Lecture Exam I Cambrian radiation / phylogeny = 2 papers Systematics intro Protista: 10 phyla...
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Lecture Exam I
• Cambrian radiation / phylogeny = 2 papers
• Systematics intro
• Protista: 10 phyla
• Animalia: Porifera, Cnidaria
• ~20 questions
• Short answer, essay, life cycle sketch
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Phylum Cnidaria•3 Classes:
– Anthozoa– Scyphozoa– Hydrozoa
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Characteristics:
• Diploblastic metazoan
• Ectoderm and endoderm separated by acellular mesoglea
• Nervous system: simple nerve net, neurons
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Development
• Planula larvae
• ciliated, motile, gastrula larva
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Radial symmetry
• Modified into biradial, quadradial, etc…
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• Saclike, partitioned, or branched w/ one opening • = mouth and anus
Gastrovascular cavity
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Alternating life history forms: polyp and medusa
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Polyp
• Radial symmetry
• Oral end = hypostome or manubrium in hydrozoans– flat pedal disc in anthozoans
• Tentacles surround mouth
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PolypGastrovascular cavity
• Circulation, digestion, distribution of Circulation, digestion, distribution of nutrientsnutrients
• Hydrozoans: coelenteron is single tubeHydrozoans: coelenteron is single tube
• Scyphozoans: four longitudinal mesenteriesScyphozoans: four longitudinal mesenteries
• Anthozoans: compartmentalized by Anthozoans: compartmentalized by mesenteriesmesenteries
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Polyp: support• Hydrostatic, water-filled coelenteron Hydrostatic, water-filled coelenteron
• Anthozoans - bits of sediment and shell Anthozoans - bits of sediment and shell fragments in column wall for supportfragments in column wall for support
• Hydrozoans - flexible horny perisac (periderm) of Hydrozoans - flexible horny perisac (periderm) of chitin from epidermischitin from epidermis
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Polyp: movement
• Retractor muscles: longitudinal fibers along mesenteries
• Circular muscles: sphincters– in tentacles and oral disc
• Most polyps sedentary or sessile
• Creep slowly w/ pedal disc musculature• http://www.youtube.com/watch?v=-UI531GMRTM
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Medusa• All cnidaria except Anthozoa
• Bell-, dish- or umbrella-shaped
• Exumbrella: convex upper (aboral) surface
• Subumbrella: concave lower (oral) surface– Mouth at center
Free-floating,
mouth down
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Medusa
•External surface: epidermis• Internal surface: gastrodermis
•Coelenteron central; extends to radial canals– Usually four radial canals, tentacles,
stomach divided by mesenteries into four gastric pouches
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• Stinging structures
•
Nematocysts
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Nematocysts: multiple types
• Generic nematocyst (all)– Double-walled capsule w/ toxic mixture of phenols +
proteins– Spines or barbs for penetration, anchor in victim
• Spirocyst (Anthozoa)– Spring-like mechanism– Adhesive tubules wrap around and stick to victim
• Ptychocyst (tube anemones)– Create capsule tubule
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Feeding and digestion
Tentacles capture prey, carry to mouth, ingest whole
Extracellular digestion in coelenteron– enzyme-producing cells
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Reproduction and DevelopmentHydrozoa
• Polyps = asexual budding
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Reproduction and DevelopmentHydrozoan
• Free-living hydromedusa
• Dioecious, release sperm or eggs
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Reproduction and DevelopmentScyphozoa
•Asexual reproduction: small polyp = scyphistoma
•Medusa from scyphistoma
• Immature medusa = ephyra
•Most species dioecious
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Reproduction and DevelopmentAnthozoa
•Exclusively polyps
•Asexual reproduction
•Fission
•Pedal laceration: pedal disc spreads, anemone moves, leaves small fragments– develop into adult
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Class Hydrozoa
Hydroids and hydromedusa
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Hydrozoa
Portuguese Man-of-war
Physalia
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Class AnthozoaAnemones, corals, sea pens
Exclusively marine
Octocorals
• Soft corals– Gastraxonacea– Helioporaceans– Sea pens and sea pansies– Protoalcyonaria– Stoloniferans– Telestaceans
Sea pen
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Class AnthozoaAnemones and true corals
• True sea anemones• True corals (stony corals)
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Class Anthozoa
• Black or thorny corals• Ceranthids or tube anemones
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Class Scyphozoa
• Jellyfish
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Box jelly
Chiropsalmus