Botanical Inventory of the Allerton Garden

David H. Lorence, Ph.D., Director of Science and Tim Flynn, Collections Manager National Tropical Botanical Garden

3530 Papalina Road Kalaheo, HI 96741

Introduction and Historical Background The Allerton Garden is situated in the lower Lawai Valley, the southern portion of a 64-square mile watershed comprising a system of streams originating in the mountains above Lawai and feeding the Lawai Stream which empties into the ocean at Lawai-kai. Before Robert Allerton and John Gregg Allerton purchased the 83 acre Lawai-kai property in 1938, it had other owners and occupants, dating from the ancient Hawaiian period (pre-1848) and the Royal Period (1848-1886) to the McBryde Period (1886-1937). With the death of John Gregg Allerton in 1986, the Allerton Garden was placed in trust and today is managed by the National Tropical Botanical Garden under an agreement with the Allerton Gardens Trust (National Tropical Botanical Garden,1997) Each set of occupants have left their mark on the flora and vegetation of Lawai Valley, often bringing in a new suite of plants, while occasionally eliminating other earlier plantings, intentionally or not. For example, Queen Emma, wife of King Kamehameha IV, is said to have introduced many flowering plants and fruit trees including bougainvillea, mango, kiawe, tamarind, Crinum lilies, and other plants. Many of these species can still be found growing in the lower Lawai valley.

The Allerton Garden is a landscaped formal garden using tropical plants as a palette with shades of green and splashes of bright color. Consequently, most of the plants the Allertons used in their garden were non-native tropical plants obtained from a variety of sources, both locally and from their world voyages. It is a well-ordered paradise where nothing appears artificial, yet nature is so under control that not a leaf is out of place. Using the line of the Lawai Stream as the spine of the garden, the Allertons created a series of garden rooms that unfold between the stream bank and the cliffs. The varied topography that includes open meadows, moist jungle ravines, and rugged cliff faces, together with Kaua`i’s rich volcanic soil, provides ideal growing conditions for a wide selection of plants and trees. Monkeypod and other flowering trees form a cool, shady canopy sheltering carefully delineated rooms with water features surrounded by rows of palms or hedge plants and swaths of green background provided by pothos or taro vine (Epipremnum pinnatum and E. aureum ), Swiss cheese plant (Monstera deliciosa), oak-leaf fig (Ficus montana), and other ground covers. Additional form and color are provided by carefully placed plants such as spider lilies (Crinum pedunculatum), Queen Emma lily (Crinum augustum), ti plants (Cordyline fruticosa) or Dracaena species of various shades, and shrubs with delightfully fragrant flowers such as the rice plant (Aglaia odorata), Chloranthus inconspicuous, and the climbing ylang-ylang (Atrabotrys uncinnatus). One section is devoted to an impressive collection of towering heliconias, Costus, Calathea, and torch gingers (Etlingera elatior), while the palmetum contains many different species of exotic and native palms. Hillsides are planted with flowering Cassia species of assorted hues or varied Plumeria cultivars, and on the lower valley cliffs overlooking the ocean are cascades of red and purple Bougainvillea and pale blue-flowered Thunbergia vines. The visitor may stroll through a golden bamboo forest, listen to the

soothing sounds of running water by the fountains, stand concealed between buttresses of the huge Moreton Bay figs (Ficus macrophylla), or admire graceful palms and phenomenal vines with brilliant flowers, all in a historical context provided by the Allertons.

As a result of human impact and introduced weedy alien plants, few native Hawaiian plants survived in the lower region of Lawai Valley except along the coastal beach area and on the inaccessible rocky slopes and cliffs unsuitable for cultivation. These few remnant native cliff plants were the `akoko (Chamaesyce celastroides var. amplectans), a`alii (Dodonaea viscosa), `ala`ala wai nui (Plectranthus parviflorus), `ala`ala wai nui (Peperomia blanda var. floribunda), ko`oko`olau (Lipochaeta connata ssp. connata), `uhaloa (Waltheria indica), and `akia (Wikstromia uva-ursi var. kauaiensis, endemic to Kaua`i). The native coastal plants able to survive were naupaka kahakai (Scaevola sericea), pohuehue or beach morning glory (Ipomoea pes-caprae), hunakai (Ipomoea imperati), pa`uohi`iaka (Jacquemontia ovalifolia), seashore rushgrass or `aki`aki (Sporobolus virginicus), and the beach pea or mohihihi (Vigna marina). These plants had to be tough to survive under conditions that were hot and dry with little soil (cliffs and rocky slopes) or subject to salt spray and waves with sand substrate (coastal). Virtually all the other areas were taken by humans for their chosen plants. Recent collections growth. During the last 20 years, under the Allerton Garden Trust—NTBG management agreement, plants of known origin have been added, greatly enriching the Allerton Gardens collections. Dozens of species have been introduced by NTBG collectors who have brought back carefully documented seeds and cuttings from archipelagos and islands in Polynesia, Micronesia, and Melanesia, and the Mascarene Islands of the Indian Ocean. Additional palms have enhanced the palmetum as well as many cycad species, all endangered. A unique collection of 19 Indo-Pacific Pandanus (hala) species has been assembled representing the largest collection of its kind in the U.S.A. As Pandanus species are either male or female, each species must be represented by a group of two or more plants to ensure both sexes are represented. Pandanus leaves are used by indigenous cultures throughout the Pacific for weaving and other purposes and certain species in this collection are rare or endangered. NTBG’s botanical expeditions to various Pacific islands have yielded a diversity of lowland tropical species that have been assembled into a diverse riparian (riverine) forest, replacing the weedy thicket of haole koa (Leucaena leucocephala) formerly along the western side of Lawai Stream. Many species in this area are of cultural, medicinal, or economic importance on Pacific islands, and this collection represents an important resource for research, conservation, and educational purposes. Ecological Restoration. The most recent focus has been initiation of a project to restore the Lawai-kai coastal zone with native Hawaiian plants, including threatened and endangered species. This restoration not only provides natural protection to the fragile coastline in the face of extreme events such as hurricanes and tsunamis, but also provides a habitat matrix (or should this be plural?) for rare and endangered coastal plants and preserves nesting habitat for the green sea turtle. During 2004-2005 Dr. David Burney, Director of Conservation and Living Collections at NTBG, received funding from the U. S. Fish and Wildlife Service for a project to restore native vegetation along the Lawai-kai beach. All necessary permits were obtained before proceeding. For the Lawai-kai restorations, the general policy regarding choices of genetic stock is that seeds,

cuttings, or other plant material are to be derived from wild individuals or garden specimens of known pedigree. If possible, the stock used should be from south coast sources. If this is not available, material used should in any case be from Kaua`i. If the taxon is extirpated on Kaua`i but believed to have been present before humans on the basis of fossil occurrence, material from nearby islands such as Ni`ihau and Nihoa was considered, e.g. Pritchardia aylmer-robinsonii, endemic to Ni`ihau. The overall purpose of this project has been to ecologically restore the Lawai-kai coastal zone, focusing efforts on the coastal strand vegetation and its interface with exotic (non-native) vegetation. Additional goals include establishing a planting of native coastal mesic forest, and to demonstrate the value of habitat restoration to the visiting public. The ecological restoration of the Lawai-kai coastal zone focuses on delivering the following conservation benefits:

• Secure one of the few nesting habitats for green sea turtles on Kaua`i. • Manage invasive weeds threatening sandy beach habitats, including removal of exotic

grasses and selected ironwood trees. • Reinstate native vegetation, providing protection against hurricanes and tsunamis. • Establish populations of threatened native plant species within restored habitats. • Implement a coastal habitat restoration project that is open to public visitation and

provides educational opportunities. • Test the application of paleoecological data as a tool for guiding ecological restoration.

Dominant beach plants are naupaka (Scaevola sericea), pa`uohi`iaka (Jacquemontia ovalifolia), and pohuehue (Ipomoea pes-caprae). These provide a matrix for threatened and endangered species now successfully growing in the coastal beach area, including dwarf naupaka (Scaevola coriacea), `ohai (Sesbania tomentosa), popolo (Solanum nelsonii), loulu palm (Pritchardia aylmer-robinsonii), and Panicum niihauense. In addition, a variety of non Threatened and Endangered species have been established including `a`ali`i (Dodonaea viscosa), hinahina kolo (Vitex rotundifolia), and Fimbristylis cymosa. The goal is that once established, the populations of native plants will become self-sustaining. A second project to re-establish a Hawaiian lowland mesic forest in the vicinity of the Allerton “Jungle Garden”. This type of forest likely existed behind the coastal vegetation over a layer of rich deep soil with abundant ground water. Mesic forest species that have been planted here include three Federally listed endangered species, hau kuahiwi (Hibiscadelphus distans), koki`o (Kokia kauaiensis), and loulu (Pritchardia aylmer-robinsonii); the rare Ochrosia kauaiensis, and various non T & E species including kou (Cordia subcordata), wili`wili (Erythrina sandwicensis), alahe`e (Psydrax odorata), naio (Myoporum sanwicense), kokio keo`keo (Hibiscus waimeae ssp. waimeae), ko`oko`olau (Bidens sandwicensis), `ahu`awa (Cyperus javanicus), and `a`ali`i (Dodonaea viscosa). In a third project, restoration of native vegetation in two plots on the banks of the Lawai Stream in the Allerton and McBryde Gardens is underway. Replacing weeds with two key native species, `ahu`awa (Cyperus javanicus) and `ae`ae (Baccopa monnieri) is expected to help prevent erosion and control runoff and sedimentation and provide habitat for the common moorhen or `alae`ula (Gallinula chloropus sandvicensis). Planting these and other endemic and

indigenous riparian and coastal wetland species will reduce non-point pollution of the bay and adjacent shoreline areas as well. Educational and Research Opportunities The potential of this project to yield new information regarding restoration techniques in tropical coastal zones is immense. For this reason, the project has been instituted in such a way that scientific information may be gleaned and students at all levels, as well as the general public and Garden staff, may learn from the results. Finally, the restored area is being incorporated into the tour offerings of the Allerton Gardens, giving visitors an opportunity to see a practical application of plant science in the coastal zone. Trails through the restoration zones are being developed as a guided walk. Importance of beach as nesting site for green sea turtles. The coastal strand at Lawai-kai is a documented nesting habitat of the honu or green sea turtle (Chelonia mydas), one of only two documented sites on Kaua`i. Don Heacock (pers. comm., October 2003) of DLNR Aquatic Resources estimates that “an average of about eight” nests were deposited there every other year over the period 1988-2000. After a 7-year hiatus turtles are nesting at Lawai-kai again. During the full moon over the weekend of 2-3 June 2007 three female turtles deposited four egg clutches after digging cavities above the high water line (Richard Hanna, pers. obs. June 2007). The Conservation Department at NTBG has made a commitment to improve the nesting habitat for the green sea turtles. Invasive trees such as ironwood (Casuarina equisetifolia) and alien grasses with dense root systems such as zoysia (Zoysia matrella var. pacifica) make digging nests in the beach sand difficult for the turtles. We are replacing them with native coastal plants with loose root systems that hold the nest in place while the turtle is digging rather than hinder her efforts. These natives include: naupaka (Scaevola sericea), pa`uohi`iaka (Jacquemontia ovalifolia), and pohuehue (Ipomoea pes-caprae). Conclusions. Approximately 858 different species, varieties, and cultivars of flowering plants and ferns are known from the Allerton Garden today, including cultivated, native, and naturalized weedy species (see Appendix 1). In addition to species introduced within the historical context of Queen Emma and the Allertons, the Garden holds a great diversity of plants with important conservation, research, and educational potential. Finally, the restoration projects have resulted in a diversification of native Hawaiian coastal, riparian, and mesic forest habitats, which now provide a matrix for conserving a diversity of rare and endangered species. Hopefully the populations of native species will become self-sustaining. Improved native coastal habitat, in turn, provides increased nesting opportunities for the green sea turtle. Scientific names in Appendix 1 follow Staples and Herbst (2005) for the cultivated species and Wagner et al. 2009 for native and naturalized species.

References Cited. National Tropical Botanical Garden. 1997. Allerton Garden Reference Manual (unpublished

document). Staples, G. W. & D. R. Herbst. 2005. A Tropical Garden Flora. Bishop Museum Press. Wagner, W.L., D.R. Herbst, and S.H. Sohmer, 1999. Manual of the flowering plants of Hawaii,

revised edition with supplement by Wagner, W.L. and D.R. Herbst. Bishop Museum Special Publication 97, Honolulu, Hawaii, USA.

[Appendix 1 follows. ]

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

S

Plants of the Allerton GardenDATE: 14 June 2007C=cultivated; N=naturalized; I=indigenous; E=endemicSTATU FAMILY GENUS SP SUBSPTAX SECTIONC Acanthaceae Fittonia albivenisC Acanthaceae Hemigraphis alternataC Acanthaceae Hemigraphis reptansC Acanthaceae Megaskapasma erythrochlamysC Acanthaceae Odontonema cuspidatumC Acanthaceae Pseuderanthemum carruthersiiC Acanthaceae Ruellia devosianaC Acanthaceae Ruellia graecizansC Acanthaceae Sanchezia speciosaC Acanthaceae Thunbergia fragransC Acanthaceae Thunbergia hybridC, N Agavaceae Agave americana cv. MarginataC Agavaceae Agave attenuataC Agavaceae Agave coucui Tour RoadC Agavaceae Agave guiengola Tour RoadC Agavaceae Agave polyacantha Tour RoadC, N Agavaceae Agave sisalanaC Agavaceae Agave sp.C Agavaceae Cordyline fruticosaC Agavaceae Dracaena dracoC Agavaceae Dracaena fragransC Agavaceae Dracaena marginataC Agavaceae Dracaena reflexa cv. VariegataC Agavaceae Dracaena reflexa cv. Song of IndiaC Agavaceae Dracaena reflexa var. linearifoliaC Agavaceae Dracaena reflexa var. reflexaN Agavaceae Furcraea foetidaC Agavaceae Nolina bigeloviiC Agavaceae Nolina nelsonii Tour RoadC Agavaceae Nolina strictaC, N Agavaceae Sansevieria cylindricaC, N Agavaceae Sansevieria trifasciataC Agavaceae Yucca gloriosaI Aizoaceae Sesuvium portulacastrum Lawai KaiC Asphodeliaceae Aloe arborescensC Asphodeliaceae Aloe barbadensisC Asphodeliaceae Aloe barberae Tour RoadC Asphodeliaceae Aloe ciliaris Tour RoadC Asphodeliaceae Aloe maculataC Asphodeliaceae Aloe saponariaC Asphodeliaceae Aloe sp. Tour RoadC Asphodeliaceae Lomatophyllum citreumC Asphodeliaceae Lomatophyllum macrum Tour RoadC Amaranthaceae Alternanthera tenellaN Amaranthaceae Amaranthus dubiusN Amaranthaceae Amaranthus lividusN Amaranthaceae Amaranthus spinosusC Amaranthaceae Charpentiera densiflora Lawai Kai RestorationC Amaranthaceae Charpentiera elliptica Visitors CenterC Amaranthaceae Chenopodium oahuense Lawai Kai RestorationC Amaranthaceae Nototrichium sandwicense Lawai Kai RestorationC Anacardiaceae Campnosperma brevipetiolata RIPARIANC Anacardiaceae Mangifera indica FRUIT ORCHARDN Anacardiaceae Schinus terebinthifoliusC Anacardiaceae Spondias dulcisC Anacardiaceae Spondias tuberosaC Annonaceae Annona glabraC Annonaceae Artabotrys uncinatusC Annonaceae Rollinia pulchrinervis

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

N Apiaceae Centella asiaticaN Apiaceae Ciclospermum leptophyllumN Apiaceae Hydrocotyl sibthorpioidesC Apocynaceae Adenium coetaneumC Apocynaceae Adenium obesumC Apocynaceae Allamanda catharticaC Apocynaceae Alstonia macrophyllaC Apocynaceae Nerium oleanderC Apocynaceae Ochrosia ellipticaC Apocynaceae Plumeria obtusaC Apocynaceae Plumeria rubraC Apocynaceae Stemmadenia littoralisC, N Apocynaceae Thevetia peruvianaC Apocynaceae Thevetia thevetioidesC Araceae Aglaonema commutatumC Araceae Alocasia cucullataC Araceae Alocasia macrorrhizaC Araceae Alocasia zebrinaC Araceae Anthurium andraeanumC Araceae Anthurium hookeriC, N Araceae Colocasia esculentaC Araceae Dieffenbachia maculataC, N Araceae Epipremnum aureum PUMP 6C, N Araceae Epipremnum pinnatumN Araceae Lemna aequinoctialisC Araceae Monstera deliciosaC Araceae Monstera siltepecanaC Araceae Philodendron erubescensC Araceae Philodendron pinnatifidumC Araceae Philodendron sp.C Araceae Pistia stratiotesC Araceae Spathiphyllum cochlearispathumC Araceae Spathiphyllum kochiiC Araceae Syngonium nodifloraC Araceae Syngonium podophyllumC Araceae Typhonodorum lindleyanum Cutting GardenC Araliaceae Munroidendron racemosumC Araliaceae Polyscias guilfoyleiC Araliaceae Polyscias nodosaC Araliaceae Polyscias scutellariaC Araliaceae Reynoldsia sandwicensis Lawai KaiC Araliaceae Schefflera actinophyllaC Araliaceae Schefflera arboricolaC Arecaceae Acoelorrhaphe wrightii Hideo Teshima PalmetumC Arecaceae Actinorhytis calapparia Hideo Teshima PalmetumC Arecaceae Adonidia merrillii Hideo Teshima PalmetumC, N Arecaceae Archontophoenix alexandrae Hideo Teshima PalmetumC Arecaceae Archontophoenix cunninghamiana Hideo Teshima PalmetumC Arecaceae Archontophoenix purpurea Hideo Teshima PalmetumC Arecaceae Archontophoenix sp. Hideo Teshima PalmetumC Arecaceae Archontophoenix sp.C Arecaceae Areca catechu Hideo Teshima PalmetumC Arecaceae Areca triandra Hideo Teshima PalmetumC Arecaceae Arenga australasicaC Arecaceae Arenga engleri Hideo Teshima PalmetumC Arecaceae Arenga hookerianaC Arecaceae Arenga obtusifolia Hideo Teshima PalmetumC Arecaceae Arenga undulatifoliaC Arecaceae Asterogyne martianaC Arecaceae Attalea butyracea Hideo Teshima PalmetumC Arecaceae Basselinia sp.C Arecaceae Beccariophoenix madagascariensis Hideo Teshima PalmetumC Arecaceae Bismarckia nobilis Hideo Teshima Palmetum

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Arecaceae Brahea armata Hideo Teshima PalmetumC Arecaceae Brahea brandegeeiC Arecaceae Brahea cf. armataC Arecaceae Calyptrocalyx pachystachys Hideo Teshima PalmetumC Arecaceae Campecarpus fulcitus Hideo Teshima PalmetumC Arecaceae Carpentaria acuminata Hideo Teshima PalmetumC Arecaceae Caryota mitisC Arecaceae Caryota sp. Hideo Teshima PalmetumC Arecaceae Caryota urensC Arecaceae Chamaerops humilisC Arecaceae Chambeyronia macrocarpa Hideo Teshima PalmetumC Arecaceae Clinostigma ponapense RiparianC Arecaceae Clinostigma samoense Hideo Teshima PalmetumC Arecaceae Clinostigma savoryana Hideo Teshima PalmetumC Arecaceae Coccothrinax barbadensis Hideo Teshima PalmetumC Arecaceae Coccothrinax fragransC Arecaceae Coccothrinax sp. Hideo Teshima PalmetumC, N Arecaceae Cocos nuciferaC Arecaceae Corypha utanC Arecaceae Cyrtostachys lakkaC Arecaceae Cyrtostachys rendaC Arecaceae Dictyosperma album Hideo Teshima PalmetumC Arecaceae Drymophloeus oliviformis Hideo Teshima PalmetumC Arecaceae Dypsis cabadae Hideo Teshima PalmetumC Arecaceae Dypsis decaryi Hideo Teshima PalmetumC Arecaceae Dypsis decipiens Hideo Teshima PalmetumC Arecaceae Dypsis lastellianaC Arecaceae Dypsis leptocheilos Hideo Teshima PalmetumC Arecaceae Dypsis lutescens Hideo Teshima PalmetumC Arecaceae Dypsis madagascariensis Hideo Teshima PalmetumC Arecaceae Elaeis guineensis Hideo Teshima PalmetumC Arecaceae Gronophyllum pinangoides Hideo Teshima PalmetumC Arecaceae Gulubia macrospadixC Arecaceae Heterospathe elata Hideo Teshima PalmetumC Arecaceae Hydriastele palauensis RiparianC Arecaceae Hyophorbe indicaC Arecaceae Hyophorbe lagenicaulis Hideo Teshima PalmetumC Arecaceae Hyophorbe verschaffeltii Hideo Teshima PalmetumC Arecaceae Hyphaene coriacea Hideo Teshima PalmetumC Arecaceae Hyphaene natalensisC Arecaceae Hyphaene thebaicaC Arecaceae Laccospadix australasicaC Arecaceae Latania loddigesii Hideo Teshima PalmetumC Arecaceae Latania lontariodesC Arecaceae Latania verschaffeltii Hideo Teshima PalmetumC Arecaceae Licuala aurantiaca Hideo Teshima PalmetumC Arecaceae Licuala grandisC Arecaceae Licuala malayana Hideo Teshima PalmetumC Arecaceae Licuala naumannii Hideo Teshima PalmetumC Arecaceae Licuala peltata Mermaid Fountain AreaC Arecaceae Licuala spinosa Mermaid Fountain AreaC Arecaceae Livistona australisC Arecaceae Livistona carinensisC Arecaceae Livistona cf. mariae Hideo Teshima PalmetumC Arecaceae Livistona chinensisC Arecaceae Livistona copernicaC Arecaceae Livistona decipensC Arecaceae Nenga wendlandianaC Arecaceae Normanbya normanbyi Hideo Teshima PalmetumC Arecaceae Nypa fruiticans RiparianC Arecaceae Oncosperma filamentosum Hideo Teshima PalmetumC Arecaceae Phoenix canariensis Hideo Teshima PalmetumC Arecaceae Phoenix reclinata Hideo Teshima Palmetum

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Arecaceae Phoenix roebelinii Hideo Teshima PalmetumC Arecaceae Pinanga kuhlii Hideo Teshima PalmetumC Arecaceae Pinanga sp. Hideo Teshima PalmetumC Arecaceae Polyandrococos caudescens Hideo Teshima PalmetumC Arecaceae Pritchardia affinis Hideo Teshima PalmetumC Arecaceae Pritchardia arecina Hideo Teshima PalmetumC Arecaceae Pritchardia aylmer-robinsonii Lawai Kai restorationC Arecaceae Pritchardia cf. hardyi Hideo Teshima PalmetumC Arecaceae Pritchardia hardyi Hideo Teshima PalmetumC Arecaceae Pritchardia hillebrandii Hideo Teshima PalmetumC Arecaceae Pritchardia kaalae Hideo Teshima PalmetumC Arecaceae Pritchardia lanaiensis Hideo Teshima PalmetumC Arecaceae Pritchardia lanigera Hideo Teshima PalmetumC Arecaceae Pritchardia lowreyana Hau ThicketC Arecaceae Pritchardia martii Hideo Teshima PalmetumC Arecaceae Pritchardia minor Hideo Teshima PalmetumC Arecaceae Pritchardia munroi Hideo Teshima PalmetumC Arecaceae Pritchardia napaliensis Hau ThicketC Arecaceae Pritchardia pacifica Hideo Teshima PalmetumC Arecaceae Pritchardia remota Hideo Teshima PalmetumC Arecaceae Pritchardia sp. Hideo Teshima PalmetumC Arecaceae Pritchardia thurstonii Hau ThicketC Arecaceae Pritchardia viscosa Hideo Teshima PalmetumC Arecaceae Pritchardia waialealeana Hideo Teshima PalmetumC Arecaceae Ptychosperma cf. keiense Hideo Teshima PalmetumC Arecaceae Ptychosperma elegans Hideo Teshima PalmetumC Arecaceae Ptychosperma lauderbachii Hideo Teshima PalmetumC Arecaceae Ptychosperma lineare Hideo Teshima PalmetumC, N Arecaceae Ptychosperma macarthurii Hideo Teshima PalmetumC Arecaceae Ptychosperma propinquum Hideo Teshima PalmetumC Arecaceae Ptychosperma sp. Hideo Teshima PalmetumC Arecaceae Ravenea rivularis AllertonC Arecaceae Rhopaloblaste augusta Hideo Teshima PalmetumC Arecaceae Rhopaloblaste sp. Hideo Teshima PalmetumC Arecaceae Roystonea regiaC Arecaceae Sabal mauritiiformis Hideo Teshima PalmetumC Arecaceae Satakentia liukiuensis Hideo Teshima PalmetumC Arecaceae Serenoa repens Hideo Teshima PalmetumC Arecaceae Syagrus amara Hideo Teshima PalmetumC Arecaceae Syagrus romanzoffiana Hideo Teshima PalmetumC Arecaceae Thrinax parvifloraC Arecaceae Thrinax radiata Hideo Teshima PalmetumC Arecaceae Trithrinax brasiliensis Hideo Teshima PalmetumC Arecaceae Veitchia joannis Hideo Teshima PalmetumC Arecaceae Veitchia merrilliiC Arecaceae Veitchia montgomeryana Hideo Teshima PalmetumC Arecaceae Verschaffeltia splendida Hideo Teshima PalmetumC Arecaceae Wodyetia bifurcata Hideo Teshima PalmetumC Aristolochiaceae Aristolochia elegansC Aristolochiaceae Aristolochia gibertiC Aristolochiaceae Aristolochia giganteaC Aspleniaceae Asplenium nidus below Diana's fountainN Asteraceae Ageratum conyzoidesC Asteraceae Artemisia australis Lawai Kai RestorationN Asteraceae Bidens pilosaC Asteraceae Bidens sandvicensis Lawai KaiN Asteraceae Conyza bonariensisN Asteraceae Crassocephalum crepidiodesN Asteraceae Cyanthillium cinereumN Asteraceae Eclipta albaN Asteraceae Emilia fosbergiiN Asteraceae Erechtites valerianifoliaI Asteraceae Lipochaeta connata subsp. connata

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Asteraceae Lipochaeta succulenta Lawai Kai RestorationN Asteraceae Parthenium hysterophorusN Asteraceae Pluchea carolinensisN Asteraceae Pluchea indicaN Asteraceae Sigesbeckia orientalisN Asteraceae Sonchus oleraceusC, N Asteraceae Sphagneticola trilobataN Asteraceae Synedrella nodifloraC Asteraceae Wilkesia gymnoxiphium Lawai KaiC Averrhoaceae Averrhoa carambola FRUIT ORCHARDC Balsaminaceae Impatiens walleranaC Begoniaceae Begonia coccineaC Bignoniaceae Crescentia cujeteC Bignoniaceae Jacaranda mimosifoliaC Bignoniaceae Kigelia pinnataC, N Bignoniaceae Macfadyena unguis-catiN Bignoniaceae Spathodea campanulataC Bignoniaceae Tabebuia roseaC Bixaceae Bixa orellana FRUIT ORCHARDC Boraginaceae Cordia sebestenaC Boraginaceae Cordia subcordata Lawai KaiC Boraginaceae Heliotropium anomalum Lawai Kai RestorationC Boraginaceae Heliotropium curassavicum Lawai Kai RestorationC Boraginaceae Heliotropium foertherianumN Brassicaceae Cardamine flexuosaC Brassicaceae Lepidium bidentatum Lawai Kai RestorationN Brassicaceae Nasturtium microphyllumC Bromeliaceae Aechmea bracteataC Bromeliaceae Aechmea fulgensC Bromeliaceae Aechmea muricataC Bromeliaceae Billbergia pyramidalisC Bromeliaceae Bromelia sp.C Bromeliaceae Pitcairnea angustifoliaC Bromeliaceae Portea petropolitanaC Bromeliaceae Tillandsia cyaneaC Bromeliaceae Tillandsia sp.N Cactaceae Cereus uruguayensisC Cactaceae Epiphyllum phyllanthusC, N Cactaceae Hylocereus costaricensisC, N Cactaceae Hylocereus undatusC, N Cactaceae Opuntia cochenilliferaN Cactaceae Opuntia ficus-indicaC, N Cactaceae Selenicereus macdonaldiaeC Campanulaceae Brighamia insignis Bromeliad eastC Campanulaceae Delissea kauaiense Lawai Kai RestorationN Cannabaceae Trema cannabina RIPARIAN FORESTN Cannaceae Canna indicaC Capparaceae Capparis sandwichiana Lawai KaiC, N Caricaceae Carica papayaN Caryophyllaceae Drymaria cordata var. pacificaC, N Casuarinaceae Casuarina equisetifoliaC, N Casuarinaceae Casuarina glaucaC Celastraceae Elaeodendron orientaleC Chloranthaceae Chloranthus inconspicuousC Chrysobalanaceae Atuna racemosa JUNGLE SIDEC Cibotiaceae Cibotium glaucum along tram roadC Clusiaceae Calophyllum inophyllum Lawai KaiC, N Clusiaceae Clusea roseaC Clusiaceae Garcinia livingstonei FRUIT ORCHARDC Clusiaceae Garcinia madruno FRUIT ORCHARDC Clusiaceae Garcinia mangostanaC Clusiaceae Garcinia xanthochymus FRUIT ORCHARDC Clusiaceae Mammea americana ORCHARD

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Clusiaceae Terminalia bentzoeC Combretaceae Terminalia carolinensis JUNGLE SIDEC, N Combretaceae Terminalia catappaC Combretaceae Terminalia glabrataC Combretaceae Terminalia littoralisC Combretaceae Terminalia nitensC Combretaceae Terminalia samoensis RIPARIANC, N Commelinaceae Callisia fragransC Commelinaceae Cochliostema odoratissimum FRUIT ORCHARDN Commelinaceae Commelina diffusaC, N Commelinaceae Dichorisandra thyrsifloraC, N Commelinaceae Tradescantia spathaceaC, N Commelinaceae Tradescantia spathacea cv. ConcolorC, N Commelinaceae Tradescantia zebrinaC Connaraceae Connarus semidecandrus RIPARIANC Convolvulaceae Ipomoea batatasI Convolvulaceae Ipomoea imperati Lawai Kai RestorationI Convolvulaceae Ipomoea indicaI Convolvulaceae Ipomoea littoralis COASTALI Convolvulaceae Ipomoea pes-caprae subsp. brasiliensis Lawai KaiN Convolvulaceae Ipomoea trilobaI Convolvulaceae Jacquemontia ovalifolia subsp. sandwicensis Lawai KaiC Convolvulaceae Porana paniculataC Costaceae Costus comosusC Costaceae Costus malortieanusC Costaceae Costus productusC Costaceae Costus villosissimusC Costaceae Costus speciousC Costaceae Costus woodsoniiC Costaceae Tapeinocheilos ananassae Cut flower gardenC Crassulaceae Kalanchoe beharensisC Crassulaceae Kalanchoe blossfeldianaC Crassulaceae Kalanchoe diagremontianumN Crassulaceae Bryophyllum pinnataC, N Crassulaceae Bryophyllum tubifloraN Cucurbitaceae Momordica charantiaI Cucurbitaceae Sicyos herbstii Lawai KaiI Cucurbitaceae Sicyos pachycarpaC Cycadaceae Cycas angulataC Cycadaceae Cycas bougainvilleana Hideo Teshima PalmetumC Cycadaceae Cycas circinalisC Cycadaceae Cycas kennedyanaC Cycadaceae Cycas ophiolitica JUNGLE SIDEC Cycadaceae Cycas revolutaC Cycadaceae Cycas thouarsiiC Cyclanthaceae Carludovica palmataC Cyclanthaceae Cyclanthus bipartitusC Cyperaceae Carex meyenii Lawai KaiC Cyperaceae Carex wahuensis Lawai Kai RestorationI Cyperaceae Cladium jamaicense Stream RestorationN Cyperaceae Cyperus gracilisN Cyperaceae Cyperus involucratusN Cyperaceae Cyperus javanicus Lawai KaiN Cyperaceae Cyperus rotundusC Cyperaceae Cyperus trachysanthos Stream RestorationC Cyperaceae Fimbristylis cymosa Lawai KaiN Cyperaceae Kyllingia brevifoliaN Cyperaceae Schoenoplectus californicusC Cyperaceae Schoenoplectus tabernaemontani StreamC Dilleniaceae Dillenia indicaC Dilleniaceae Dillenia suffruticosaC Ebenaceae Diospyros blancoiC Ebenaceae Diospyros sandwicensis Visitors Center

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C, N Euphorbiaceae Aleurites moluccanaC Euphorbiaceae Chamaesyce celastroides Lawai KaiI Euphorbiaceae Chamaesyce celastroides var. amplectansC Euphorbiaceae Chamaesyce degeneri Lawai Kai RestorationN Euphorbiaceae Chamaesyce hirtaN Euphorbiaceae Chamaesyce hypericifoliaC Euphorbiaceae Codiaeum variegatumC Euphorbiaceae Euphorbia antiquorumN Euphorbiaceae Euphorbia heterophyllaC, N Euphorbiaceae Euphorbia lacteaC, N Euphorbiaceae Euphorbia tirucalliC Euphorbiaceae Excoecaria cochinchininensisC Euphorbiaceae Jatropha integerrimaC Euphorbiaceae Macaranga mappaC Euphorbiaceae Pedilanthus tithymaloidesN Euphorbiaceae Ricinus communisC Fabaceae Acacia koaC Fabaceae Acacia koaia Lawai Kai RestorationC Fabaceae Acacia simplexC Fabaceae Bauhinia cumingianaC Fabaceae Bauhinia forficataC Fabaceae Bauhinia monandraC Fabaceae Bauhinia punctataC Fabaceae Bauhinia variegata var. candidaC Fabaceae Brownea coccineaN Fabaceae Canavalia catharticaC Fabaceae Canavalia napaliensis Lawai KaiC Fabaceae Cassia fistulaC Fabaceae Cassia grandisC Fabaceae Cassia javanicaN Fabaceae Chamaecrista nictitans subsp. patellaria var. glabrataN Fabaceae Crotalaria pallidaC Fabaceae Cynometra ramifloraC Fabaceae Delonix regiaC Fabaceae Dendrolobium umbellatumC Fabaceae Derris trifoliata COASTAL FORESTN Fabaceae Desmanthus virgatusN Fabaceae Desmodium incanumN Fabaceae Desmodium trifoliumC Fabaceae Erythrina crista-galliC Fabaceae Erythrina sandwicensis Lawai Kai RestorationN Fabaceae Leucaena leucocephalaN Fabaceae Mimosa pudica var. unijugaC Fabaceae Mucuna novoguieensisC Fabaceae Mucuna toyoshimae RiparianC Fabaceae Ormosia monospermaC Fabaceae Phyllocarpus septentrionalisC, N Fabaceae Pithecellobium dulceC, N Fabaceae Prosopis pallidaC, N Fabaceae Samanea samanC Fabaceae Schleinitzia fosbergii RiparianN Fabaceae Senna surrattensisC Fabaceae Sesbania tomentosa Lawai Kai RestorationC Fabaceae Sophora tomentosa COASTALC Fabaceae Strongylodon macrobotrysC Fabaceae Tamarindus indicaI Fabaceae Vigna marina Lawai KaiC Gentianaceae Fagraea berteroanaC Gentianaceae Fagraea galilai RiparianC Goodeniaceae Scaevola coriacea BEACHI Goodeniaceae Scaevola taccada Stream RestorationC Heliconiaceae Heliconia acuminata Heliconia AreaC Heliconiaceae Heliconia aemygdiana

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Heliconiaceae Heliconia aurantiacaC Heliconiaceae Heliconia bihai cv. Claw 1C Heliconiaceae Heliconia bihai cv. Claw 2C Heliconiaceae Heliconia bihai cv. KamehamehaC Heliconiaceae Heliconia bihaiC Heliconiaceae Heliconia caribaea cv. CreamC Heliconiaceae Heliconia caribaea cv. PurpureaC Heliconiaceae Heliconia caribaea x bihai cv. Jacquinii CUTTING GARDENC Heliconiaceae Heliconia caribaea x bihai cv. CriswickC Heliconiaceae Heliconia caribaea x bihai cv. KawuchiC Heliconiaceae Heliconia caribaea x bihaiC Heliconiaceae Heliconia chartaceaC Heliconiaceae Heliconia collinsianaC Heliconiaceae Heliconia cv. GuyanaC Heliconiaceae Heliconia cv. Tropic FleurC Heliconiaceae Heliconia imbricata CUTTING GARDENC Heliconiaceae Heliconia indica cv. SpectabilisC Heliconiaceae Heliconia mariae CUTTING GARDENC Heliconiaceae Heliconia metallicaC Heliconiaceae Heliconia orthotricha cv. Pink FuzzyC Heliconiaceae Heliconia orthotricha cv. SheC Heliconiaceae Heliconia orthotrichaC Heliconiaceae Heliconia pendulaC Heliconiaceae Heliconia platystachysC Heliconiaceae Heliconia psittacorum cv. AndromedaC Heliconiaceae Heliconia psittacorum cv. ParakeetC Heliconiaceae Heliconia psittacorum CUTTING GARDENC Heliconiaceae Heliconia cv. Golden TorchC Heliconiaceae Heliconia rostrata CUT FLOWER GARDENC Heliconiaceae Heliconia sp CUT FLOWER GARDENC Heliconiaceae Heliconia sp. CUTTING GARDENC Heliconiaceae Heliconia sp. CUTTING GARDENC Heliconiaceae Heliconia sp. CUTTING GARDENC Heliconiaceae Heliconia sp. CUTTING GARDENC Heliconiaceae Heliconia sp. Heliconia AreaC Heliconiaceae Heliconia solomonensisC Heliconiaceae Heliconia spathocircinata CUTTING GARDENC Heliconiaceae Heliconia standleyiC Heliconiaceae Heliconia strictaC Heliconiaceae Heliconia sylvestrisC Heliconiaceae Heliconia talamanacana BEACHC Heliconiaceae Heliconia tarumaensisC Heliconiaceae Heliconia wagnerianaC Heliconiaceae Heliconia wagneriana or stricta CUTTING GARDENC Heliconiaceae Heliconia wagneriana or stricta CUTTING GARDENC Heliconiaceae Heliconia x nickeriensisC Hernandiaceae Hernandia nymphaefolia COASTALC Iridaceae Dietes grandifloraC Iridaceae Neomarica caerulaC Iridaceae Neomarica gracilisC Lamiaceae Callicarpa candicans RiparianC Lamiaceae Callicarpa elegans RIPARIAN FORESTC Lamiaceae Clerodendrum buchananiC Lamiaceae Clerodendrum indicumC Lamiaceae Leonurus sibiricusC Lamiaceae Plectranthus amboincusI Lamiaceae Plectranthus parviflorusC Lamiaceae Premna obtusifolia RiparianC Lamiaceae Tectona grandisC Lamiaceae Vitex cofassus RIPARIAN FORESTC Lamiaceae Vitex negundo East side of streamC Lamiaceae Vitex rotundifolia Lawai KaiC Lauraceae Cinnamomum burmannii

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Lauraceae Cinnamomum camphoraC Lauraceae Cinnamomum zeylanicum FRUIT ORCHARDC Lauraceae Laurus nobilis FRUIT ORCHARDC Lauraceae Persea americanaC Lecythidaceae Barringtonia asiaticaC Lecythidaceae Barringtonia edulis RiparianC Lecythidaceae Barringtonia racemosaC Liliaceae Agapanthus praecox subsp. orientalisC Liliaceae Chlorophytum comosumC Liliaceae Crinum augustumC Liliaceae Crinum mauritianum CUTTING GARDENC, N Liliaceae Crinum pedunculatumN Liliaceae Hippeastrum puniceumC Liliaceae Liriope muscariC Liliaceae Ophiopogon japonicusC Loganiaceae Geniostoma kusaiense RiparianC Loganiaceae Neubergia celebica ALLERTONC Lomariopsidaceae Nephrolepis cordifoliaC Lythraceae Lagerstroemia speciosaC Lythraceae Pemphis acidula COASTALC Lythraceae Punica granatum FRUIT ORCHARDC Maesaceae Maesa carolinensis ALLERTONC Malpighiaceae Galphimia gracilisC Malpighiaceae Malpighia coccigeraC Malpighiaceae Malpighia glabraC Malpighiaceae Malpighia punicifolia FRUIT ORCHARDC Malpighiaceae Stigmaphyllon ciliatumN Malvaceae Abutilon grandifoliumC Malvaceae Brachychiton acerifoliumC Malvaceae Chorisia speciosaC Malvaceae Commersonia bartramia RIPARIANC Malvaceae Dombeya cf. wallichii PANDANETUMC Malvaceae Durio graveolensC Malvaceae Durio zibethinusC Malvaceae Gossypium tomentosum Lawai KaiC Malvaceae Hibiscadelphus distans Lawai Kai RestorationC Malvaceae Hibiscus arnottianus Visitor's CenterC Malvaceae Hibiscus fragilisC Malvaceae Hibiscus genevii Hibiscus AlleyC Malvaceae Hibiscus glaberC Malvaceae Hibiscus kokio subsp. kokioC Malvaceae Hibiscus kokioC Malvaceae Hibiscus rosa-sinensisC Malvaceae Hibiscus scottiiC, N Malvaceae Hibiscus tiliaceusC Malvaceae Hibiscus waimeae Lawai KaiC Malvaceae Kokia drynarioides Lawai Kai RestorationC Malvaceae Kokia kauaiensis Lawai Kai RestorationC Malvaceae Malachra alceifoliaN Malvaceae Malvastrum coromandelianum subsp. C Malvaceae Malvaviscus arboreusC Malvaceae Matisia cordataC Malvaceae Pachira aquaticaI Malvaceae Sida fallax Lawai KaiC Malvaceae Sterculia foetidaC, N Malvaceae Thespesia populnea JUNGLE SIDEN Malvaceae Triumfetta semitrilobaI Malvaceae Waltheria indicaC Marantaceae Calathea burle-marxii cv. Ice BlueC Marantaceae Calathea crotaliferaC Marantaceae Calathea luteaC Marantaceae Donax cannaeformis CUTTING GARDENC Marantaceae Marantachloa flexuosa

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Marantaceae Pleiostachya pruinosaC Marcgraviaceae Norantea guianensisC Melastomataceae Medinella magnificaC Meliaceae Aglaia odorataC Meliaceae Aphanamixis polystachyaC Meliaceae Xylocarpus granatum RIPARIANC Menispermaceae Pachygone ledermannii RiparianC Monimiaceae Tambourissa peltataC Moraceae Artocarpus altilisC Moraceae Artocarpus altilis x mariannensisC Moraceae Artocarpus heterophyllus FRUIT ORCHARDC Moraceae Artocarpus lingnanensis Fruit OrchardC Moraceae Ficus carica FRUIT ORCHARDC Moraceae Ficus cf. virensC Moraceae Ficus elasticaC Moraceae Ficus lyrataC Moraceae Ficus macrophyllaC Moraceae Ficus microcarpaC Moraceae Ficus montanaC Moraceae Ficus pseudopalmaC Moraceae Ficus pumilaC Moraceae Ficus tinctoria West side of streamC Moraceae Morus alba FRUIT ORCHARDC Moraceae Treculia africana FRUIT ORCHARDC Moringaceae Moringa oleifera FRUIT ORCHARDC Musaceae Musa acuminataC Musaceae Musa ornataC Musaceae Musa uranoscopus CUTTING GARDENC Musaceae Musa velutinaC Musaceae Musella lasiocarpa CUTTING GARDENC Myristicaceae Horsfieldia nunu RIPARIANC Myrsinaceae Tapeinosperma sp. RiparianC, N Myrtaceae Eucalyptus degluptaC Myrtaceae Eugenia palumbis RIPARIANC Myrtaceae Eugenia uniflora FRUIT ORCHARDC Myrtaceae Feijoa sellowiana FRUIT ORCHARDC Myrtaceae Metrosideros polymorpha Lawai Kai RestorationC Myrtaceae Myrciaria cauliflora FRUIT ORCHARDC Myrtaceae Myrciaria vexator FRUIT ORCHARDC Myrtaceae Pimenta dioicaC, N Myrtaceae Psidium cattleianumC, N Myrtaceae Psidium guajava FRUIT ORCHARDC Myrtaceae Syzygium aromaticumC Myrtaceae Syzygium cuminiC Myrtaceae Syzygium malaccenseC Nelumbonaceae Nelumbo nuciferaN Nyctaginaceae Boerhavia coccineaC Nyctaginaceae Boerhavia repens Lawai KaiC Nyctaginaceae Bougainvillea glabraN Nyctaginaceae Mirabilis jalapaC Nymphaeaceae Nymphaea nouchaliC Nymphaeaceae Nymphaea sp.C Ochnaceae Ochna kirkiiC Oleaceae Jasminum sambacC Oleaceae Osmanthus fragransC Oleaceae Osmanthus insularis RiparianC Orchidaceae Brassavola nodosaC Orchidaceae Dendrobium sp.C Orchidaceae Epidendrum sp.C Orchidaceae Oncidium sphacelatumC Orchidaceae Vanda sp.C Oxalidaceae Averrhoa carambolaN Oxalidaceae Oxalis corniculata

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

N Oxalidaceae Oxalis debilisC Pandanaceae Freycinetia cummingianaC Pandanaceae Pandanus amaryllifolius PANDANETUMC Pandanaceae Pandanus aurantiacus PANDANETUMC Pandanaceae Pandanus balansae PANDANETUMC Pandanaceae Pandanus baptistii PANDANETUMC Pandanaceae Pandanus dubius PANDANETUMC Pandanaceae Pandanus fragrans PandanetumC Pandanaceae Pandanus kanehirae PANDANETUMC Pandanaceae Pandanus mei PANDANETUMC Pandanaceae Pandanus pedunculatusC Pandanaceae Pandanus polycephalus PandanetumC Pandanaceae Pandanus sp. PANDANETUMC Pandanaceae Pandanus sp. PANDANETUMC Pandanaceae Pandanus sp. POLYNESIAN INTRO. C Pandanaceae Pandanus tectorius PANDANETUMC Pandanaceae Pandanus tectorius laevis PANDANETUMC Pandanaceae Pandanus utilis PANDANETUMC Pandanaceae Pandanus vandermeerschii PANDANETUMC Pandanaceae Pandanus veitchii PANDANETUMC Pandanaceae Pandanus whitmeeanus PANDANETUMC Pandanaceae Sararanga philippinensis PANDANETUMC Papaveraceae Argemone glauca Lawai KaiC Passifloraceae Adenia keramanthusC Passifloraceae Passiflora vitifoliaC Phyllanthaceae Antidesma kusaiense West side of streamN Phyllanthaceae Phyllanthus debilisN Phyllanthaceae Phyllanthus tenellusN Phytolaccaceae Rivinia humilisI Piperaceae Peperomia blanda var. floribundaN Piperaceae Peperomia pellucidaC Pittosporaceae Pittosporum napaliensisN Plantaginaceae Plantago majorC Plumbaginaceae Plumbago zeylanica Lawai KaiC Poaceae Bambusa vulgarisN Poaceae Brachiaria muticaN Poaceae Chloris barbataI Poaceae Chrysopogon aciculatusN Poaceae Cynodon dactylonN Poaceae Dactylis glomerataN Poaceae Digitaria insularisN Poaceae Echinochloa crus-galliN Poaceae Eragrostis tenellaN Poaceae Eragrostis variabilis Lawai KaiN Poaceae Eremochloa ophiuroidesI Poaceae Heteropogon contortus Lawai KaiN Poaceae Melinis repensN Poaceae Oplismenus hirtellusN Poaceae Panicum maximumC Poaceae Panicum niihauense Lawai KaiN Poaceae Pennisetum clandestinumN Poaceae Pennisetum purpureumC Poaceae Saccharum officinarumN Poaceae Sporobolus dianderN Poaceae Sporobolus indicusN Poaceae Sporobolus virginicus Lawai KaiN Poaceae Stenotaphrum secundatumN Poaceae Thysanolaena maximaN Poaceae Zoysia matrella var. pacificaC Podocarpaceae Podocarpus elatusC, N Polygonaceae Antigonon leptopusC Polygonaceae Coccoloba uviferaC Polygonaceae Rumex albescens Lawai Kai

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C, N Polypodiaceae Microsorum grossusC Polypodiaceae Microsorum punctatum cv. CristatumC Polypodiaceae Platycerium alcicorneC Polypodiaceae Platycerium bifurcatumC Polypodiaceae Platycerium sp.C Pontedariaceae Eichornia crassipesN Portulacaceae Portulaca oleraceaN Portulacaceae Portulaca pilosaN Portulacaceae Talinum paniculatumC Primulaceae Lysimachia mauritiana Lawai Kai RestorationC Primulaceae Myoporum sandwicense Lawai KaiC, N Proteaceae Grevillea robustaC Proteaceae Macadamia integrifolia FRUIT ORCHARDN Pteridaceae Adiantum raddianumC Pteridaceae Doryopteris decipiens Lawai Kai RestorationN Pteridaceae Pellaea viridisN Pteridaceae Pityrogramma calomelanosC Rhamnaceae Colubrina asiatica JUNGLE SIDEC Rhizophoraceae Bruguiera gymnorhiza West side of streamC Rosaceae Osteomeles anthyllidifolia Lawai KaiC Rubiaceae Atractocarpus fitzalaniC Rubiaceae Bikkia comptonii RiparianC, N Rubiaceae Coffea arabica ALLERTONC Rubiaceae Cubanola dominguensis CUTTING GARDENC Rubiaceae Gardenia aubryi RIPARIANC Rubiaceae Guettarda heterosepala RIPARIAN FORESTC Rubiaceae Guettarda speciosa East side of streamC Rubiaceae Ixora casei RIPARIAN FORESTC Rubiaceae Ixora triantha RiparianC Rubiaceae Kadua haupuensis Lawai Kai RestorationC Rubiaceae Kadua littoralis Lawai KaiC Rubiaceae Kadua st.-johnii Lawai KaiC,N Rubiaceae Morinda citrifolia JUNGLE SIDEC Rubiaceae Morinda citrifolia var. potteriC Rubiaceae Morinda jasminoidesC Rubiaceae Mussaenda raiateensis RIPARIAN FORESTN Rubiaceae Paederia foetidaC Rubiaceae Psychotria hobdyi 3CC Rubiaceae Psychotria leptothyrsa RIPARIAN FORESTC Rubiaceae Psychotria sp. RIPARIANC Rubiaceae Psychotria sp. RIPARIANC Rubiaceae Psychotria sp. RIPARIANC Rubiaceae Psychotria sp. RiparianC Rubiaceae Psydrax odorata Lawai KaiI Ruppiaceae Ruppia maritimaC Rutaceae Citrofortunella microcarpa FRUIT ORCHARDC Rutaceae Citrus aurantiifolia FRUIT ORCHARDC Rutaceae Citrus jambhiri FRUIT ORCHARDC Rutaceae Citrus limon FRUIT ORCHARDC Rutaceae Citrus maxima FRUIT ORCHARDC Rutaceae Citrus mitis ORCHARDC Rutaceae Citrus paradisi FRUIT ORCHARDC Rutaceae Citrus reticulata FRUIT ORCHARDC Rutaceae Citrus sinensis FRUIT ORCHARDC Rutaceae Citrus x nobilis FRUIT ORCHARDC Rutaceae Citrus x paradisi FRUIT ORCHARDC Rutaceae Citrus x tangeloC Rutaceae Clausena lansiumC Rutaceae Fortunella sp. FRUIT ORCHARDC Rutaceae Melicope mucronulata Lawai Kai RestorationC Rutaceae Murrya paniculataC Rutaceae Triphasia trifoliaC Rutaceae Zanthoxylum dipetalum Lawai Kai

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Salicaceae Dovyalis hebecarpa FRUIT ORCHARDC Santalaceae Santalum freycinetianum Lawai Kai RestorationC Sapindaceae Allophylus timorensis RIPARIAN FORESTC Sapindaceae Dimocarpus longan FRUIT ORCHARDC Sapindaceae Dodonaea viscosa Lawai KaiC Sapindaceae Euphoria longan FRUIT ORCHARDC Sapindaceae Harpullia pendulaC Sapindaceae Litchi chinensis FRUIT ORCHARDC Sapindaceae Nephelium lappaceum FRUIT ORCHARDC Sapindaceae Sapindus oahuensis Lawai Kai RestorationC Sapotaceae Chrysophyllum cainitoC Sapotaceae Manilkara zapota FRUIT ORCHARDC Sapotaceae Pouteria cainito FRUIT ORCHARDC Sapotaceae Pouteria campechiana FRUIT ORCHARDC Sapotaceae Pouteria cf. obovata RIPARIAN FORESTC Sapotaceae Pouteria sapota FRUIT ORCHARDC Sapotaceae Pouteria viridis FRUIT ORCHARDC Sapotaceae Synsepalum dulcificum FRUIT ORCHARDI Scrophulariaceae Bacopa monnieriC Solanaceae Brugmansia x candidaC Solanaceae Lycium sandwicense Lawai KaiC Solanaceae Solandra maximaN Solanaceae Solanum americanumE Solanaceae Solanum nelsonii Lawai KaiN Solanaceae Solanum seaforthianumC Strelitziaceae Phenakospermum guyannenseC Strelitziaceae Ravenala madagascariensisC Strelitziaceae Strelitzia reginaeC Thelypteridaceae Cyclosorus interruptus Lawai KaiN Thelypteridaceae Thelypteris parasiticusC Thymelaeaceae Phalaria dispermaE Thymelaeaceae Wikstroemia uva-ursi var. kauaiensisC Urticaceae Dendrocnide latifolia RiparianC Urticaceae Elatosperma calcareum RiparianC Urticaceae Pilea cadiereiN Urticaceae Pilea microphyllaC Urticaceae Pipturus kauaiensis Steam RestorationN Verbenaceae Lantana camaraN Verbenaceae Stachytarpheta urticifoliaC Woodsiaceae Diplazium esculentumC Xanthorrhoeaceae Xanthorrhoea minorC Zamiaceae Bowenia serrulata DIANAC Zamiaceae Dioon eduleC Zamiaceae Dioon mejiaeC Zamiaceae Encephalartos altensteiniiC Zamiaceae Encephalartos arenariusC Zamiaceae Encephalartos cupidusC Zamiaceae Encephalartos dyerianusC Zamiaceae Encephalartos eugene-maraisiiC Zamiaceae Encephalartos feroxC Zamiaceae Encephalartos friderici-guilielmiC Zamiaceae Encephalartos ghellinckiiC Zamiaceae Encephalartos gratus HARRY'S VALLEYC Zamiaceae Encephalartos horridusC Zamiaceae Encephalartos humilisC Zamiaceae Encephalartos inopinusC Zamiaceae Encephalartos ituriensisC Zamiaceae Encephalartos laevifoliusC Zamiaceae Encephalartos lebomboensisC Zamiaceae Encephalartos lehmaniiC Zamiaceae Encephalartos longifoliusC Zamiaceae Encephalartos natalensisC Zamiaceae Encephalartos ngoyanus

Cultivated, native, and naturalized flowering plants and ferns of the Allerton Garden

Compiled by Tim Flynn and David Lorence, NTBG

C Zamiaceae Encephalartos paucidentatusC Zamiaceae Encephalartos princepsC Zamiaceae Encephalartos transvenosusC Zamiaceae Encephalartos trispinosusC Zamiaceae Encephalartos villosusC Zamiaceae Lepidozamia peroffskyana ALLERTONC Zamiaceae Stangeria eriopus HARRY'S VALLEYC Zamiaceae Zamia cremnophila below Diana's fountainC Zamiaceae Zamia lindenii BAMBOO GROVEC Zamiaceae Zamia loddogesiiC Zamiaceae Zamia pseudoparasiticaC Zamiaceae Zamia pumilaC Zamiaceae Zamia roezli ALLERTONC Zamiaceae Zamia sp. CYCAD AREAC Zingiberaceae Alpinia bilamellataC Zingiberaceae Alpinia calcarataC Zingiberaceae Alpinia carolinensis CUTTING GARDENC Zingiberaceae Alpinia formosana forma variegataC Zingiberaceae Alpinia nieuwenhuiziiC Zingiberaceae Alpinia purpurata cv. Eileen McDonaldC Zingiberaceae Alpinia purpurata cv. Jungle KingC Zingiberaceae Alpinia purpurata cv. Jungle QueenC Zingiberaceae Alpinia purpurata cv. TahitianC Zingiberaceae Alpinia purpurataC Zingiberaceae Alpinia sp. CUTTING GARDENC Zingiberaceae Alpinia sp. CUTTING GARDENC Zingiberaceae Alpinia sp. CUTTING GARDENC Zingiberaceae Alpinia sp. CUTTING GARDENC Zingiberaceae Alpinia sp. Heliconia AreaC Zingiberaceae Alpinia sp. Heliconia AreaC Zingiberaceae Alpinia vittataC Zingiberaceae Alpinia zerumbet Heliconia AreaC Zingiberaceae Alpinia zerumbet cv. VariegataC Zingiberaceae Etlingera elatiorC Zingiberaceae Etlingera sp.C Zingiberaceae Hedychium coccineum JUNGLE SIDEC Zingiberaceae Hedychium gardnerianumC Zygophyllaceae Guaiacum officinale