Involvement of phytohormone and c ell wall …...M. Asahina et al-1 FH 0CQR QRK2 "%#(PS7,W ^* 0 E [...
Transcript of Involvement of phytohormone and c ell wall …...M. Asahina et al-1 FH 0CQR QRK2 "%#(PS7,W ^* 0 E [...
M. Asahina et al-1
FH�0CQR��QRK2��"%#( PS7,W�^*�
�
ƁƟİɅś ɚ#3=+5<+;3815+;8�)//;+<+5�HÒȄŚ ��
�
ɞŊÇĬĴ�ƵňĴȲ�¡�����¹�ĴǓ� K������ƎƃljĵȴĺʼnȘȳĈ��
�ɞǝƤĬĴ� ƷĐƶĤǣ� K�������� ȀĝljfVoʼnĭƳĈ���
�
Involvement of phytohormone and cell wall metabolism on the tissue-reunion of incised tissue of plant.
Masashi ASAHINA1, Weerasak PITAKSARINGKARN2 and Shinobu SATOH2
Keywords; Arabidopsis, cell division, incised stem, phytohormone, tissue-reunion 1; Department of Biosciences, Teikyo University. 1-1, Toyosatodai, Utsunomiya, Tochigi,
320-8551, JAPAN. 2; Faculty of Life and Environment Sciences, University of Tsukuba. 1-1-1, Tennoudai, Tsukuba,
Ibaraki, 305-8572, JAPAN. �
�
�������
� ƑƯnƖJkĪƶĤmŒɋ�ǮP_ĄW|ÁhɚamǁƷxƙǺ�ĩý[`ɚƶĤlȮŜ\g
N|ɘ$/3.�+8.�$9<<��əIamyOkƶĤȋėmɛfhM|ÞĻnɚɑxȅkjmƯƵǂºƷ
ƯǂȋėɚôķxŬUƃkjmÉƭǂȋėǜlyegōTȜY[}|IɕǜƑƯhnɚȧŋɚ¼
Żìɇ\cǧǹċĨSüƯǧǹmyOlçpŬNJ^|YinkNSɚŬUƃly|ÌÚÓimŬ
NJxíŶ[}cǩDzmÙŗɚɆ\rSŐš[}|ɃmŘDŽmǪĊmɃlnɚÕĪǂlɇ}cǩDz
ċĨSçŌŬNJ^|YiSNjz}gN|��)+65/;������%=9..+;.���!-�>66A������9667+88�+8.�
�69-57+88������%3/1/6�+8.�(/;,/5/������?+8�./;�%-299=�/=�+6������$3-2/;.<98�/=�+6������
)+81�+8.��9667+88�����IǿSȲìǂlíŶ[}|iɚíŶ[}cǩDznɚǧǹìȈ�çȽ\g
į�}cǩDz�ìý[`ɚámǩDzċĨ�ǀĊ[`|YihÚÓƙǺSĖŗ^|ɘ�6+3<27+8�/=�+6�
����$/3.�+8.�$9<<�����5/>-23�/=�+6���əIǩDzSçƷºǀĊ^|ɃlnɚǧǹĦmçƷ
mÌɚ¦³�°��®mŔƷǂŐšly|ǧǹȾmȩǫɚǯǟƇǩDzmŷƷiȩǪiNec��
���kǁƷƴșSȏļ[}|YiSĢď[}gQ{ɘ%+-2<�����9667+88����69-57+88�
���əɚamȬǕnCÞĻmȓNjɚDǧǹìȈmȔȜɚEǧǹmìýɚFǧǹȾmŬNJiLjÆÔƹ
mšǚɚGĥƞºìýmÜƜRzk|¼ȩmƷƵăŜRzk|iǶPz}|�ę ɞ�<+238+�/=�+6�
��ƁƟİ ���IƅǗhnɚY}thlŢJSȆegTc�·�����míŶǻǿ�ƹNc
ìIJƷƯĴǂȐƊRzŕz}cſȤmǪƌ�ÁŘlǨË\cNI�
�
�
�
植物科学最前線 6: 31 ( 2015)
BSJ- Revi ew 6: 31 ( 2015)
M. Asahina et al-2
�
���'�! � 0CTU�QRK2�
����>A:LVE�
� Y}thlŢJnɚÔƯƷƸlQNgĚħƿĻ�ĖȰ^|džǂhŬUƃSȆ�}gN|�²�
´x�¬��ƹNgíŶ[}cǸȟSǀĊ^|ȬǕ�ƷƵĴǂlȐƊ\ɘę��əɚIJȂRzÖǬ
[}|�§¶´¹SDŽńmǧǹìȈȽılřɍhM{ɚċžlǧǹŬNJlßV¨��¹mĊš�
×Ȫ^|Yi�ŽzRl\cItcɚľǟƧlyegÖǬ[}|©�ǦkjmƮƙáǦwɚǩDz
ǀĊlřɍkYiwǏ\cɘ�<+238+�/=�+6����������ƁƟݺÒȄ�� əI�
植物科学最前線 6: 32 ( 2015)
BSJ- Revi ew 6: 32 ( 2015)
M. Asahina et al-3
� \R\ɚ�²�´x�¬�hnɚǀĊȬǕ�ȯÏĴǂºìIJƷƯĴǂlȐƊ^|YinĘɈh
Meccvɚ�·�����mǻǿ�ƹN|ŷȍmĸɓǣ�ǎǚ\cɘ�<+238+�/=�+6��əI�
�·�����mǻǿhnɚíŶ[}|iɔǩDzmǧǹSǧǹìȈ�çȽ^|Sɘę��əɚYm
žlǿƷȂxǿɌ�íɁ^|iǧǹìȈSɀĻ[}|YiSìRecIYmƴșlĽ^|ƑƯ©
µ°¹mŒɋlfNgƒȑ\ciY~ɚ�»��¹ĪƷŨÀly{ĕĶíɁly|ɀĻSĖŗ^
|Yiɚ�»��¹ƔŝȡȥɀĻó&����=;339.9,/8B93-�+-3.��ŨÀ\cÚÓxɚƔŝȡȥƚ
űÓɘ!� ���əhǩDzǀĊSŧñ[}|YiSìRecItcɚ�»��¹æƷȹmƫķɚĂp
�»��¹Ȕľŝ¦·°»�»�$����'% ŐțȞůÓ�%+,+=383�/=�+6������ƹNcȐƊRzɚ
íŶLJŔmíŶȲ½Ýl�»��¹Sȃǘ\gN|YiSǏ[}cɘ�<+238+�/=�+6��əItcɚ
íŶêƵŔɚƑƯÓ�ƗlÛ\ɚíŶȲÑ�¾ÝlÑdz\gƐğ\cģĊlnǀĊnƷ]|Sɚ
Þą�½Ýl\gƑƯÓ�Ɛğ^|iíŶȲmɌǼÝlQW|�$����'% Qyp �A-����'%ɘǧǹ
ìȈm¬»�»əmǁƴmƪĿiålɚɔmǧǹìȈSŧñ[}cɘę�əI�
� �
� νmǪƌRzɚ�·�����íŶǻǿmǩDzǀĊlnɚ�»��¹mƔŝȡȥiȃǘSř
ɍhM|YiSîŽ\cI�
�
����QRK2�^*��X.49�3;�
� �·�����ǻǿmǩDzǀĊȬǕlQW|ȯÏIJǁƴmĩý�ŽzRl^|cvɚ¬��·
�¶�ƣ�ƹNcȐƊ�ȆkecI�·�����ǻǿmǛ¼ǠȾtcnǛÅǠȾmǻǿl¬�
�·��¥�ƹNgíŶêƵ�Ź\ɚ¼ķžȾŔlíŶȲ�ĎuǤ��77 mǻǿ�í{ë\g$"�
植物科学最前線 6: 33 ( 2015)
BSJ- Revi ew 6: 33 ( 2015)
M. Asahina et al-4
�ũë\ɚȐƊlƹNcI�
� ¬��·�¶�ȐƊlyegċķ\cȯÏIJǵmÁRzɚǀĊȬǕòÿƂhǁƴ^|ȞèėIJ
lƥdž\gȐƊ�ȆecǪƌɚ"��ĜȞèėIJ¥�´»lŃ^| � ���ɚQyp�$���#� ĜȞ
èėIJ¥�´»lŃ^| � ����SɚíŶźŔRz�źŔl�cegɚǀĊȲưƾǂkǁƴ
�Ǐ\gN|YiSŽzRikecɘ�<+238+�/=�+6��əI�
� [zlH�$�%�& ƣ��3;+=<>�/=�+6�����ƹNgY}zmƙǺ�ŧñ\cŐțȞůÓxƚűĩ
ƾÓ � ����hnǩDzǀĊSɀĻ[}gNcYiRzɚY}zmȞèėIJnǩDzǀĊlřɍkƙ
Ǻ�ŪegN|YiSǏ[}cIƛlɚíŶɊm½¾mǩDz�a}b}ìö\gȯÏIJǁƴ�ȕ
rciY~ɚ � ���níŶɊm½Ýhɚ� ����níŶɊm¾ÝhưƾǂlǁƴS½ż^|Yi
SîŽ\cɘę əIY}zȞèñŖėIJmǁƴlĽ^|�»��¹mŒɋ�ȕrciY~ɚ
� ���mǁƴn�»��¹lyegƝlɚ� ����mǁƴnȚlɚa}b}ñŖ[}gN|Yi
SŽzRikecIY}zmǪƌRzɚÞm½Ȳhn�»��¹Sȃǘ^|Yilyeg � ���
SȔľ[}ɚÞm¾Ȳhn�»��¹SƍƩ^|Yilyeg � ����SȔľ[}|YiSŽz
Rikecɘ�<+238+�/=�+6��ɞƁƟİ��əI�
� �
�����$�#(]_��&(�^* �)��(@6-Z�
� íŶɛźŔm�·�����íŶǻǿmǀĊȲlQNgɚ��¶¹Ċšlȿ�|�� ��
·¦·¢¹�µª¹ȶĊšȵǦ�� ����ɚQyp�±�°¹ȶĊšlȿ�|´«����»��
�����Qyp �����SŎVǁƴ\gNcYiRzɚ��¶¹ɚ�±�°¹ȶmȿÀlfNgwƒȑ
\cI�
植物科学最前線 6: 34 ( 2015)
BSJ- Revi ew 6: 34 ( 2015)
M. Asahina et al-5
�
�
植物科学最前線 6: 35 ( 2015)
BSJ- Revi ew 6: 35 ( 2015)
M. Asahina et al-6
�
t_ɚ��¶¹ɉşĄŝĩƾÓ �� ��ƹNgċƖmíŶêƵ�ȆeciY~ɚíŶȲÑmǧǹì
ȈºÐȼlĩýSƷ]ɚƝŋkǩDzǀĊSȜYzkRecItcɚ�� �hnǻǿíŶȲlQW|
� ���mǁƴSŧñ[}gNcYiRzɚ � ���n��¶¹lyegƝlñŖ[}gN|Yi
SŽzRikecI�
� ƛlɚ�±�°¹ȶ¯�µ�ǻǿlŨÀ^|iɚ� ����mǁƴSȔľ[}|YiSìRecI
[zlɚ�±�°¹ȶȿȩĩƾÓhnƝŋkǩDzǀĊSƷ]kNYiɚǻǿíŶŔl�±�°¹
ȶmæƷȹSĥù\gNcYikjRzɚǀĊȲlQW| � ����mǁƴn�±�°¹ȶlye
gwȕǠ[}gN|YiSǏ[}c��<+238+�/=�+6��ɚƁƟİ���I�
� ¼Ÿɚ�»��¹ŜǞėIJhM| �"��"��ÅȷĩƾÓmíŶǻǿhnɚǩDzǀĊSŎVɀĻ[
}|YiSìRecɘę�əItcɚ�"��"��ÅȷĩƾÓ�ƹNcǁƴȐƊmǪƌɚ � ����i
� ����míŶǻǿhmǁƴSŧñ[}|YiɚɉíŶǻǿlQW|$�#��� mǁƴnɚȸƷĜ
iƟȠ\g½ż\gN|YiSìRecɘę����əI[zlɚ�"��"��mǀĊȲhnɚ�±�°
¹ȶƷĊšȯÏIJm¼ǖhM| � ��mǁƴSɚŎVŧñ[}gN|YiSìRecɘę��əIÎ
½mǪƌy{ɚ�·�����íŶǻǿmǩDzǀĊlQNgɚ�±�°¹ȶnȷȋkßT�ƀ\
gN|Yiɚ�»��¹ly|�"���º$�#��� ȞèėIJi�±�°¹ȶƷĊšȯÏIJmǁƴȕ
Ǡlɚ ��i ���SȷȋkėIJi\gßNgN|YiSǏ[}cɘ#3=+5<+;3815+;8�/=�+6�� +əI�
� νmǪƌɚ�·�����íŶǻǿmǩDzǀĊȬǕhnɚǻǿíŶlyegƷ]c�»��
¹ɚ��¶¹ɚ�±�°¹ȶiNecƑƯ©µ°¹m���´¹�Sɚ � ���%� ����Ȟèė
IJmǁƴ�Ë\gɚǧǹìȈǜmȯÏIJǁƴ�ñŖ\gN|ćǺŝSǏ[}cɘę�əI�
植物科学最前線 6: 36 ( 2015)
BSJ- Revi ew 6: 36 ( 2015)
M. Asahina et al-7
�����PS78N�,W �(�5��'�%�(XM\O�1G/V\O[-9�^*�
� ǩDzǀĊžlnǧǹìȈxǧǹŐŠm÷ǂkĩýSƷ]|YiRzɚǧǹĦīǢmĩý�ȐƊ
\ciY~ɚ̈ ��¹ŝmìņ\c�³£�¹Sĥù\gN|YiSîŽ\cɘ#3=+5<+;3815+;8�
/=�+6�� ,əI¼ŸH¬��·�¶�ƣly{íŶŔL�źdžlǁƴm¤»��ȣP|ǧǹĦȿ
ȩȯÏIJmȐƊ�ȆeciY~ɚċķ[}cȯÏIJǵmÁln¨��¹kjmǧǹĦīǢ�ìȐ
^|ȵǦmȯÏIJSīŴĎt}gNcSɚǧǹĦīǢmĊšlȿ�|ȯÏIJnsi�jĎt}g
NkRecItcɚǧǹĦ¬�´���īǢhM|��·�µ�¹i�µ·»�mȾmƠǦǪĊ
�íŶ^|���¢¹�¹��$!� #� ����i��·�µ�¹ȻȾmfkUůP�ȆO�¹�Ĝ�
�·�µ�¹ȞǔȵǦ�ùƠìȐȵǦ�*A6916>-+8�/8.9=;+8<16>-9<A6+</�2A.;96+</��������nɚ
ĉJɚíŶŔɛźdžiɜźdžlŎVǁƴ\gQ{ɚǧǹĦ���¸»�mÙɒiçƕǡSǀĊȬ
ǕlȿÀ\gN|ćǺŝSǏĒ[}cI�
aYhɚ�����mǀĊȬǕlQW|ǁƴ�ȒǧlȕrciY~ɚ�����mǁƴS�»��¹ly
egƝlñŖ[}gQ{ɚ � ��������ŐțȞůÓĂp � ����ƚűÓhn �����ȯÏIJmǁƴ
Sŧñ[}|Yiɚ��%�� � ���ȬõǁƴÓhnȦlǁƴS½ż\gN|YiSìRec
ɘ#3=+5<+;3815+;8�/=�+6�� ,əIƛlɚ�����iLjċŝmɕN �����imÅȷƚűĩƾÓmí
ŶǻǿlQW|ǀĊȬǕ�ȕrciY~ɚ��������ÅȷĩƾÓhnǩDzǀĊȬǕhmǧǹìȈS
ŎVɀĻ[}gNcYiRzɚ��������nǩDzǀĊȬǕlQW|ɔǩDzmǧǹìȈlȿÀ\gN
|YiSǏ[}cɘę�əI�
tcɚ�µ�¥�������6/-=;9:29;/<3<�!9,363=A�%230=��<<+A�lQNg �"��� �¹¢
�S ��������m¦·°»�»ɎĞlǪĊ^|Yiɘę�əɚQyp�·�����mȂlQNg�
植物科学最前線 6: 37 ( 2015)
BSJ- Revi ew 6: 37 ( 2015)
M. Asahina et al-8
�
��·¡��´�®�Ë\g � ����¼Ȭǂlǁƴ[`|i ���������m¦·°»�»ƦŝS
ɕt|YiSǏ[}cɘęəIνmǪƌRzɚ�»��¹hȔľ[}c�"��� lyegLJ
ŬñŖ�ĄWgǁƴ\c*&���� S��·�µ�¹mçDZ�ȧ\gǀĊȬǕmǧǹìȈlȿÀ
\gN|YiSǏ[}cɘ#3=+5<+;3815+;8�/=�+6�� ,əI�
植物科学最前線 6: 38 ( 2015)
BSJ- Revi ew 6: 38 ( 2015)
M. Asahina et al-9
��+?�<D�
� νy{ɚ�·�����íŶǻǿmǩDzǀĊlřɍmȞèėIJiƑƯ©µ°¹QypǧǹĦ
ÍȖmßTSŽzRikecIǩDzǀĊmyOkƑƯmƀ^|çƷǺønŬUƃi\gwðƹ[
}ɚƌȁɐxƌƘmǽmƷƸlƮVgnkzkNȢƓŦȇikegN|ɘŀŐ��əI\R\ɚ
ŬUƃȎđŝmìIJ¯���®lȿ^|ȒǧnŽzRikegNkNIŐŠĴǂNjȌSȒ\Vŕ
z}gN|¼ŸɚƷƵĴǂɚìIJƷƯĴǂNjȌnƄd¿ȝ\gN|mSƴƱhM|ItcɚǩDz
ȾºÚÓȾmȎđŝly{ǀĊSŒɋ�ĄW|cvɚŬUƃlQNgnɚamȎđŝiƦNJŝSɚ
ĸƹŝmɊlQNgweiwĬTkēɏikegN|IƴěthlƕȨ½mLjȭɚǩDzìýmƚ
űɚÅƛÍȖƸƯmLjȭǜSǶPz}gN|SɚŬUƃȎđŝmìIJ¯���®lȿ^|Ȓǧn
ŽzRikegNkNIÊŔɚŬUƃxÞĻlQW|ƑƯǩDzƢǀmìIJǂ¯���®mȐŽ�
džū\gNTcNiǶPgN|I�
�
��WY�
� ƅǍǙ�ȫȆ^|lŏc{ɚƵýĴǍǙŤºǑȗûƢâƷɚƈþĬĴºŅąØƛȱâƷɚȊȗ
đőâƷɚƗŅɗÈâƷɚġƲĬĴºɕƃàâƷɚƸƓŦȇǰĊǍǙŤºãƻłɂâƷɚČĆŁ
ĬĴºnjɖƬơâƷHŊÇĬĴºƗƻijɄâƷɚŅƏÂđâƷlnɚǍǙƆŵmZŮÖHZūľ
�ɌTt\cIŘy{şȖƼ\½Xt^ItcɚǍǙ�ålȪvgV}t\cɚǝƤĬĴºƉŇ
ĔĮĀĨɘƴºŊÇĬĴəɚƈäÃčɚƨƠǴƺ[�HĂpƑƯƷƵĴǍǙĹmǃ[�ɚŊÇĬĴº
ƑƯƷƵĴǍǙĹmǃ[�lQǐƼ\½Xt^I�
� ƅǍǙm¼ȲnɚǓĴǍǙȉúȺºǾťǍǙ�ɘ��� ��əɚưķɎĞǍǙºƑƯ¯´��®i
ĕĶmǁƷ�ŲP|ŞĢǭŖǣ���� �ɚŷĴȇɎĞǍǙºƑƯǧǹĦmŞĢêƵ���®
ɘ� �əɚǒǚĬĴţƽǂǍǙĠDžŐšŲŰÄƓºƑƯ���´¤¹mƷƵƙǺmȐŽiam
Ŝƹɘ%� əɚŊÇĬĴƵňĴȲųǷǍǙŭȪưïȉúȺɘ�����ɚ�� ���əmŲŰ�ĄWg
ȆNt\cI�
�
=JBI�
ƁƟİɅśɘ��əƑƯíŶǩDzmǀĊlQW|ƑƯ©µ°¹QypȞèėIJmœöIƑƯmƷ
ȼȕǠ� �ɝ ����
Asahina M, Azuma K, Pitaksaringkarn W, Yamazaki T, Mitsuda N, Ohme-Takagi M, Yamaguchi S, Kamiya Y, Okada K, Nishimura T, Koshiba T, Yokota T, Kamada H and Satoh S (2011) Spatially selective hormonal control of RAP2.6L and ANAC071 transcription factors involved in tissue reunion in
Arabidopsis. Proc Natl Acad Sci USA: 108 (38) 16128-16132.
Asahina M, Gocho Y, Kamada H and Satoh S (2006) Involvement of inorganic elements in tissue-reunion in the hypocotyl cortex of Cucumis sativus. J Plant Res 119:337-342.
Asahina M, Iwai H, Kikuchi A, Yamaguchi S, Kamiya Y, Kamada H and Satoh S (2002) Gibberellin
produced in the cotyledon is required for cell division during tissue-reunion inthe cortex of cut cucumber and tomato hypocotyls. Plant Physiol 129: 201-210.
ƁƟİɅśºÒȄŚɘ� əíŶǩDzmǀĊi�§¶´¹mȿÀIƑƯmƷȼȕǠ���ɝ��� �
植物科学最前線 6: 39 ( 2015)
BSJ- Revi ew 6: 39 ( 2015)
M. Asahina et al-10
Asahina M, Yamauchi Y, Hanada A, Kamiya Y, Kamada H, Satoh S and Yamaguchi S (2007) Effects
of the removal of cotyledons on endogenous gibberellin levels in hypocotyls of young cucumber and
tomato seedlings. Plant Biotechnol 24:99-106. Flaishman MA, Loginovsky K and Lev-Yadun S (2003) Regenerative xylem in inflorescence stems of
Arabidopsis thaliana. J Plant Growth Regul 22:253-258.
Hiratsu K, Matsui K, Koyama T and Ohme-Takagi M (2003) Dominant repression of target genes by chimeric repressors that include the EAR motif, a repression domain, in Arabidopsis. Plant J 34:733-739.
Ikeuchi M, Sugimoto K, Iwase A (2013). Plant Callus: Mechanisms of Induction and Repression. Plant
Cell. 25(9): 3159–3173. Kollmann R and Glockmann C (1985) Studies on graft unions.I. Plasmodesmata between cells of plants
belonging to different unrelated taxa. Protoplasma 124: 224-235.
Mattsson J, Sung ZR and Berleth T (1999) Responses of plant vascular systems to auxin transport inhibition. Development 126: 2979-2991.
ŀŐéƷɘ��əŬUƃmƷƵIƑƯmƷȼȕǠ� ɝ����� �
Pitaksaringkarn W, Ishiguro S, Asahina M, Satoh S (2014a). ARF6 and ARF8 contribute to tissue reunion in incised Arabidopsis inflorescence stems. Plant Biotechnol. 31, 49–53.
Pitaksaringkarn W, Matsuoka K, Asahina M, Miura K, Sage-Ono K, Ono M, Yokoyama R, Nishitani K, Ishii T, Iwai H, Satoh S (2014b). XTH20 and XTH19 regulated by ANAC071 under auxin flow are involved in cell proliferation in incised Arabidopsis inflorescence stems. Plant J. 80, 604–614.
Reid JB and Ross JJ (2011) Regulation of tissue repair in plants. Proc Natl Acad Sci USA: 108 (42)
17241-17242. Sabatini S, Beis D, Wolkenfelt H, Murfett J, Guilfoyle T, Malamy J, Benfey P, Leyser O, Bechtold N,
Weisbeek P and Scheres B (1999) An Auxin-Dependent Distal Organizer of Pattern and Polarity in the
Arabidopsis Root. Cell 99: 463-472. Sachs T (2000) Integrating cellular and organismic aspects of vascular differentiation. Plant Cell Physiol
41: 649-656.
Siegel BA and Verbeke JA (1989) Diffusible factors essential for epidermal cell redifferentiation in Catharanthus roseus. Science 244: 580-582.
Stoddard FL and McCully ME (1980) Effects of excision of stock and scion organs on the formation of
the graft union in coleus: A histological study. Bot Gaz 141: 401-412. van der Schoot C, Dietrich MA, Storms M, Verbeke JA and Lucas WJ (1995) Establishment of a
cell-to-cell communication pathway between separate carpels during gynoecium development. Planta
195: 450-455. Walker DB (1975) Postgenital carpel fusion in Catharanthus roseus (Apocynaceae). I. Light and scanning
electron microscopic study of gynoecial ontogeny. Am J Bot 62: 457-467.
Wang Y and Kollmann R (1996) Vascular differentiation in the graft union of in-vitro grafts with different compatibility. Structural and functional aspects. J Plant Physiol 147: 521-533.
植物科学最前線 6: 40 ( 2015)
BSJ- Revi ew 6: 40 ( 2015)