Mealybugs & Mealybug Look-Alikes of the …...for some of major agronomic crops in the southeastern...

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Mealybugs & Mealybug Look-Alikes of the Southeastern United States

Transcript of Mealybugs & Mealybug Look-Alikes of the …...for some of major agronomic crops in the southeastern...

Page 1: Mealybugs & Mealybug Look-Alikes of the …...for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of

Mealybugs & Mealybug Look-Alikes of the Southeastern United States

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This publication was produced and distributed by USDA-CSREES Integrated Pest Management •Centers in cooperation with the National Plant Diagnostic Network, APHIS, the National Plant Board, and the Land Grant Universities. USDA-CSREES Critical Needs grant 2005-1763-02 ‘Early Detec-tion and Rapid Response System for the Pink Hibiscus Mealybug through Education, Training, and Implementation of Integrated Pest Management’, by J. VanKirk, G. Wisler, R. Hammerschmidt, and S. Ratcliffe, funded graphic design and printing for this publication. Content development was funded through the USDA-CSREES Southern Plant Diagnostic Network, the University of Florida, Institute of Food & Agricultural Sciences, and the Florida Department of Agriculture & Consumer Services, Division of Plant Industry.

Publication Information

For further information regarding the development of this document, contact Amanda Hodges at •[email protected](352)392-1901ext.199

Susan Ratcliffe• , University of Illinois, NCIPMC, Project Coordinator

Scott Martin• , University of Illinois, NCIPMC, Graphic Design

321INCHES5 64321MM

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Amanda Hodges• , SPDN, University of Florida, Institute of Food & Agricultural Sciences, Ento-mology & Nematology Department

Gregory Hodges• , Florida Department of Agriculture & Consumer Services, Division of Plant Industry

Lyle Buss• , University of Florida, Institute of Food & Agricultural Sciences, Entomology & Nema-tology Department

Lance Osborne• , University of Florida, Institute of Food & Agricultural Sciences, Mid-Florida Research & Education Center

Authors

Photo CreditsLyle Buss, University of Florida, Institute of Food & Agricultural Sciences, Entomology & •Nematology Department - 001c,001d,005,008,009,010,011,012,013,014a,014c,015,016,017,019a,019b,020a,021a,022a,022b,023b,026,027,028,029a,029b,030,031,032,034,036,037,038a,039,CoverPhoto

Florida Department of Agriculture & Consumer Services, Division of Plant Industry •001a,002,004,007,014b,018,020b,021b,023a,024,025,033,035,040

Lance Osborne, University of Florida, Institute of Food & Agricultural Sciences, Mid-Florida •Research & Education Center - 001b

Catharine Mannion, University of Florida, Institute of Food & Agricultural Sciences, Tropical •Research & Education Center - 003, 006

Mike Merchant, Texas A&M University, Texas AgriLife Extension Service - • 038b

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Page 001Mealybugs (Family Pseudococcidae)

Page 022Cottony Cushion Scale & Related Species (Family Margarodidae)

Page 024Soft Scales (Family Coccidae)

Page 038Felt Scales (Family Eriococcidae)

Page 040Cochineal Scales (Family Dactylopiidae)

Mealybugs & Mealybug Look-Alikes of the Southeastern United States

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What are mealybugs?Mealybugsareplantpeststhatfeedbypiercing-suckingmouthparts.Theyareclassifiedin•thefamilyPseudococcidae.Mealybugs,alongwithsoftscales(Coccidae)andarmoredscales(Diaspididae),arethemostcommonfamiliesofscaleinsects(Hemiptera:Coccoidea).

Approximately11ofthe21speciesofexoticscalesandmealybugsthathaveestablishedinthe•United States during the past 20 years are currently considered problematic pests. Four of these speciesaremealybugs:pinkhibiscusmealybug,miscanthusmealybug,papayamealybug,andvinemealybug.

Mostexoticscalesandmealybugsofconcernfeedonawiderangeofplantsspecies,andtheyare•capableofrapidpopulationexplosions.

Awarenessofcommonfieldcharacteristicspromotesearlydetectionforpestsofconcern.•

Mealybugs of the Southeastern United States

Why are mealybugs important?

How do you identify mealybugs?Thecommonname“mealybug”isderivedfromthefinepowderyto“mealy”waxthatcoversthe•body.Adultfemalemealybugsarewinglessandthebestlifestageforidentification.Adultmalemealybugsarewinged,buttheyarerarelyseenexceptwithspecificchemical-basedtrappingsystems.Mostadultfemalemealybugsaresmall(afewmillimeterstotypicallylessthan1/2inch),and a hand lens is useful for scouting. The immature or active “crawler” stage of mealybugs is typically less than half a millimeter and may be readily dispersed to new plants via wind currents.

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Body shape, size, and color.•Thenumberofwaxfilamentsprotrudingfromthesideofthebody.•Presenceandlengthofwaxfilamentsattheendofthebody(i.e.,terminalwaxfilaments).•Colorofeggs(ifpresent).•Presenceofanovisac(awaxymasscoveringtheeggs).•Stripes on the body.•Coloroffluidswhencrushed.•

Thefollowingcharacteristicsareusefulforfieldidentification:

Becausemealybugsfeedonplantjuicestheyproducehoneydew.Honeydew(asugary,sticky•substance)promotesthegrowthofblacksootymold.Sootymoldmayinhibitplantphotosynthesis.Other signs of plant damage may include stunted plant growth, wilting, and crinkled leaves. Mealy-bugscommonlyfeedonnewplantgrowth.Thepinkhibiscusmealybugalsoinjectsatoxicsalivawhile feeding, and a characteristic known as “bunchy top” is often seen on its preferred host plant, hibiscus.

Plant damage

Chemical and biological control options may be available for some mealybug and scale insect •species.Ageneralwebsitefocusingonmealybugsisavailableathttp://mrec.ifas.ufl.edu/LSO/Mealybug.htm.Biologicalcontroloptionsoftenincludespecies-specificparasitoids,andchemi-cal recommendations may vary depending on pest species or local recommendations. Your local countyextensionserviceisthebestsourceforinformationregardingchemicalcontrolsinyourarea.

General mealybug integrated pest management

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Both biological and chemical control options are available for pink hibiscus mealybug. Once this •pest has established in the landscape, biological control options are most often recommended. The most common natural enemies of pink hibiscus mealybug include the mealybug destroyer, Cryptolaemus montrouzieeri, and the parasitic wasps, Anagyrus kamali and Gyranusoidea indica. Incontrast,chemicaloptionsmaybetheonlychoiceinacommercial/nurserysetting.

General mealybug integrated pest management

Pink hibiscus mealybug is considered a regulatory pest of concern; subsequently, there is a zero tol-•erance for shipment of infested plant material. Please refer to the Pink Hibiscus Mealybug website http://mrec.ifas.ufl.edu/lso/pinkmealybug.htmforthelatestup-to-dateinformationonpinkhibiscusmealybug management.

Mealybugsinthisdeckhavebeenclassifiedasthefollowing:•

Warning! This deck is not a comprehensive listing of all mealybugs. Although useful as a •fieldscreeningtool,fieldidentificationisnotdefinitivefornewcounty,host,state,orconfi-dentialrecords.Slidemountingofspecimensandidentificationbyaspecialistisnecessaryforspecies-levelconfirmation.

Warning

Exotic:NotnativetotheUnitedStates,buthasinvaded.1. Native:NativetotheUnitedStates,orduetoitswidespreaddistributionandlong-termestablish-2. ment in the United States, its origin is uncertain.

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001001E X O T I C

Maconellicoccus hirsutus

Pink Hibiscus Mealybug

Common Hosts

Known Southeastern Distribution

Field RecognitionBodycolorpink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopinkincolor.Ovisacsarepresentcoveringpinktoorange eggs. Feeding from pink hibiscus mealybug can cause twisted or distorted foilage. High populations may result in leaf drop.

Over 200 known hosts occur, but the most common host detected to-date is Hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern U.S. if estab-lished populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.

EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

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Pink Hibiscus MealybugMaconellicoccus hirsutus

Bunchy Top

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001001E X O T I C

Maconellicoccus hirsutus

Pink Hibiscus Mealybug

Common Hosts

Known Southeastern Distribution

Field RecognitionBodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.

More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.

EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

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Pink Hibiscus MealybugMaconellicoccus hirsutus

Droplet of Red Body Fluid

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001001E X O T I C

Maconellicoccus hirsutus

Pink Hibiscus Mealybug

Field Recognition

Known Southeastern Distribution

Common Hosts

Bodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.

More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.

EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

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Pink Hibiscus MealybugMaconellicoccus hirsutus

White Wax on Abdomen

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001001E X O T I C

Maconellicoccus hirsutus

Pink Hibiscus Mealybug

Field Recognition

Known Southeastern Distribution

Common Hosts

Bodypink,about3mmlong,notofewlateral(side)waxfilaments,bodyfluidredtopink.Ovisacsarepresentcoveringpinktoorangeeggs.Feeding from pink hibiscus mealybug can cause twisted or distorted foil-age. High populations may result in leaf drop.

More than 200 known hosts occur, but the most common host detected to date is hibiscus. Pink hibiscus mealybug could be a problematic pest for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of hibiscus, is of particular concern.

EstablishedinFlorida(2002)andlimitedpopulationsdetectedinLouisiana(2006)andTexas(2007).

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Pink Hibiscus MealybugMaconellicoccus hirsutus Eggs

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Common Hosts

Known Southeastern Distribution

Field Recognition

Jatropha, Hibiscus, and Plumeria.

Established in Florida.

Bodyyellow,2-3mmlong,withmanylateral(side)waxfilaments.Ovisacspresentwithyellowtogreenishyelloweggs.Waxpatternonbodylackinganystripesonitsuppersurface(i.e.,dorsum).

Papaya MealybugParacoccus marginatus

E X O T I C002002

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Papaya MealybugParacoccus marginatus

Yellow Immatures

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Field Recognition

Common Hosts

Known Southeastern Distribution

Jatropha, Hibiscus, and Plumeria.

Established in Florida.

Bodyyellow,2-3mmlong,withmanylateral(side)waxfilaments.Ovisacspresentwithyellowtogreenishyelloweggs.Waxpatternonbodylackinganystripesonitsuppersurface(i.e.,dorsum).

Papaya MealybugParacoccus marginatus

E X O T I C002002

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Papaya MealybugParacoccus marginatus

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Field Recognition

Common Hosts

Known Southeastern Distribution

N A T I V E003003

Wide host range, may vary in local areas but common on Croton, Coleus, and Hibiscus.

Considered cosmopolitan in greenhouses and interior landscapes throughout the southeastern United States. Outdoor populations estab-lished in Florida.

Bodygrayunderneaththewaxysurfaceandwithmanylateralwaxfila-ments. Body has two darker lengthwise strips on the body surface and size is about 3 mm long. Ovisacs present with yellow eggs.

Madeira MealybugPhenacoccus madeirensis

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Madeira MealybugPhenacoccus madeirensis Lengthwise Dark Stripes

Many Lateral Wax Filaments

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Common Hosts

Field Recognition

Known Southeastern Distribution

004004N A T I V E

Phenacoccus solenopsis

Solenopsis Mealybug

Bodygray,about4mmlong,withmanylateralwaxfilaments.Ovisacsnotpresent.Waxontopofbodysurfaceappearsasacottonymass,horizontaldarkstripesatthetipoftheendofthebody(i.e.,theabdomenoppositeofthefeedingmouthparts).

Florida and Mississippi.

Most common on Hibiscus and other malvaceous hosts.

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Solenopsis MealybugPhenacoccus solenopsis

Dark Stripes

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Common Hosts

Known Southeastern Distribution

Field Recognition

005005E X O T I C

Ferrisia virgata

Striped Mealybug

Widehostrange,commononcopperleaf(Acalypha), Alternanthera, silverbuttonwood(Conocarpus),andHibiscus.

Alabama, Florida, Georgia, Louisiana, Mississippi.

Bodygray,about4-5mmlong,withoutside(lateral)wasfilaments.Two“thick”waxfilamentsarisingfromtipofabdomen.Ovisacsnotpresent.Twodarklengthwisestripesontopsurfaceofbody.Thincrystal-likefila-ments protruding from sides and top of body.

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Striped MealybugFerrisia virgata Crystal-Like Filaments

Two Filaments Protruding from Abdomen

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Common Hosts

Known Southeastern Distribution

Field Recognition

006006

Widehostrange,commononCitrus.SPECIALNOTE:ThePacificmealybug(Planococcus minor)cannotbeseperatedinfieldfromcitrusmealybug.TherearenoknownpopulationsofPacificmealybugoccurringintheUnitedStates.ThePacificmealybugisalsoknownasthe passionvine mealybug.

Common on ornamental plants in interior landscapes and greenhouses. Does occur outdoors in Florida.

Body color light yellow to grayish yellow, about 3 mm long, with many side(lateral)filaments.Ovisacspresent,eggsyellow.Onecentralizeddarkstripeondorsum(topsurfaceofbody).

Planococcus citri

Citrus MealybugE X O T I C

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Citrus MealybugPlanococcus citri

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Common Hosts

Known Southeastern Distribution

Field Recognition

007007E X O T I C

Pseudococcus longispinus

Longtailed Mealybug

Body gray in adults, about 3 mm long, yellowish in immature stages, withmanylateralwaxfilaments.Fourlongfilamentsarisingfromtipof abdomen, middle pair being very long. One centralized dark stripe located in center of abdomen. No ovisac.

Common species found in interiorscapes and greenhouses. Does occur outdoors in Florida.

Wide host range, common on Sago palms in Florida.

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Longtailed MealybugPseudococcus longispinus

Lengthwise Dark Stripes

Four Anal Wax Filaments

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Common Hosts

Field Recognition

Known Southeastern Distribution

008008E X O T I C

Polyphagous species, most common in Florida on tropical fruit.

Not common, does occur outdoors in Florida.

Bodylightorangetopink,about3mmlong,withmanylateralfilaments.Twolongfilamentsarisingfromtipofabdomen.Bodyfluidsbrownish.Ovisac present, but not entirely covering body.

Pseudococcus jackbeardsleyi

Jack Beardsley Mealybug

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Jack Beardsley MealybugPseudococcus jackbeardsleyi

Two Anal Filaments

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Known Southeastern Distribution

Common Hosts

Field Recognition

009

Polyphagous species, common on cacti.

Not common, occurs in Florida.

Bodypinktolightpurple,about3mmlong,withmanyside(lateral)filaments.Verysimilar in appearance to Jack Beardsley mealybug, but body tends to be more pinkish.Bodyisoval.Twolonganalfilamentsarisingfromtheabdomen.Nostripe on top surface of body. Eggs light colored, gray to yellow. Ovisac present, but not entirely covering body.

Pseudococcus viburni

Obscure Mealybug009N A T I V E

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Obscure MealybugPseudococcus viburni

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Field Recognition

Known Southeastern Distribution

Common Hosts

010010E X O T I C

Literature reports that this mealybug can occur in cacti. However, in Florida the most common hosts are Portulaca and Alternathera species.

Established in Florida.

Bodyredtopink,about3mmlong,withoutlateralfilaments.Bodyfluidsreddish. Oval to round-shaped body. No ovisac, but produces large amountsofwax.Feedingtendstoclusteratnodalregionsofplants.

Hypogeococcus pungens

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Hypogeococcus pungens

Adult Female

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Common Hosts

Known Southeastern Distribution

Field Recognition

011

Polyphagous species, common in Florida on roots of many of the palms.

Florida and Louisiana.

Bodygray,about2-3mmlong,withmanylateralwaxfilaments.Nostripes on body. Occurs both above and below ground.

Dysmicoccus brevipes

Pineapple MealybugE X O T I C011

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Pineapple MealybugDysmicoccus brevipes

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Field Recognition

Known Southeastern Distribution

Common Hosts

012012N A T I V E

Oracella acuta

Acute Mealybug

Restrictedtothefollowingpines:loblolly,longleaf,shortleaf,andVirginiapines.

Florida, Georgia, Kentucky, Louisiana, Mississippi, North Carolina, SouthCarolina,Texas,andVirginia.

Bodyredtopink,about3mmlong,withoutside(lateral)waxfilaments.Generallyfoundbothunderneathbarkandonneedlesofhosts.

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Acute MealybugOracella acuta

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Field Recognition

Known Southeastern Distribution

Common Hosts

013013E X O T I C

Noxious Bamboo Mealybug

Restricted to bamboos.

Common species, occurring throughout Southeastern region where bamboo is grown.

Adultbodybrown,about2-3mmlong;immaturestages(i.e.,crawlers)yellow. Generally found at the nodal regions of various bamboos. Sooty moldoccurringatthenodalregionsandlongwaxfilamentsarising from nodal areas is a common symptom.

Antonina pretiosa

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Noxious Bamboo MealybugAntonina pretiosa

Long Wax Filaments

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Field Recognition

Known Southeastern Distribution

Common Hosts

014Nipaecoccus nipae

Polyphagous species, most common on palms.

Florida and Louisiana.

Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.

Coconut MealybugE X O T I C014

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Coconut MealybugNipaecoccus nipae

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Field Recognition

Known Southeastern Distribution

Common Hosts

014Nipaecoccus nipae

Polyphagous species, most common on palms.

Florida and Louisiana.

Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.

Coconut MealybugE X O T I C014

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Coconut MealybugNipaecoccus nipae

Male Puparia

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Field Recognition

Known Southeastern Distribution

Common Hosts

014Nipaecoccus nipae

Polyphagous species, most common on palms.

Florida and Louisiana.

Bodymaroontored,about3mmlong,withfewside(lateral)waxfila-ments.Dorsalwaxfilamentsoccurring.Bodyroundtooval.Noovisacproduced, males common.

Coconut MealybugE X O T I C014

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Coconut MealybugNipaecoccus nipae

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Field Recognition

Known Southeastern Distribution

Common Hosts

015015E X O T I C

Most commonly encountered on sugarcane but Andropogon and Miscanthus grasses also may be hosts.

Established in Florida.

Bodypinktored,about4mmlong,bodyelongatedandlackinglateralwaxfilaments.Usuallyfoundonstalksorbeneathsheathsonbladesofhost plants.

Pink Sugarcane MealybugSaccharicoccus sacchari

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Pink Sugarcane MealybugSaccharicoccus sacchari

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Field Recognition

Known Southeastern Distribution

Common Hosts

016016E X O T I C

Varies,butgenerallypolyphagous.SPECIALNOTE:Therearemanyspecies within both Rhizoecus and Ripersiella. All may be considered pestsandsomemayrequirequarantineactions.Expertidentificationisrequired.

Varies depending on species but many are common throughout the Southeastern region.

Verysmallmealybugs(1to2mminlength),bodywhitetoyellow-ishwhite,lackingside(lateral)waxfilaments.Rootswithinfestationsofgroundmealybugsgenerallyhaveareasofwhitewaxpresentandmealybugs may be visible with use of a hand lens.

Ground MealybugsRhizoecus & Ripersiella

N A T I V E&

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Ground MealybugsRhizoecus & Ripersiella

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E X O T I C Field Recognition

Known Southeastern Distribution

Common Hosts

017Vryburgia amaryllidis

Lily Bulb Mealybug017

Commonlyinterceptedonbulbplants(lilybulbmealybug)andjadeplants(short-leggedmealybug).

Smallmealybugs(2-3mmlong),redtopurple.Lackingside(lateral)waxfilaments.Twothickwaxfilamentsarisingfromtipofabdomen.

Not established in Southeastern region, but established in California.

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Lily Bulb MealybugVryburgia amaryllidis

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Field Recognition

Known Southeastern Distribution

Common Hosts

018

Commonlyinterceptedonbulbplants(lilybulbmealybug)andjadeplants(short-leggedmealybug).

Not established in Southeastern region, but established in Arizona and California.

Vryburgia brevicruris

Short-Legged Mealybug

Smallmealybugs(2-3mmlong),redtopurple.Lackingside(lateral)waxfilaments.Twothickwaxfilamentsarisingfromtipofabdomen.

018E X O T I C

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Short-Legged MealybugVryburgia brevicruris

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Field Recognition

Known Southeastern Distribution

Common Hosts

019019N A T I V E

Most common host is fakahatchee and muhlygrass.

Established in Florida.

Bodygraywithwhitewax,about2-3mmlong,somewaxseemstobefilamentous.Nolateralwaxfilamentsproduced.

Stemmatomerinx acircula

Page 57: Mealybugs & Mealybug Look-Alikes of the …...for some of major agronomic crops in the southeastern United States if established populations are nearby. Cotton, a close relative of

Stemmatomerinx acircula

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Field Recognition

Known Southeastern Distribution

Common Hosts

019019N A T I V E

Most common host is fakahatchee and muhlygrass.

Established in Florida.

Bodygraywithwhitewax,about2-3mmlong,somewaxseemstobefilamentous.Nolateralwaxfilamentsproduced.

Stemmatomerinx acircula

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Stemmatomerinx acircula

Ovisac

Body

Wax FilamentsCrawler

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Field Recognition

Known Southeastern Distribution

Common HostsBamboo.

Established in Florida.

Bodygrayishpink,about3mmlong,withlargeamountsofwhitewaxvisibleonhostplant.Bodywithfewside(lateral)waxfilaments.Noovisacproduced.SPECIALNOTE:Thisspeciesisfoundunderneath sheaths of bamboo. It can be a severe pest.

020020E X O T I C

Palmicultor lumpurensis

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Palmicultor lumpurensisAdult

Crawlers

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Field Recognition

Common Hosts

Known Southeastern Distribution

Bamboo.

Established in Florida.

Bodygrayishpink,about3mmlong,withlargeamountsofwhitewaxvisibleonhostplant.Bodywithfewside(lateral)waxfilaments.Noovisacproduced.SPECIALNOTE:Thisspeciesisfoundunderneath sheaths of bamboo. It can be a severe pest.

020020E X O T I C

Palmicultor lumpurensis

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Palmicultor lumpurensis

Infested Bamboo

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Field Recognition

Known Southeastern Distribution

Common Hosts

021021E X O T I C

Various palms.

Established in Florida.

Bodyreddishbrowntopink,about3mmlong,withside(lateral)waxfilaments.Noovisacproduced.

Palmicultor browni

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Palmicultor browni

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Field Recognition

Known Southeastern Distribution

Common Hosts

021021E X O T I C

Various palms.

Established in Florida.

Bodyreddishbrowntopink,about3mmlong,withside(lateral)waxfilaments.Noovisacproduced.

Palmicultor browni

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Palmicultor browni

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Known Southeastern Distribution

Common Hosts

022022E X O T I C

Icerya genistae

Superficiallyresemblesthecottonycushionscale.Bodylightorangetopink,legsblack.Bodymaybecoveredwithawhitewax.Longovisacproduced(3-5xbodylength)andhelderecttohorizontalfrombody.

Established in Florida.

Wide host range, commonly collected on legumes.

Field Recognition

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Icerya genistae

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Known Southeastern Distribution

Common Hosts

022022E X O T I C

Icerya genistae

Superficiallyresemblesthecottonycushionscale.Bodylightorangetopink,legsblack.Bodymaybecoveredwithawhitewax.Longovisacproduced(3-5xbodylength)andhelderecttohorizontalfrombody.

Established in Florida.

Wide host range, commonly collected on legumes.

Field Recognition

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Icerya genistae

Immatures

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023023E X O T I C Field Recognition

Known Southeastern Distribution

Common HostsPolyphagous species, common on Citrus and Pittosporum.

A common species. Distribution scattered throughout the Southeastern region at different periods.

Bodyisorangewithsomewhitewaxpresentontopsideofbody.Legsareblack.Producesagroovedovisacthatisaboutthesamesize(or2x)asthebodyandgenerallytapersdowntowardthehostsubstrate.

Cottony Cushion ScaleIcerya purchasi

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Cottony Cushion ScaleIcerya purchasi

Ladybugs Feeding on Cottony Cushion Scale

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023023E X O T I C Field Recognition

Known Southeastern Distribution

Common HostsPolyphagous species, common on Citrus and Pittosporum.

A common species. Distribution scattered throughout the Southeastern region at different periods.

Bodyisorangewithsomewhitewaxpresentontopsideofbody.Legsareblack.Producesagroovedovisacthatisaboutthesamesize(or2x)asthebodyandgenerallytapersdowntowardthehostsubstrate.

Cottony Cushion ScaleIcerya purchasi

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Cottony Cushion ScaleIcerya purchasi

Immatures

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Field Recognition

Known Southeastern Distribution

Common Hosts

024

Maples, hollies, and dogwoods.

Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.

Body light brown to yellowish brown with young adults, dark brown on olderfemales.Body2.5-4.5mmlong.Ovisacproducedandgenerally found on underside of leaves.

Pulvinaria acericola

Cottony Maple Leaf ScaleN A T I V E024

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Cottony Maple Leaf ScalePulvinaria acericola

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Field Recognition

Known Southeastern Distribution

Common Hosts

025Pulvinaria ericicola

Azalea, lowbush blueberry and rusty lyonia.

Alabama, Florida, and Virginia.

Bodyredtoyellowandcoveredinglassywax.Body1.5-3.5mmlong.Ovisacproducedand2-4xaslongasbody.

Cottony Azalea Scale025N A T I V E

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Cottony Azalea ScalePulvinaria ericicola

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Field Recognition

Known Southeastern Distribution

Common Hosts

026E X O T I C026 Pulvinaria psidii

Green Shield Scale

Wide host range, common on Pittosporum, Citrus, Ficus species, and mango.

Alabama, Florida, Georgia, and Mississippi.

Adultfemalelightgreen,body2.0-4.5mmlong,ovisacproducedandwithoutnoticeablegrooves,approximately3xlengthofbody.Bodycoloris usually green but may seem somewhat brown depending on the host plant.

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Green Shield ScalePulvinaria psidii

Ants Tending Scales for Honeydew

Ovisac

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Field Recognition

Known Southeastern Distribution

Common Hosts

027E X O T I C027 Pulvinaria urbicola

Urbicola Soft Scale

Body of adult female brown to green, body 1.5 - 3.0 mm long, ovisac producedandwithoutnoticeablegrooves,approximately1.5-3xbodylength. Occurs on foliage but also near root line.

Alabama,Florida,Louisiana,andTexas.

Common on seagrape, guava, and Alternanthera species.

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Urbicola Soft ScalePulvinaria urbicola

Grooved Ovisac

Brown Body

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Field Recognition

Known Southeastern Distribution

Common Hosts

028N A T I V E028

Neopulvinaria innumerabilis

Cottony Maple Scale

Largescale(3-6mmlong)withcolorpatterndifferingdependingonhost, often blending in with host bark. Ovisac produced, but grooves areusuallynotclearlyvisible.Ovisac2xbodylength.Generallyfoundon woody portions of host plant material.

Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, Tennessee, and Virginia.

Polyphagous, common on dogwoods, maples and pecans.

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Cottony Maple ScaleNeopulvinaria innumerabilis

Ovisac

Female Body

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Common Hosts

Known Southeastern Distribution

Field Recognition

029E X O T I C029 Philephedra tuberculosa

Philephedra Scale

Largescale(3-6mmlong).Youngfemalesaregreensimilartogreenshieldscale.Ovisacsarelong(3-5xbodylength)andusuallyhavea“wet” appearance.

FloridaandTexas.

Wide host range but common on Acalypha, Codiaeum, and Conocarpus species.

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Philephedra ScalePhilephedra tuberculosa

Ovisac

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Field Recognition

Known Southeastern Distribution

Common Hosts

029E X O T I C029 Philephedra tuberculosa

Philephedra Scale

Largescale(3-6mmlong).Youngfemalesaregreensimilartogreenshieldscale.Ovisacsarelong(3-5xbodylength)andusuallyhavea“wet” appearance.

FloridaandTexas.

Wide host range but common on Acalypha, Codiaeum, and Conocarpus species.

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Philephedra ScalePhilephedra tuberculosa

Green Female Body

Immature Scales

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Field Recognition

Known Southeastern Distribution

Common Hosts

030E X O T I C030 Protopulvinaria pyriformis

Pyriform Scale

Body varies from tan to green to brown. Body about 2.5 mm long, pyriform. Ovisac produced, small and located beneath the scale insect. Long triangular anal plates visible in middle of top surface of body. Found on foliage of hosts.

Alabama, Florida, Georgia, Louisiana, North Carolina, South Carolina, Texas,andVirginia.

Wide host range, commonly found on Pittosporum, Ficus species, Schefflera, and tropical fruit.

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Pyriform ScaleProtopulvinaria pyriformis

Ovisac

Triangular Anal Plates

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Common Hosts

Known Southeastern Distribution

Field Recognition

031E X O T I C031 Milviscutulus mangiferae

Mango Shield Scale

Very similar to that of pyriform scale, body yellowish green, body size 2 - 3.5 mm long, pyriform. Ovisac produced and located beneath adult female. Found on foliage of hosts.

FloridaandTexas.

Polyphagous species, commonly found on Citrus, Ficus, Gardenia, jasmine and tropical fruit.

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Mango Shield ScaleMilviscutulus mangiferae

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Common Hosts

Known Southeastern Distribution

Field Recognition

032E X O T I C032 Ceroplastes ceriferus

Indian Wax Scale

Bodyred,coveredinathickwhitewetwax,normallywithananteriorprojecting “horn.” Body size 3-12 mm long. Found on woody portions of host plants.

Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.

Wide host range, common on hollies, camellias, and magnolias.

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Indian Wax ScaleCeroplastes ceriferus

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Field Recognition

Known Southeastern Distribution

Common Hosts

033E X O T I C033 Ceroplastes rusci

Fig Wax Scale

Similar in appearance to barnacle scale. Body red, covered in a thick pinkishwhitewax.Waxcoveringseemsplatedfromlateralview.Body2-3 mm long. Occurring on foliage and woody portions of host plant material.

Florida.

Wide host range, most commonly collected on Ficus species.

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Fig Wax ScaleCeroplastes rusci

Plated Wax Cover

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Field Recognition

Known Southeastern Distribution

Common Hosts

034E X O T I C034 Ceroplastes cirripediformis

Barnacle Scale

Bodymaroontored,coveredinathickbuff-coloredwax.Withoutananterior projecting horn. Appearing “plated” from lateral view. Body 1-6 mm long. Immatures maroon. Found on woody portions of host.

Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina, SouthCarolina,andTexas.

Wide host range, common on Ficus species, Pittosporum, hollies and Citrus.

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Barnacle ScaleCeroplastes cirripediformis

Maroon Immature Stages

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Field Recognition

Known Southeastern Distribution

Common Hosts

035E X O T I C035 Ceroplastes dugesii

Duges Wax Scale

Bodyred,coveredinthickgrayishwhitetobuffcoloredwaxcover.Body3-8 mm long. Found on woody portions of host plants.

Florida.

Wide host range, common on Bursera and Piper species.

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Duges Wax ScaleCeroplastes dugesii

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Field Recognition

Known Southeastern Distribution

Common Hosts

036E X O T I C036 Ceroplastes floridensis

Florida Wax Scale

Bodyred,coveredinathickgrayishtopinkishwhitewax.Generallyflat.Body2-4mmlong.Immaturesoftenresemblesmallwhitestars.Foundon twigs and foliage of hosts.

Alabama, Florida, Georgia, Louisiana, Mississippi, North Carolina, SouthCarolina,Texas,andVirginia.

Wide host range, common on hollies, Schefflera, Pittosporum, cycads, Citrus, and tropical fruit.

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Florida Wax ScaleCeroplastes floridensis

Adult Female

Immatures

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Field Recognition

Known Southeastern Distribution

Common Hosts

037E X O T I C037 Ceroplastes rubens

Red Wax Scale

Bodyred,coveredinathickredtopinkwax.Body2mmlong.Waxpentagonal in dorsal view. Occurring on foliage of host plants.

Florida.

Wide host range, common on Citrus and other tropical fruit.

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Red Wax ScaleCeroplastes rubens

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038

Common Hosts

Known Southeastern Distribution

Field RecognitionN A T I V E038 Eriococcus azaleae

Azalea Bark Scale

Body dark red to purple. Eggs and immature crawler stages pink. Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).Generallyfoundinnodalareasor“forks”oftwigsandbranches.

Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.

Wide host range but most commonly collected on Azalea.

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Azalea Bark ScaleEriococcus azaleae

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038

Common Hosts

Known Southeastern Distribution

Field RecognitionN A T I V E038 Eriococcus azaleae

Azalea Bark Scale

Body dark red to purple in color. Eggs and immature crawler stages pink.Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).Generallyfoundinnodalareasor“forks”oftwigsand branches.

Alabama, Arkansas, Florida, Georgia, Louisiana, Mississippi, NorthCarolina,SouthCarolina,Tennessee,Texas,andVirginia.

Wide host range but most commonly collected on Azalea.

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Azalea Bark ScaleEriococcus azaleae

Adult Female

Pink Immatures

Adult Female

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039N A T I V E039 Eriococcus quercus

Oak Eriococcin

Field RecognitionBodyisdarkredtopurple.Bodyapproximately3-4mmlong,ovisacproducedandisashinywhite(almostsilvery).

Known Southeastern DistributionAlabama,Florida,Georgia,Louisiana,Mississippi,Texas,andVirginia.

Common HostsOaks.

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Oak EriococcinEriococcus quercus

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040Dactylopius confusus040

N A T I V E

California Cochineal Scale

Field RecognitionBodymaroontopurple,coveredwithsticky-stringywax.Bodyfluidsred. Found on pads of cacti.

Known Southeastern DistributionFlorida,Georgia,andTexas.

Common HostsOpuntia cacti.

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California Cochineal ScaleDactylopius confusus