Full NameAvian polyomavirusB-lymphotropic polyomavirusMW polyomavirusKI polyomavirusBaboon polyomavirusBat polyomavirusBK polyomavirusBornean orangutan polyomavirusBovine polyomavirusCalifornia sea lion polyomavirus 1Canary polyomavirusCrow polyomavirusChimpanzee polyomavirusFinch polyomavirusGoose hemorrhagic polyomavirusGorilla polyomavirusHamster polyomavirusHuman polyomavirus 6Human polyomavirus 7Human polyomavirus 9JC polyomavirusMerkel cell polyomavirusMurine pneumotropic polyomavirusMurine polyomavirusPan troglodytes verus polyomavirus 1aPan troglodytes verus polyomavirus 2aSimian virus 40Squirrel monkey polyomavirusSumatran orangutan polyomavirusTrichodysplasia spinulosa-associated polyomavirusMultimammate mouse polyomavirusWU polyomavirusBandicoot papillomatosis carcinomatosis virus type 1Bandicoot papilomatosis carcinomatosis virus type 2Japanese eel endothelial cells-infection virusCentropristis striata polyomavirus Meles Meles Polyomavirus 1 STL PolyomavirusBaboon Polyomavirus 2Rabbit kidney vacuolating virus

Athymic rat polyomavirusDolphin polyomavirusAfrican elephant polyomavirus 1Cynomolgus polyomavirusVervet monkey polyomavirus

Incomplete Genomic Sequence PolyomavirusesAthymic rat polyomavirusRabbit kidney vacuolating virusCynomolgus polyomavirusBaboon polyomavirus 2

Abbreviation Alternative Name APyV Budgerigar fledgling disease polyomaLPyV African green monkey polyomavirusMWPyVKIPyV KISA12 Simian agent 12BATPyV Myotis polyomavirus VM-2008BKPyV BK, BKVOraPyV1BPyVSLPyVCaPyVCPyVCHPyVFPyVGHPyVGggPyVHaPyVHPyV6HPyV7HpyV9JCPyV JC, JCVMCPyV MCVMPtV KilhamMPyVPtvPyV1aPtvPyV2aSV40SqPyVOraPyV2TSPyVMasPyV Mastomys polyomavirusWUPyVBPCV1BPCV2

BassPyV1 Black Sea Bass Polyomavirus 1MmelPyV1 European Badger PolyomavirusSTLPyVPpyVRKV

Rat-PyVDolPyVAelPyV-1CyPVVmPyV1

RatPyV Papovaviral sialoadenitisRKVCyPyVBPyV2

Genome Date DiscoveredNC_004764 1981NC_004763 1979NC_018102.1 2012NC_009238 2007NC_0007611 1963NC_011310 2009NC_001538 1971NC_013439 2010NC_001442 1977NC_013796 2010NC_017085.1 2010NC_007922 2006NC_014743 2005NC_007923 2006NC_004800 2000HQ385752.1NC_001663 1967NC_014406 2010NC_014407 2010NC_015150 2011NC_001699 1971NC_010277 2008NC_001505 1953NC_001515 1953HQ385746 2011HQ385948 2011NC_001669 1960NC_009951 2008FN356901.1 2010NC_014361 2010NC_025895.1 2011NC_009539 2007NC_010107 2007NC_010817 2008NC_015123 2011NC_025790 2015KP644239 2015 NC_020106 2013AB767295 1989AB435232 1966

1984 NC_025899 2013NC_022519 2013

1999NC_025898 2013

CitationBozeman, L. H., Davis, R. B., Gaudry, D., Lukert, P. D., Fletcher, O. J., & Dykstra, M. J. (1981). Characterization of a papovavirus isolated from fledgling budgerigars. Avian diseases, 972-980.zur Hausen H, Gissmann L (1979) Lymphotropic papovaviruses isolated from African green monkey and human cells. Med Microbiol Immunol 167: 137–153Siebrasse, Erica A., et al. "Identification of MW polyomavirus, a novel polyomavirus in human stool." Journal of virology 86.19 (2012): 10321-10326.Allander, Tobias, et al. "Identification of a third human polyomavirus." Journal of virology 81.8 (2007): 4130-4136.Malherbe, H. H., & Harwin, R. (1963). The cytopathic effects of vervet monkey viruses. South African medical journal= Suid-Afrikaanse tydskrif vir geneeskunde, 37, 407.Misra, Vikram, et al. "Detection of polyoma and corona viruses in bats of Canada." Journal of General Virology 90.8 (2009): 2015-2022.Gardner, S., Field, A., Coleman, D., & Hulme, B. (1971). New human papovavirus (BK) isolated from urine after renal transplantation. The Lancet, 297(7712), 1253-1257.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.Shah, K. V., Reissig, M., Daniel, R. W., & Bellhan, F. Z. (1977). Congenital transmission of a papovavirus of the stump-tailed macaque. Science, 195(4276), 404-406.Colegrove, Kathleen M., et al. "Polyomavirus infection in a free-ranging California sea lion (Zalophus californianus) with intestinal T-cell lymphoma." Journal of veterinary diagnostic investigation 22.4 (2010): 628-632.Halami, M. Y., Dorrestein, G. M., Couteel, P., Heckel, G., Müller, H., & Johne, R. (2010). Whole-genome characterization of a novel polyomavirus detected in fatally diseased canary birds. Journal of General Virology, 91(12), 3016-3022.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Johne, R., Enderlein, D., Nieper, H., & Müller, H. (2005). Novel polyomavirus detected in the feces of a chimpanzee by nested broad-spectrum PCR. Journal of virology, 79(6), 3883-3887.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Guerin, J. L., Gelfi, J., Dubois, L., Vuillaume, A., Boucraut-Baralon, C., & Pingret, J. L. (2000). A novel polyomavirus (goose hemorrhagic polyomavirus) is the agent of hemorrhagic nephritis enteritis of geese. Journal of virology, 74(10), 4523-4529.

Graffi, A., Schramm, T., Bender, E., Bierwolf, D., & Graffi, I. (1967). On a new virus containing skin tumor in golden hamster. Archiv für Geschwulstforschung, 30(4), 277.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Scuda, Nelly, et al. "A novel human polyomavirus closely related to the african green monkey-derived lymphotropic polyomavirus." Journal of virology 85.9 (2011): 4586-4590.Padgett, B., Zurhein, G., Walker, D., Eckroade, R., & Dessel, B. (1971). Cultivation of papova-like virus from human brain with progressive multifocal leucoencephalopathy. The Lancet, 297(7712), 1257-1260.Feng, Huichen, et al. "Clonal integration of a polyomavirus in human Merkel cell carcinoma." Science 319.5866 (2008): 1096-1100.Kilham, L., & Murphy, H. W. (1953). A pneumotropic virus isolated from C3H mice carrying the Bittner milk agent. Experimental Biology and Medicine, 82(1), 133-137.Gross, L. (1953). A filterable agent, recovered from Ak leukemic extracts, causing salivary gland carcinomas in C3H mice. Experimental Biology and Medicine, 83(2), 414-421.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Sweet, B. H., & Hilleman, M. R. (1960). The vacuolating virus, SV 40. Experimental Biology and Medicine, 105(2), 420-427.Verschoor, E. J., Groenewoud, M. J., Fagrouch, Z., Kewalapat, A., van Gessel, S., Kik, M. J., & Heeney, J. L. (2008). Molecular characterization of the first polyomavirus from a New World primate: squirrel monkey polyomavirus. Journal of General Virology, 89(1), 130-137.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.van der Meijden, Els, et al. "Discovery of a new human polyomavirus associated with trichodysplasia spinulosa in an immunocompromized patient." PLoS pathogens 6.7 (2010): e1001024.Orba, Yasuko, et al. "Detection and characterization of a novel polyomavirus in wild rodents." Journal of General Virology 92.4 (2011): 789-795.Gaynor, Anne M., et al. "Identification of a novel polyomavirus from patients with acute respiratory tract infections." PLoS pathogens 3.5 (2007): e64.Woolford, L., Rector, A., Van Ranst, M., Ducki, A., Bennett, M. D., Nicholls, P. K., ... & O'Hara, A. J. (2007). A novel virus detected in papillomas and carcinomas of the endangered western barred bandicoot (Perameles bougainville) exhibits genomic features of both the Papillomaviridae and Polyomaviridae. Journal of virology, 81(24), 13280-13290.Bennett, M. D., Woolford, L., Stevens, H., Van Ranst, M., Oldfield, T., Slaven, M., ... & Nicholls, P. K. (2008). Genomic characterization of a novel virus found in papillomatous lesions from a southern brown bandicoot (Isoodon obesulus) in Western Australia. Virology, 376(1), 173-182.Mizutani, T., Sayama, Y., Nakanishi, A., Ochiai, H., Sakai, K., Wakabayashi, K., ... & Ono, S. I. (2011). Novel DNA virus isolated from samples showing endothelial cell necrosis in the Japanese eel, Anguilla japonica. Virology, 412(1), 179-187.Peretti A, FitzGerald PC, Bliskovsky V, Pastrana DV, Buck CB. 2015. Genome sequence of a fish-associated polyomavirus, black sea bass (Centropristis striata) polyomavirus 1. Genome Announc 3(1):e01476-14. doi:10.1128/genomeA.01476-14.Hill, S., Murphy, A., Cotten, M., Palser, A., Benson, P., Lesellier, S., ... & Pybus, O. (2015). Discovery of a polyomavirus in European badgers (Meles meles) and the evolution of host range in the Polyomaviridae. Journal of General Virology, vir-0.Lim, E. S., Reyes, A., Antonio, M., Saha, D., Ikumapayi, U. N., Adeyemi, M., ... & Wang, D. (2013). Discovery of STL polyomavirus, a polyomavirus of ancestral recombinant origin that encodes a unique T antigen by alternative splicing. Virology, 436(2), 295-303.Gardner, S. D., Knowles, W. A., Hand, J. F., & Porter, A. A. (1989). Characterization of a new polyomavirus (Polyomavirus papionis-2) isolated from baboon kidney cell cultures. Archives of virology, 105(3-4), 223-233.Ito, Y., Hsia, S., & Evans, C. A. (1966). Rabbit kidney vacuolating virus: extraction of infectious DNA. Virology, 29(1), 26-31.

Ward, J. M., Lock, A., Collins, M. J., Gonda, M. A., & Reynolds, C. W. (1984). Papovaviral sialoadenitis in athymic nude rats. Laboratory animals, 18(1), 84-89.Anthony, S. J., Leger, J. A. S., Navarrete-Macias, I., Nilson, E., Sanchez-Leon, M., Liang, E., ... & Lipkin, W. I. (2013). Identification of a novel cetacean polyomavirus from a common dolphin (Delphinus delphis) with Tracheobronchitis. PloS one, 8(7), e68239.Stevens, H., Bertelsen, M. F., Sijmons, S., Van Ranst, M., & Maes, P. (2013). Characterization of a Novel Polyomavirus Isolated from a Fibroma on the Trunk of an African Elephant (Loxodonta africana). PloS one, 8(10), e77884.van Gorder, M. A., Della Pelle, P., Henson, J. W., Sachs, D. H., Cosimi, A. B., & Colvin, R. B. (1999). Cynomolgus polyoma virus infection: a new member of the polyoma virus family causes interstitial nephritis, ureteritis, and enteritis in immunosuppressed cynomolgus monkeys. The American journal of pathology, 154(4), 1273-1284.Yamaguchi, H., Kobayashi, S., Ishii, A., Ogawa, H., Nakamura, I., Moonga, L., ... & Orba, Y. (2013). Identification of a novel polyomavirus from vervet monkeys in Zambia. Journal of General Virology, 94(Pt 6), 1357-1364.

Bozeman, L. H., Davis, R. B., Gaudry, D., Lukert, P. D., Fletcher, O. J., & Dykstra, M. J. (1981). Characterization of a papovavirus isolated from fledgling budgerigars. Avian diseases, 972-980.zur Hausen H, Gissmann L (1979) Lymphotropic papovaviruses isolated from African green monkey and human cells. Med Microbiol Immunol 167: 137–153Siebrasse, Erica A., et al. "Identification of MW polyomavirus, a novel polyomavirus in human stool." Journal of virology 86.19 (2012): 10321-10326.Allander, Tobias, et al. "Identification of a third human polyomavirus." Journal of virology 81.8 (2007): 4130-4136.Malherbe, H. H., & Harwin, R. (1963). The cytopathic effects of vervet monkey viruses. South African medical journal= Suid-Afrikaanse tydskrif vir geneeskunde, 37, 407.Misra, Vikram, et al. "Detection of polyoma and corona viruses in bats of Canada." Journal of General Virology 90.8 (2009): 2015-2022.Gardner, S., Field, A., Coleman, D., & Hulme, B. (1971). New human papovavirus (BK) isolated from urine after renal transplantation. The Lancet, 297(7712), 1253-1257.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.Shah, K. V., Reissig, M., Daniel, R. W., & Bellhan, F. Z. (1977). Congenital transmission of a papovavirus of the stump-tailed macaque. Science, 195(4276), 404-406.Colegrove, Kathleen M., et al. "Polyomavirus infection in a free-ranging California sea lion (Zalophus californianus) with intestinal T-cell lymphoma." Journal of veterinary diagnostic investigation 22.4 (2010): 628-632.Halami, M. Y., Dorrestein, G. M., Couteel, P., Heckel, G., Müller, H., & Johne, R. (2010). Whole-genome characterization of a novel polyomavirus detected in fatally diseased canary birds. Journal of General Virology, 91(12), 3016-3022.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Johne, R., Enderlein, D., Nieper, H., & Müller, H. (2005). Novel polyomavirus detected in the feces of a chimpanzee by nested broad-spectrum PCR. Journal of virology, 79(6), 3883-3887.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Guerin, J. L., Gelfi, J., Dubois, L., Vuillaume, A., Boucraut-Baralon, C., & Pingret, J. L. (2000). A novel polyomavirus (goose hemorrhagic polyomavirus) is the agent of hemorrhagic nephritis enteritis of geese. Journal of virology, 74(10), 4523-4529.

Graffi, A., Schramm, T., Bender, E., Bierwolf, D., & Graffi, I. (1967). On a new virus containing skin tumor in golden hamster. Archiv für Geschwulstforschung, 30(4), 277.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Scuda, Nelly, et al. "A novel human polyomavirus closely related to the african green monkey-derived lymphotropic polyomavirus." Journal of virology 85.9 (2011): 4586-4590.Padgett, B., Zurhein, G., Walker, D., Eckroade, R., & Dessel, B. (1971). Cultivation of papova-like virus from human brain with progressive multifocal leucoencephalopathy. The Lancet, 297(7712), 1257-1260.Feng, Huichen, et al. "Clonal integration of a polyomavirus in human Merkel cell carcinoma." Science 319.5866 (2008): 1096-1100.Kilham, L., & Murphy, H. W. (1953). A pneumotropic virus isolated from C3H mice carrying the Bittner milk agent. Experimental Biology and Medicine, 82(1), 133-137.Gross, L. (1953). A filterable agent, recovered from Ak leukemic extracts, causing salivary gland carcinomas in C3H mice. Experimental Biology and Medicine, 83(2), 414-421.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Sweet, B. H., & Hilleman, M. R. (1960). The vacuolating virus, SV 40. Experimental Biology and Medicine, 105(2), 420-427.Verschoor, E. J., Groenewoud, M. J., Fagrouch, Z., Kewalapat, A., van Gessel, S., Kik, M. J., & Heeney, J. L. (2008). Molecular characterization of the first polyomavirus from a New World primate: squirrel monkey polyomavirus. Journal of General Virology, 89(1), 130-137.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.van der Meijden, Els, et al. "Discovery of a new human polyomavirus associated with trichodysplasia spinulosa in an immunocompromized patient." PLoS pathogens 6.7 (2010): e1001024.Orba, Yasuko, et al. "Detection and characterization of a novel polyomavirus in wild rodents." Journal of General Virology 92.4 (2011): 789-795.Gaynor, Anne M., et al. "Identification of a novel polyomavirus from patients with acute respiratory tract infections." PLoS pathogens 3.5 (2007): e64.Woolford, L., Rector, A., Van Ranst, M., Ducki, A., Bennett, M. D., Nicholls, P. K., ... & O'Hara, A. J. (2007). A novel virus detected in papillomas and carcinomas of the endangered western barred bandicoot (Perameles bougainville) exhibits genomic features of both the Papillomaviridae and Polyomaviridae. Journal of virology, 81(24), 13280-13290.Bennett, M. D., Woolford, L., Stevens, H., Van Ranst, M., Oldfield, T., Slaven, M., ... & Nicholls, P. K. (2008). Genomic characterization of a novel virus found in papillomatous lesions from a southern brown bandicoot (Isoodon obesulus) in Western Australia. Virology, 376(1), 173-182.Mizutani, T., Sayama, Y., Nakanishi, A., Ochiai, H., Sakai, K., Wakabayashi, K., ... & Ono, S. I. (2011). Novel DNA virus isolated from samples showing endothelial cell necrosis in the Japanese eel, Anguilla japonica. Virology, 412(1), 179-187.Peretti A, FitzGerald PC, Bliskovsky V, Pastrana DV, Buck CB. 2015. Genome sequence of a fish-associated polyomavirus, black sea bass (Centropristis striata) polyomavirus 1. Genome Announc 3(1):e01476-14. doi:10.1128/genomeA.01476-14.Hill, S., Murphy, A., Cotten, M., Palser, A., Benson, P., Lesellier, S., ... & Pybus, O. (2015). Discovery of a polyomavirus in European badgers (Meles meles) and the evolution of host range in the Polyomaviridae. Journal of General Virology, vir-0.Lim, E. S., Reyes, A., Antonio, M., Saha, D., Ikumapayi, U. N., Adeyemi, M., ... & Wang, D. (2013). Discovery of STL polyomavirus, a polyomavirus of ancestral recombinant origin that encodes a unique T antigen by alternative splicing. Virology, 436(2), 295-303.Gardner, S. D., Knowles, W. A., Hand, J. F., & Porter, A. A. (1989). Characterization of a new polyomavirus (Polyomavirus papionis-2) isolated from baboon kidney cell cultures. Archives of virology, 105(3-4), 223-233.Ito, Y., Hsia, S., & Evans, C. A. (1966). Rabbit kidney vacuolating virus: extraction of infectious DNA. Virology, 29(1), 26-31.

Ward, J. M., Lock, A., Collins, M. J., Gonda, M. A., & Reynolds, C. W. (1984). Papovaviral sialoadenitis in athymic nude rats. Laboratory animals, 18(1), 84-89.Anthony, S. J., Leger, J. A. S., Navarrete-Macias, I., Nilson, E., Sanchez-Leon, M., Liang, E., ... & Lipkin, W. I. (2013). Identification of a novel cetacean polyomavirus from a common dolphin (Delphinus delphis) with Tracheobronchitis. PloS one, 8(7), e68239.Stevens, H., Bertelsen, M. F., Sijmons, S., Van Ranst, M., & Maes, P. (2013). Characterization of a Novel Polyomavirus Isolated from a Fibroma on the Trunk of an African Elephant (Loxodonta africana). PloS one, 8(10), e77884.van Gorder, M. A., Della Pelle, P., Henson, J. W., Sachs, D. H., Cosimi, A. B., & Colvin, R. B. (1999). Cynomolgus polyoma virus infection: a new member of the polyoma virus family causes interstitial nephritis, ureteritis, and enteritis in immunosuppressed cynomolgus monkeys. The American journal of pathology, 154(4), 1273-1284.Yamaguchi, H., Kobayashi, S., Ishii, A., Ogawa, H., Nakamura, I., Moonga, L., ... & Orba, Y. (2013). Identification of a novel polyomavirus from vervet monkeys in Zambia. Journal of General Virology, 94(Pt 6), 1357-1364.

Bozeman, L. H., Davis, R. B., Gaudry, D., Lukert, P. D., Fletcher, O. J., & Dykstra, M. J. (1981). Characterization of a papovavirus isolated from fledgling budgerigars. Avian diseases, 972-980.

Malherbe, H. H., & Harwin, R. (1963). The cytopathic effects of vervet monkey viruses. South African medical journal= Suid-Afrikaanse tydskrif vir geneeskunde, 37, 407.

Gardner, S., Field, A., Coleman, D., & Hulme, B. (1971). New human papovavirus (BK) isolated from urine after renal transplantation. The Lancet, 297(7712), 1253-1257.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.

Colegrove, Kathleen M., et al. "Polyomavirus infection in a free-ranging California sea lion (Zalophus californianus) with intestinal T-cell lymphoma." Journal of veterinary diagnostic investigation 22.4 (2010): 628-632.Halami, M. Y., Dorrestein, G. M., Couteel, P., Heckel, G., Müller, H., & Johne, R. (2010). Whole-genome characterization of a novel polyomavirus detected in fatally diseased canary birds. Journal of General Virology, 91(12), 3016-3022.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Johne, R., Enderlein, D., Nieper, H., & Müller, H. (2005). Novel polyomavirus detected in the feces of a chimpanzee by nested broad-spectrum PCR. Journal of virology, 79(6), 3883-3887.Johne, R., Wittig, W., Fernández-de-Luco, D., Höfle, U., & Müller, H. (2006). Characterization of two novel polyomaviruses of birds by using multiply primed rolling-circle amplification of their genomes. Journal of virology, 80(7), 3523-3531.Guerin, J. L., Gelfi, J., Dubois, L., Vuillaume, A., Boucraut-Baralon, C., & Pingret, J. L. (2000). A novel polyomavirus (goose hemorrhagic polyomavirus) is the agent of hemorrhagic nephritis enteritis of geese. Journal of virology, 74(10), 4523-4529.

Graffi, A., Schramm, T., Bender, E., Bierwolf, D., & Graffi, I. (1967). On a new virus containing skin tumor in golden hamster. Archiv für Geschwulstforschung, 30(4), 277.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Schowalter, Rachel M., et al. "Merkel cell polyomavirus and two previously unknown polyomaviruses are chronically shed from human skin." Cell host & microbe 7.6 (2010): 509-515.Scuda, Nelly, et al. "A novel human polyomavirus closely related to the african green monkey-derived lymphotropic polyomavirus." Journal of virology 85.9 (2011): 4586-4590.Padgett, B., Zurhein, G., Walker, D., Eckroade, R., & Dessel, B. (1971). Cultivation of papova-like virus from human brain with progressive multifocal leucoencephalopathy. The Lancet, 297(7712), 1257-1260.

Gross, L. (1953). A filterable agent, recovered from Ak leukemic extracts, causing salivary gland carcinomas in C3H mice. Experimental Biology and Medicine, 83(2), 414-421.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.

Verschoor, E. J., Groenewoud, M. J., Fagrouch, Z., Kewalapat, A., van Gessel, S., Kik, M. J., & Heeney, J. L. (2008). Molecular characterization of the first polyomavirus from a New World primate: squirrel monkey polyomavirus. Journal of General Virology, 89(1), 130-137.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.van der Meijden, Els, et al. "Discovery of a new human polyomavirus associated with trichodysplasia spinulosa in an immunocompromized patient." PLoS pathogens 6.7 (2010): e1001024.

Woolford, L., Rector, A., Van Ranst, M., Ducki, A., Bennett, M. D., Nicholls, P. K., ... & O'Hara, A. J. (2007). A novel virus detected in papillomas and carcinomas of the endangered western barred bandicoot (Perameles bougainville) exhibits genomic features of both the Papillomaviridae and Polyomaviridae. Journal of virology, 81(24), 13280-13290.Bennett, M. D., Woolford, L., Stevens, H., Van Ranst, M., Oldfield, T., Slaven, M., ... & Nicholls, P. K. (2008). Genomic characterization of a novel virus found in papillomatous lesions from a southern brown bandicoot (Isoodon obesulus) in Western Australia. Virology, 376(1), 173-182.Mizutani, T., Sayama, Y., Nakanishi, A., Ochiai, H., Sakai, K., Wakabayashi, K., ... & Ono, S. I. (2011). Novel DNA virus isolated from samples showing endothelial cell necrosis in the Japanese eel, Anguilla japonica. Virology, 412(1), 179-187.Peretti A, FitzGerald PC, Bliskovsky V, Pastrana DV, Buck CB. 2015. Genome sequence of a fish-associated polyomavirus, black sea bass (Centropristis striata) polyomavirus 1. Genome Announc 3(1):e01476-14. doi:10.1128/genomeA.01476-14.Hill, S., Murphy, A., Cotten, M., Palser, A., Benson, P., Lesellier, S., ... & Pybus, O. (2015). Discovery of a polyomavirus in European badgers (Meles meles) and the evolution of host range in the Polyomaviridae. Journal of General Virology, vir-0.Lim, E. S., Reyes, A., Antonio, M., Saha, D., Ikumapayi, U. N., Adeyemi, M., ... & Wang, D. (2013). Discovery of STL polyomavirus, a polyomavirus of ancestral recombinant origin that encodes a unique T antigen by alternative splicing. Virology, 436(2), 295-303.Gardner, S. D., Knowles, W. A., Hand, J. F., & Porter, A. A. (1989). Characterization of a new polyomavirus (Polyomavirus papionis-2) isolated from baboon kidney cell cultures. Archives of virology, 105(3-4), 223-233.

Anthony, S. J., Leger, J. A. S., Navarrete-Macias, I., Nilson, E., Sanchez-Leon, M., Liang, E., ... & Lipkin, W. I. (2013). Identification of a novel cetacean polyomavirus from a common dolphin (Delphinus delphis) with Tracheobronchitis. PloS one, 8(7), e68239.Stevens, H., Bertelsen, M. F., Sijmons, S., Van Ranst, M., & Maes, P. (2013). Characterization of a Novel Polyomavirus Isolated from a Fibroma on the Trunk of an African Elephant (Loxodonta africana). PloS one, 8(10), e77884.van Gorder, M. A., Della Pelle, P., Henson, J. W., Sachs, D. H., Cosimi, A. B., & Colvin, R. B. (1999). Cynomolgus polyoma virus infection: a new member of the polyoma virus family causes interstitial nephritis, ureteritis, and enteritis in immunosuppressed cynomolgus monkeys. The American journal of pathology, 154(4), 1273-1284.Yamaguchi, H., Kobayashi, S., Ishii, A., Ogawa, H., Nakamura, I., Moonga, L., ... & Orba, Y. (2013). Identification of a novel polyomavirus from vervet monkeys in Zambia. Journal of General Virology, 94(Pt 6), 1357-1364.

Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.

Guerin, J. L., Gelfi, J., Dubois, L., Vuillaume, A., Boucraut-Baralon, C., & Pingret, J. L. (2000). A novel polyomavirus (goose hemorrhagic polyomavirus) is the agent of hemorrhagic nephritis enteritis of geese. Journal of virology, 74(10), 4523-4529.

Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.Leendertz, F. H., Scuda, N., Cameron, K. N., Kidega, T., Zuberbühler, K., Leendertz, S. A. J., ... & Ehlers, B. (2011). African great apes are naturally infected with polyomaviruses closely related to Merkel cell polyomavirus. Journal of virology, 85(2), 916-924.

Verschoor, E. J., Groenewoud, M. J., Fagrouch, Z., Kewalapat, A., van Gessel, S., Kik, M. J., & Heeney, J. L. (2008). Molecular characterization of the first polyomavirus from a New World primate: squirrel monkey polyomavirus. Journal of General Virology, 89(1), 130-137.Groenewoud, M. J., Fagrouch, Z., van Gessel, S., Niphuis, H., Bulavaite, A., Warren, K. S., ... & Verschoor, E. J. (2010). Characterization of novel polyomaviruses from Bornean and Sumatran orang-utans. Journal of General Virology, 91(3), 653-658.

Woolford, L., Rector, A., Van Ranst, M., Ducki, A., Bennett, M. D., Nicholls, P. K., ... & O'Hara, A. J. (2007). A novel virus detected in papillomas and carcinomas of the endangered western barred bandicoot (Perameles bougainville) exhibits genomic features of both the Papillomaviridae and Polyomaviridae. Journal of virology, 81(24), 13280-13290.Bennett, M. D., Woolford, L., Stevens, H., Van Ranst, M., Oldfield, T., Slaven, M., ... & Nicholls, P. K. (2008). Genomic characterization of a novel virus found in papillomatous lesions from a southern brown bandicoot (Isoodon obesulus) in Western Australia. Virology, 376(1), 173-182.

Peretti A, FitzGerald PC, Bliskovsky V, Pastrana DV, Buck CB. 2015. Genome sequence of a fish-associated polyomavirus, black sea bass (Centropristis striata) polyomavirus 1. Genome Announc 3(1):e01476-14. doi:10.1128/genomeA.01476-14.Hill, S., Murphy, A., Cotten, M., Palser, A., Benson, P., Lesellier, S., ... & Pybus, O. (2015). Discovery of a polyomavirus in European badgers (Meles meles) and the evolution of host range in the Polyomaviridae. Journal of General Virology, vir-0.Lim, E. S., Reyes, A., Antonio, M., Saha, D., Ikumapayi, U. N., Adeyemi, M., ... & Wang, D. (2013). Discovery of STL polyomavirus, a polyomavirus of ancestral recombinant origin that encodes a unique T antigen by alternative splicing. Virology, 436(2), 295-303.

Anthony, S. J., Leger, J. A. S., Navarrete-Macias, I., Nilson, E., Sanchez-Leon, M., Liang, E., ... & Lipkin, W. I. (2013). Identification of a novel cetacean polyomavirus from a common dolphin (Delphinus delphis) with Tracheobronchitis. PloS one, 8(7), e68239.

van Gorder, M. A., Della Pelle, P., Henson, J. W., Sachs, D. H., Cosimi, A. B., & Colvin, R. B. (1999). Cynomolgus polyoma virus infection: a new member of the polyoma virus family causes interstitial nephritis, ureteritis, and enteritis in immunosuppressed cynomolgus monkeys. The American journal of pathology, 154(4), 1273-1284.

Woolford, L., Rector, A., Van Ranst, M., Ducki, A., Bennett, M. D., Nicholls, P. K., ... & O'Hara, A. J. (2007). A novel virus detected in papillomas and carcinomas of the endangered western barred bandicoot (Perameles bougainville) exhibits genomic features of both the Papillomaviridae and Polyomaviridae. Journal of virology, 81(24), 13280-13290.

van Gorder, M. A., Della Pelle, P., Henson, J. W., Sachs, D. H., Cosimi, A. B., & Colvin, R. B. (1999). Cynomolgus polyoma virus infection: a new member of the polyoma virus family causes interstitial nephritis, ureteritis, and enteritis in immunosuppressed cynomolgus monkeys. The American journal of pathology, 154(4), 1273-1284.