberschlesien

SCXMAR1A

NAHxBRA1A

SCxNAH

HM1xNAH

FVC_18-1

FVC_18-18

FVC_30-1

FVC_30-2

FVC_30-3

FVC_30-4

FVC_30-5

FVC_8-1

FVC_8-2

FVC_8-3

FVC_8-4

FVC_8-5

FVC_17-20

FVC_16-1

Neunan

Haverland

Ruby

Cumberland_Triumph

Oregon

OSC_7

Rose_Ettersburg

Van_Deman

Clark

Cal_Z_11

Cal_Z_9

Prinz_Julius_Ernst

FVC_11-1

FVC_11-2

FVC_11-3

FVC_11-4

FVC_11-5

FVC_11-6

FVC_33-1

FVC_10-1

FVC_10-2

FVC_28-1

Hoffman

Seedling#4Howard

Cyclone(Old)

Dunlap

OSC_18

OSC_59

WSU_14

Corvallis

Ettersburg_20

Bismarck

Excelsior

Clyde

Eurisko

Pan_American

Lulu

Ettersburg_80

Easypicker

Laxton's_Noble

Noble

Seedling#1Howard

Redheart

Progressive

Duluth

Pickerproof

Royal_Sovereign

Howard_17

Sieger

US_836

US_3358

Klondike

Cal_145.38a

Cal_145.52

Rockhill

Hansa

Gem

Burgundy

Narcissa

NJ_1051

US_543

US_634

Dorsett

Fairfax

Suwannee

Tenn_Beauty

Blakemore

Midland

Senga_Sengana

OSC_767

US_253

NY_4626

Southland

Pathfinder

Deutsch_Evern

Valentine

Cal_43.2

Glenheart

Rockhill_selfed

Twentieth_Century

Trumpeter

Robinson

OSC_346

Sparkle

Brightmore

OSC_456

US_3763

Redgauntlet

Talisman

US_3374

Fairland

MDUS_430

Temple

OSC_1490

OSC_1491

OSC_305

OSC_821

Fairpeake

Redstar

MDUS_683

Cal_21.9

Cal_86.6

OSC_1629

WSU_1149

US_253_OP

Cal_7.20

Cal_7.9a

Cal_720

Cal_86.5

Cal_68.24

Cal_68.25

Cal_135.18

Cal_179.15

Cal_179.2

Cal_67.5

Cal_10.21

NC__1065

Massey

Tenn_Shipper

WSU_828

WSU_438

WSU_450

Cal_144.2

Cal_144.21

Cal_32.50

Cal_32.63

Cal_66.2

NY_316

Red_Rich

Redrich

OSC_913

OSC_1366

NJ_953

NJ_925

Klonmore

Rotkappchen

US_2506

US_2267

Ada_Herzberg

NC_1729

(FairfaxXF.ovalis)

Cheyenne2

Cal_74.3

BxD

Cal_143.32

Koro103

Crimson_King

OSC_1765

Northwest

Juspa

OSC_2136

OSC_1509

OSC_1757

OSC_1816

US_4127

MDUS_1972

Sierra

Cal_161.1

OSC_2100

Cal_A.08

Cal_186.1

Cal_177.19

Cal_177.21

Cal_474.2

Cal_159.1

NC__1053

MDUS_2101

Stelemaster

WSU_157

WSU_175

Cal_564.6

Cal_564.7

Cal_337.2

Cal_337.3

Cal_249.5

Cal_222.11

Cal_222.4

Geneva

Earlidawn

Redglow

NY_254

OSC_1815

OSC_1776

(Tenn_Shipper_x_Midland)

Dixieland

Pocahontas

Jerseybelle

(Klonmore_x_Midland)

MDUS_2359

US_4144

Guardsman

Vibrant

Redcoat

Georg_Soltwedel?

US_3919

Macherauchs_Dauerernte

Repita

Veestar

Cal_213.4

Cal_378.16

Cheyenne3

(MidlandxCheyenne2)

Protem

Cal_543.14

Revada

Cyclone

Ourown

Ozark_Beauty

Beaversweet

Puget_Beauty

Gorella

OSC_2012

NC__1768

NC_1759

NC_1767

NC_1768

Surecrop

Lassen

Cal_103.22

Cal_468.4

Cal_588.1

Albritton

US_4152

WSU_685

Columbia

Cal_370.14

Cal_37.20-45

Cal_35.102-5

Cal_403.8

Shasta

S.65122

NY_844

OSC_2066

Donner

OSC_2122

((KlonmxMidl)x(TennShipxMidl))

Raritan

MDUS_2188

Bounty

K61-87

Macherauchs_Fruhernte

Regina

Belrubi

NY_E58

MDUS_2386

Midway

Ostara

Rabunda

Cal_491.1

MDUS_2321

Cal_470.9

Cal_490.9

NC_2665

(RockhillxCheyenne3)

Cal_37.2-11

Cal_638.45

Cal_544.3

MDUS_2787

Beaverbelle

Totem

Tamella

Induka

OSC_2242

Siletz

Vale

MDUS_3082

MDUS_3184

Lassenxself

Cal_51.51-1

OSC_2234

OSC_2315

Cal_168.1

Cal_168.3

Cal_39.96-18

NC_2967

Earlibelle

MDUS_2611

Sunrise

Rainier

Shuksan

Cal_752.23

Cal_752.31

Cal_46.5-1

Cal_55.32-1

Cal_460.11

Solana

Fort_Laramie

MDUS_2713

WSU_1532

OSC_2338

OSC_2261

OSC_2371

Marlate

Holiday

NJ_357

MDUS_2650

PocahontasXRegina

(MachFruherntexSengana)

NY_1221

Guardian

Redchief

Hapil

MDUS_2966

MDUS_2927

Hummi_GentoxOstara

MDUS_2992

MDUS_2905

Atlas

NC_2839

Cal_648.2

Cal_794.19

Cal_800.13

Apollo

MDUS_2856

Marmolada(Onebor)

Ogallala

Korona

OSC_2414

OSC_2931

Molalla

Cal_5649-1

OSC_2467

Cal_611.4

WSU_1985

Cal_42.8-16

Cheam

OSC_2981

Cal_54.124-2

Cal_63.7-101

MDUS_3364

OSC_2475

OSC_2769

Earliglow

Elsanta

Ark_5063

Delite

((MFruherxSSengana)xGeorgSolt)

Honeoye

Jewel

MDUS_4463

(GorellaxMDUS3184)

MDUS_4377

MDUS_4380

MDUS_4386

Governor_Simcoe

SJ85189

MDUS_3700

Hood

Etna

Lateglow

MDUS_3816

Cal_604.4

MDUS_3694

US_4419

Cal_42.17-18

ORUS_3306

ORUS_3316

ORUS_3320

NC_3140

MDUS_3413

SARGENTOCTOPLOID

Nanaimo

Linn

OSC_3727

Benton

Cal_829.9

Cal_39.117-4

OSC_4357

OSC_3624

OSC_3628

OSC_3551

OSC_3554

OSC_3577

Tioga

Cal_53.10-2

MDUS_4258

Olympus

OSC_2853

OSC_3044

Cal_59.51-11

Cardinal

Ark_5431

Macherauchs_Marieva

WIUS_8008

Joliette

Dana

Cal_66.96-101

OSC_3965

MDUS_4616

RedgauntletxKorona

K88-4

NYUS_119

Cal_814.6

NETHBIN

MDUS_4587

MSU_9-12

MSU_9-9

Allstar

MSU_9-5

MSU_9-8

Delmarvel

ORUS_850-4

ORUS_850-48

Cal_52.16-13

Cal_52.16-15

Wiltguard

OSC_4681

OSC_4682

Redcrest

WSU_1697

Bountiful

BC_41-14R-5

OSC_4445

Tufts

Cal_58.45-11

Cal_59.41-1

Cal_59.41-14

WSU_1574

OSC_3596

Tyee

WA_88097-47

OSC_3991

Aiko

OSC_3040

ORUS_1091-1

K68-58

ArKing

Micmac

BelrubixMacherauchs_Marieva

Scott

BC_91-14-31

FL_69-1302

Sumas

Harunoka

WSU_1651

((GorellaxMDUS3184)xTioga)

MNUS_52

OSC_4474

OSC_4816

OSC_4817

OSC_4916

Mara_des_Bois

Rapella

Evangeline

OSC_2732

ORUS_746-4

WSU_1945

BC_77-2-72

ORUS_984-49

Sequoia

Cal_65.65-601

Cal_71.98-604

Cal_71.98-605

WSU_1623

Redgem

Firecracker

OSC_4863

OSC_4275

Kent

K72-9

Glooscap

K74-5

(WiltguardxGorella)

Gariguette

WIUS_8002

OSC_4867

FL_71-189

Whonnock

E-30324

ORUS_973-1

Shuswap

ORUS_740-7

Idea

Winona

OSC_2975

Puget_Reliance

Cal_64.57-108

Cal_70.3-117

Cal_65.63-601

USEB_18

ORUS_869-13

ORUS_923-316

OSC_2950

OSC_6058

ORUS_1376-16

Pajaro

Brighton

K79-5

K79-4

Annapolis

(Redgauntx(WiltgxGorella))

Favour

MNUS_99

ORUS_986-129

Florida_Belle

L'Acadie

E-30397

Hecker

ORUS_1239R-21

OSC_3746

OSC_3845

WA_87010-7

Douglas

Muir

Selva

Cal_69.62-103

Tribute

ORUS_1267-236

ORUS_1267-250

NW_90005-1

OSC_3653

ORUS_1722-2

Cal_72.361-105

B144

Cal_77.3-603

K87-5

K86-19

Cavendish

Tiogax(Redgauntx(WiltgxGorel))

FavourxNY844

Mesabi

ORUS_1083-135

Aptos

Dover

Sweet_Charlie

ORUS_1769-1

Stolo

ORUS_2240-1

ORUS_2676-1

ORUS_3315

ORUS_3317

ORUS_3318

L'Authentique_Orleans

Nyohou

ORUS_750-1

OSC_4727

NW_90054-36

Irvine

Seascape

Fern

Parker

ORUS_1723-3

OSC_4319

ORUS_1735-1

Camarosa

Chandler

Cabot

Providence

MSU_49

WA_94023-1

NW_90055-35

MSU_9-15

MSU_9-16

MSU_9-7

ORUS_3308

ORUS_3326

ORUS_2369-1

ORUS_1947-2

Independence

ORUS_1076-124

ORUS_1069-3

Laguna

Anaheim

Cal_83.25-2

ORUS_767-4

ORUS_2180-1

EarliglowxChandler

Evita

Oso_Grande

MSU_56

(Providence_self)

Capitola

Darselect

MSU_9-13

MSU_9-2

MSU_9-6

ORUS_3307

ORUS_3323

ORUS_2262-2

ORUS_1754-1

ORUS_1963-5

ORUS_2159-2

MSU_9-1

MSU_9-10

MSU_9-3

Puget_Summer

NW_90049-19

Valley_Red

Cuesta

Pinnacle

ORUS_1009-1

ORUS_1027-1

CF1116

Everest

Strawberry_Festival

ORUS_1827-2

MSU_9-11

MSU_9-14

MSU_9-17

MSU_9-4

Florence

ORUS_2016-1

Tillamook

ORUS_2427-1

ORUS_2427-4

ORUS_1384-3

ORUS_1391-1

WSU_2833

ORUS_2490-1

ORUS_2742-1

ORUS_2781-1

ORUS_2493-2

ORUS_3304

ORUS_3305

ORUS_3321

ORUS_3324

ORUS_3325

ORUS_3277

ORUS_3278

ORUS_3279

ORUS_3314

ORUS_3319

ORUS_3322

Acknowledgements

This project is supported by the

Specialty Crops Research Initiative of USDA‟s

National Institute of Food and Agriculture

Questions?