FRI—UW—8615December 1986

PUGET SOUND DREDGE DISPOSAL ANALYSIS (PSDDA)DISPOSAL SITE INVESTIGATIONS: PHASE 1 TRAWL STUDIES IN SARATOGA

PASSAGE, PORT GARDNER, ELLIOTT BAY AND COMMENCEMENT BAY, WASHINGTON

Principal Investigators

Paul A. Dinnel, David A. Armstrong, Bruce S. Miller, Robert F. DonnellyFisheries Research Institute, School of Fisheries

University of Washington, Seattle

Part I

Crab and Shrimp Studies

by

Paul A. Dinnel, David A. Armstrong, and Anthony Whiley

Part II

Demersal Fish Studies

by

Robert F. Donnelly, Bruce S. Miller, Robert R. Lauth, and Shelley C. Clarke

FINAL REPORT

31 December 1986

for

Washington Sea Grant Program in Cooperation withSeattle District, U.S. Army Corps of Engineers

Seattle, Washington

Approved:

Submitted 31 December 1986 ___ ‘4’,~~j~pbert Franiis, Director

TABLE OF CONTENTS

Page

LIST OF FIGURES iv

LIST OF TABLES viii

LIST OF APPENDICES ix

ACKNOWLEDGEMENTS xiv

EXECUTIVE SUMMARY xv

PART I: Crab and Shrimp Studies 1

INTRODUCTION 1

METHODS 2

Beam Trawl 2

Otter Trawl 2

Sample Sites and Stations 4

RESULTS 11

Dungeness Crab 11

Shrimp 24

DISCUSSION AND CONCLUSIONS 40

Dungeness Crab 40

Shrimp 44

PART II: Demersal Fish Studies 51

INTRODUCTION 51

MATERIALS AND METHODS 52

Laboratory Processing of Fish 58

Flatfish Diseases 58

Environmental Measurements 59

Data Analysis 59

RESULTS 60

11

Commencement Bay.62

Elliott Bay 72

Saratoga Passage 77

Port Gardner 82

DISCUSSION AND CONCLUSIONS 88

The Research Otter Trawl for Documenting

Fish Assemblage 88

Commencement Bay 92

Elliott Bay 93

Saratoga Passage 95

Port Gardner 96

LITERATURE CITED 98

APPENDIX 104

Part I 104

Part II 151

111

LIST OF FIGURES

Part I

No. Page

1. Diagrams of the beam trawl and otter trawl used in thisstudy

2. Map of Puget Sound showing the general locations of thepreliminary PSDDA disposal sites in the Main Basin 5

3. Map of Saratoga Passage showing the beam trawl and ottertrawl sampling stations in and around the preliminarySaratoga Passage disposal site 6

4. Beam trawl and otter trawl sample stations in Port Gardner.

5. Maps of Elliott Bay showing the beam trawl and otter trawlsampling stations by season in and around proposed disposalsites in Elliott Bay 9

6. Maps of Commencement Bay showing the beam trawl and ottertrawl sampling stations by season in and around the twopreliminary disposal sites in Commencement Bay 10

7. Maps of Saratoga Passage showing the approximate beam trawlDungeness crab and shrimp catches at each sample station byseason 12

8. Dungeness crab size-frequency histograms by season for crabscaught in Saratoga Passage 13

9. Comparative average densities of Dungeness crab at the NavyDisposal Site and the two control sites in Port Gardner byseason and by trawl type 15

10. Distribution of male Dungeness crabs caught by beam trawl inPort Gardner during the seasonal sampling in 1986 17

11. Distribution of female Dungeness crabs caught by beam trawlin Port Gardner during the seasonal sampling in 1986 18

12. Distribution by depth of all Dungeness crabs caught by beamtrawl in Port Gardner during seasonal sampling in 1986 20

13. Carapace width—frequency histograms of all male and femaleDungeness crabs caught by beam trawl in Port Gardner duringseasonal sampling in 1986 21

14. Carapace width—frequency histograms for all male Dungenesscrabs caught by beam trawl in Port Gardner during seasonalsampling in 1986 22

iv

No. Page

15. Carapace width—frequency histograms for all female Dungenesscrabs caught by beam trawl in Port Gardner during seasonalsampling in 1986 23

16. Maps of Elliott Bay showing seasonal beam trawl catches ofDungeness crab and shrimp at the sampling stations in ElliottBay 25

17. Beam and otter trawl shrimp catches by species and by seasonfor PSDDA Site 2 in Saratoga Passage 27

18. Distribution of shrimp (all species combined) caught by beamtrawl in Port Gardner during seasonal sampling in 1986 29

19. Distribution by depth of shrimp (all species combined) caughtby beam trawl in Port Gardner during seasonal sampling in1986 30

20. Number of shrimp caught per hectare by both beam and ottertrawls in the three preliminary disposal sites in PortGardner during seasonal sampling in 1986 31

21. Beam and otter trawl shrimp catches by site, by species and byseason for the two Elliott Bay proposed disposal sites 33

22. Maps of Commencement Bay showing beam trawl catches ofshrimp at the sampling stations in Commencement Bay 34

23. Beam and otter trawl shrimp catches by site, by species and byseason for the proposed disposal sites in Commencement Bay. . . 35

24. Distribution by depth and by species for all beam trawl—caught shrimp, all areas (except Port Gardner) and seasonscombined 37

25. Average carapace lengths by species and by depth ranges forall shrimp caught, all seasons and areas (except PortGardner) combined 38

26. Length—frequency histograms for all spot prawn and side—stripe shrimp caught during the three sampling seasons, allareas (except Port Gardner) combined 39

27. Length—frequency histograms for all coonstripe and humpbackshrimp caught during the three sampling seasons, all areas(except Port Gardner) combined 41

28. Length—frequency histograms for all smooth pink and pinkshrimp caught during the three sampling seasons, all areas(except Port Gardner) combined 42

V

No. Page

29. Annual commercial shrimp landing from Puget Sound (includingHood Canal) from 1935 to 1982 46

30. Map of Western Washington showing areas of commercial shrimpproduction from late 1800’s to mid—1930’s 47

Part II

1. Map of Commencement Bay showing locations sampled for bottom—fish on June 13 (summer) and September 8 (autumn) 54

2. Map of Elliott Bay showing locations sampled for bottomfish onJuly 3 (summer) and September 9 (autumn) 55

3. Map of Saratoga Passage showing locations sampled for bottom—fish on July 1 (summer) 56

4. Map of Port Gardner showing the station sampled for bottom—fish 57

5. Number (abundance) of fish caught in Commencement Bay, shownby station and season 63

6. Biomass (in grams) of fish caught in Commencement Bay, shownby station and season 63

7. Number (abundance) of English sole, Dover sole, and ratfishcaught in Commencement Bay during summer and autumn, shownby station 65

8. Biomass (in grams) of English sole, Dover sole, and ratfishcaught in Commencement Bay during summer and autumn 66

9. Species richness (total number of species) of fish caught inCommencement Bay, shown by station and season 67

10. Species diversity (H’) of fish caught in Commencement Bay,shown by station and season 67

11. Length frequency of English sole, shown by sex, caught inCommencement Bay during summer and autumn at 40 m 68

12. Length frequency of English sole, shown by sex, caught inCommencement Bay during autumn at 20 m 69

13. Number (abundance) of fish caught in Elliott Bay shown bystation and season 73

14. Biomass (in grams) of fish caught in Elliott Bay, shown bystation and season 73

vi

No. Page

15. Number (abundance) of English sole, Dover sole, slendersole, ratfish and blackbelly eelpout caught in Elliott Bayduring summer and autumn, shown by station 74

16. Biomass (in grams) of English sole, Dover sole, slendersole, ratfish and blackbelly eelpout caught in Elliott Bayduring summer and autumn, shown by station 75

17. Species richness (total number of species) of fish caughtin Elliott Bay, shown by station and season 76

18. Species diversity (H’) of fish caught in Elliott Bay, shownby station and season 76

19. Length frequency of English sole, shown by sex, caught atPSDDA 2, reference 1, during autumn in Elliott Bay 78

20. Number (abundance) of fish caught in Saratoga Passage duringsummer, shown by station 81

21. Biomass (in grams) of fish caught in Saratoga Passage duringsummer, shown by station 81

22. Number (abundance) of English sole, Dover sole, slendersole, Pacific hake and ratfish caught in Saratoga Passageduring summer, shown by station 83

23. Biomass (in grams) of English sole, Dover sole, slendersole, Pacific hake and ratfish caught in Saratoga Passageduring summer, shown by station 83

24. Species richness (total number of species) of fish caughtin Saratoga Passage during summer, shown by station 84

25. Species diversity (H’) of fish caught in Saratoga Passageduring summer, shown by station 84

26. Number (abundance) of fish caught in Port Gardner, shownby station and season 86

27. Biomass (in grams) of fish caught in Port Gardner, shownby station and season 87

28. Species diversity of fish (H’) caught in Port Gardner, shownby station and season 89

vii

LIST OF TABLES

Part I

No. Page

1. Average shrimp catches, lengths and weights (wet biomass)for all shrimp caught by otter trawl in the proposed PSODAdisposal sites in Saratoga Passage, Port Gardner, ElliottBay and Commencement Bay during all sample months(combined), 1986 45

2. Estimated average shrimp catches per hectare from ottertrawis conducted in selected areas of Hood Canal and PugetSound from 1967 to 1979 49

3. Shrimp weights per hectare from the Elliott Bay preliminarydisposal sites as estimated from the otter trawl catches inJune and September 1986 50

Part II

1. List of bottomfish species caught by otter trawl duringthis study 61

2. Percent incidence and sample size of blood worm Philometrasp. infection in flatfish caught in Commencement Bay, shownby species, station and season 70

3. Environmental measurements of temperature, salinity, andwater clarity in Commencement Bay during autumn by station. . 71

4. Percent incidence and sample size of blood worm Philometrasp. infection in flatfish caught in Elliott Bay, shown byspecies, station and season 79

5. Environmental measurements of water temperature, dissolvedoxygen, salinity and water clarity in Elliott Bay, by stationand season 80

6. Percent incidence and sample size of bloodworm Philonietra sp.infection in flatfish caught at Saratoga Passage duringsummer, shown by station and species 85

7. Percent incidence and sample size of bloodworm Philometra sp.infection in flatfish, shown by species, station and seasonat Port Gardner 90

8. Measurements of temperature, salinity and water clarity bystation and season at Port Gardner 91

viii

LIST OF APPENDICES

Part I

AppendixTable No. Page

1. Saratoga Passage beam trawl station location data 105

2. Summary of Port Gardner beam trawl station location data . 106

3. Elliott Bay beam trawl station location data 109

4. Commencement Bay beam trawl station location data 110

5. Dungeness crab catches per hectare in Saratoga Passageduring February and June 1986 112

6. Beam trawl shrimp catches per hectare in Saratoga Passageduring February and June 1986, and in Elliott and Commencementbays during February, June and September 1986 113

7. Number of shrimp caught per hectare by otter trawl in SaratogaPassage in June, and in Elliott and Commencement bays in Juneand September 1986 115

8. Beam trawl catches of Dungeness crab from Port Gardner duringwinter 117

9. Dungeness crab densities per hectare calculated from beamtrawl catches in Port Gardner during April 1986 121

10. Dungeness crab densities per hectare calculated from beamtrawl catches in Port Gardner during June 1986 . 125

11. Dungeness crab densities per hectare calculated from beamtrawl catches in Port Gardner during September 1986 130

12. Dungeness crab densities per hectare calculated from beamtrawl catches at extra stations in Port Gardner duringSeptember 1986 134

13. Commercial shrimp densities per hectare in Port Gardnercalculated from beam trawl catches in February and April1986 135

14. Densities per hectare of Dungeness crabs and commercialshrimp calculated from otter trawl catches from the Februaryand April cruises in Port Gardner 139

15. Dungeness crab densities per hectare calculated from ottertrawl catches in Port Gardner in June and early July 1986. . 141

ix

AppendixTable No. Page

16. Commercial shrimp densities per hectare calculated from beamand otter trawls in Port Gardner in June and early July1986 143

17. Shrimp densities per hectare calculated from both beam andotter trawl catches in Port Gardner during September 1986. . • 147

18. Shrimp densities per hectare calculated from both beam andotter trawl catches at extra stations in Port Gardner duringSeptember 1986 150

Part II

Appendix

APPENDIX A Analysis of fish data collected by beam trawls inCommencement Bay, Elliott Bay, and Saratoga Passageduring 1986 152

Table 1. Abundance, biomass, species richness and speciesdiversity of fish caught by beam trawl in Commencement Bay during summer 1986 154

Table 2. Abundance, biomass, species richness and speciesdiversity of fish caught by beam trawl in ElliottBay by season 154

Table 3. Abundance, biomass, species richness and speciesdiversity of fish caught by beam trawl in SaratogaPassage during summer 155

APPENDIX B Length—frequency histograms of abundant,non—commercially or recreationally important fishcaught in Commencement Bay and Elliott Bay 156

Figure 1. Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during summerat 156 m in Commencement Bay 157

Figure 2. Length frequency of otter trawl caught ratfish,shown by sex, during summer at PSDDA 1 in Commencement Bay 157

Figure 3. Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during autumnat PSDDA 1 in Commencement Bay 158

Figure 4. Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during summerat PSDDA 1 reference site in Elliott Bay 158

x

Appendix Page

Figure 5. Length frequency of otter trawl caught ratfish,shown by sex, during autumn at PSDDA 2, referencesite 1 in Elliott Bay 159

Figure 6. Length frequency of otter trawl caught ratfish,shown by sex and life history stage, duringautumn at PSDDA 2, reference site 2 in ElliottBay 159

Figure 7. Length frequency of otter trawl caught slendersole, shown by sex and life history stage, duringautumn at PSDDA 1 in Elliott Bay 160

Figure 8. Length frequency of otter trawl caught slendersole males during summer at PSDDA 2 in ElliottBay 160

Figure 9. Length frequency of otter trawl caught slendersole, shown by sex, during autumn at PSDDA 2 inElliott Bay 161

APPENDIX C Abundance and biomass (and range at multiple samplestations) of otter trawl caught fish by station andspecies in Commencement Bay, Elliott Bay, andSaratoga Passage 162

Table 1. Abundance and range at multiple sample stations ofotter trawl—caught fish by station and species inCommencement Bay on June 13, 1986 163

Table 2. Biomass (in grams) and range at multiple samplestations of otter trawl—caught fish by station andspecies in Commencement Bay on June 13, 1986 165

Table 3. Abundance and range at multiple sample stations ofotter trawl—caught fish by station and species inCommencement Bay on September 8, 1986 167

Table 4. Biomass (in grams) and range at multiple samplestations of otter trawl—caught fish by station andspecies in Commencement Bay on September 8, 1986. 169

Table 5. Abundance and range of multiple sample stations forotter trawl—caught fish by station and species inElliott Bay on July 3, 1986 171

Table 6. Biomass (in grams) and range at multiple samplestations for otter trawl—caught fish by station andspecies in Elliott Bay on July 3, 1986 172

xi

Appendix Page

Table 7. Abundance and range at multiple sample stationsof otter trawl—caught fish by station and speciesin Saratoga Passage on July 1, 1986 173

Table 8. Biomass (in grams) and range at multiple samplestations of otter trawl caught fish by station andspecies in Saratoga Passage on July 1, 1986 175

Table 9. Number (abundance) of fish, biomass (in grams) andrange at Navy and PSDDA sites of otter trawl caughtfish by station and species in Port Gardner onFebruary 12 and 13, 1986 177

Table 10. Number (abundance) of fish, biomass (in grams) andrange at Navy and PSDDA sites for otter trawlcaught fish, by station and species in Port Gardneron April 18 and 21, 1986 179

Table 11. Number (abundance) of fish, biomass (in grams) andrange at Navy and PSDDA sites for otter trawlcaught fish by station and species in Port Gardneron June 30 and July 2, 1986 181

Table 12. Number (abundance) of fish, biomass (in grams) andrange at Navy and PSDDA sites for otter trawlcaught fish by station and species in Port Gardneron September 11 and 15, 1986 183

APPENDIX D Abundance and biomass (and range of multiple samplestations) of beam trawl—caught fish by station andspecies in Commencement Bay, Elliott Bay andSaratoga Passage 185

Table 1. Abundance and range of multiple sample stations ofbeam trawl—caught fish by station and species inCommencement Bay during July 1986 186

Table 2. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Commencement Bay during July 1986 187

Table 3. Abundance and range at multiple sample stations ofbeam trawl—caught fish by station and species inElliott Bay during June 1986 188

Table 4. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Elliott Bay during June 1986 189

xii

Appendix Page

Table 5. Abundance and range at multiple sample stationsof beam trawl—caught fish by station and speciesat Elliott Bay during September 1986 190

Table 6. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Elliott Bay during September 1986 191

Table 7. Abundance and range of multiple sample stations ofbeam trawl—caught fish by station and species inSaratoga Passage during June 1986 192

Table 8. Biomass (in grams) and range at. multiple samplestations of beam trawl—caught fish by station andspecies in Saratoga Passage during June 1986 193

APPENDIX E Number of flatfish per hectare caught by otter trawl inCommencement Bay, Elliott Bay, Saratoga Passage, andPort Gardner, shown by season, station and species. 194

Table 1. Number of each flatfish species per hectare caught byotter trawl at each station in Commencement Bayduring summer and autumn 1986 195

Table 2. Number of each flatfish species per hectare caught byotter trawl at each station in Elliott Bay duringsummer and autumn 1986 196

Table 3. Number of each flatfish species per hectare caught byotter trawl at each station in Saratoga Passage duringsummer 1986 197

Table 4. Number of each flatfish species per hectare caught byotter trawl at each station in Port Gardner duringwinter 1986 198

Table 5. Number of each flatfish species per hectare caught byotter trawl at each station in Port Gardner duringspring 1986 199

Table 6. Number of each flatfish species per hectare caught byotter trawl at each station in Port Gardner duringsummer 1986 200

Table 7. Number of each flatfish species caught by otter trawlat each station in Port Gardner during autumn1986 201

xiii

ACKNOWLEDGMENTS

The work was supported by the Washington Sea Grant Program in cooperation

with the Seattle District, U.S. Army Corps of Engineers. We appreciate the

valuable assistance of Louie Echols, Washington Sea Grant Program; Frank

Urabeck, Warren Baxter, Steve Martin and David Kendall of the U.S. Army Corps

of Engineers; and members of the PSDDA Disposal Site Work Group (DSWG). All

trawl work was conducted on board the R.V. Kittiwake skippered by Charles

Eaton. Assistance in the field and laboratory was provided by Russ McMillan,

Tom Wainwright, Greg Jensen, Dan Doty, Debbie Dole, Loann Hallurn, and Sandy

O’Neil. Carol Sisley, Carla Norwood and Kathy Boaz provided assistance with

report preparation.

xiv

EXECUTIVE SUMMARY

The multi—agency Puget Sound Dredge Disposal Analysis (PSDDA)

Program has been delegated the task of evaluating, selecting, monitoring

and managing sites within the inland waters of Washington State for

long—term, unconfined disposal of uncontaminated dredged materials. The

Disposal Site Work Group (DSWG) of PSDDA was assigned the responsibility

of selecting unconfined, open water disposal sites in central Puget

Sound. DSWG selected seven preliminary disposal sites within five Zones

of Siting Feasibility (ZSF’s) in central Puget Sound (Saratoga Passage,

Port Gardner, Elliott Bay and Commencement Bay) based on 19 selection

factors covering physical parameters, human uses and historical

biological resource data. Final site selection is now dependent, in

part, on site—specific trawl investigations for biological resources in

and around each of the five ZSF’s.

This document is the final technical report detailing the results

of trawl studies conducted in each of the five ZSF’s during 1986. This

report is divided into two parts: Part I summarizes the results of

trawling conducted with a small 3—ni research beam trawl especially

useful for capturing Dungeness crab, shrimp and small bottomfish.

Part II details the results of trawling conducted with a research

(7.6 m) otter trawl primarily designed to capture bottomfish of all

sizes.

Initial trawis in central Puget Sound identified three faunal

groups of specific importance to Puget Sound commercial and sport

fisheries: Dungeness crab, pandalid shrimp, and bottomfish (especially

flatfish, Pacific hake, cod, and rockfish). Each of these resources has

been analyzed in this report to provide the best possible biological

xv

data base for the final site selection process and to provide a

“baselin&’ of information for future monitoring of these disposal sites.

It should be noted that the Port Gardner data presented in this report

are essentially abstracted from cruise reports from a closely related

project (Navy Homeport Project). The final analyses of the Port Gardner

data will be available at a later date.

Generally, Dungeness crab were found to be absent from Commencement

Bay, of only minor concern in Elliott Bay and Saratoga Passage, and a

major resource (especially females) in Port Gardner, where this species

will be of primary concern in site selection and future monitoring.

Shrimp were ubiquitous throughout the areas sampled. Shrimp

populations were highly variable depending on such factors as site,

species, depth, season and habitat type. Shrimp populations were

generally insignificant as commercial or sport resources in all of the

five ZSF’s with the possible exception of the inner Elliott Bay ZSF.

Shrimp in this particular area may prove to be a siting concern, although

this area is also heavily impacted by gilinet fishing for salmon, ship

navigation and anchorage, and toxic contaminants in the nearby Duwamish

Waterways.

Bottomfish were sampled by a research trawl and it is important to

understand that the data generated are not comparable to that generated

by commercial trawls, upon which the Washington Department of Fisheries

bases its “flatfish index.~’

Bottomfish were low in abundance, biomass and species diversity at

the Commencement Bay PSDDA sites. In contrast, bottomfish were highest

in abundance, biomass and species diversity at the PSDDA sites in

xvi

Elliott Bay, when compared with the other locations sampled in Elliott

Bay. Saratoga Passage PSDDA sites were not adequately sampled to make

a concluding statement, but WDF studies have previously indicated they

may be an important area to some commercial fishes. Preliminary analysis

of Port Gardner bottomfish indicates that biomass and abundance

decrease with depth and towards the mouth of Port Gardner, and are at

maximum values during the winter.

xvii

PUGET SOUND DREDGE DISPOSAL ANALYSIS (P5DDA)DISPOSAL SITE INVESTIGATIONS

PART I

Crab and Shrimp Studies

by

Paul A. Dinnel, David A. Armstrong, and Anthony Whiley

INTRODUCTION

In January 1986, the Disposal Site Work Group (DSWG) of the Puget Sound

Dredge Disposal Analysis (PSDDA) team, selected preliminary preferred and

alternative sites for the unconfined disposal of dredged materials in the main

basin of Puget Sound (Phase I area). Initial site selections were based on

information gathered from limited field studies conducted within the ZSF’s

(Zones of Siting Feasibility) and existing information from each ZSF.

Selection of final preferred and alternative disposal sites required more

detailed evaluations of important physical factors and biological resources in

and around the identified sites. One of the key factors in choosing final

sites will be an evaluation of important benthic and epibenthic fisheries

resources including Dungeness crab (Cancer magister), commercial (Pandalid)

shrimp and bottomfish.

The purpose of this report is to describe the findings of the trawling

studies conducted in and around each of the preliminary PSDDA disposal sites

during February, April, June and September of 1986. The trawls conducted

during these seasons consisted of beam trawls known to be effective for

capturing Dungeness crab but which also sampled shrimp and smaller bottomfish

incidental to crabs. Demersal fauna were additionally sampled by a medium-

2

sized otter trawl especially effective in capturing larger bottomfish and

shrimp.

METHODS

Beam Trawl

All trawling was conducted aboard the 16 m research vessel Kittiwake.

Dungeness crabs were sampled with a 3 m beam trawl (Figure 1, top; Qunderson

and Ellis 1986) previously used elsewhere in Puget Sound (Dinnel et al. 1985a,

1985b, 1986; Weitkamp et al. 1986). The beam trawl was towed at each station

approximately 232 meters (1/8 nautical mile) at a target ground speed and time

of 2.5 km/hr (1 .5 knots) for 5.5 minutes which yielded an area swept by the

net (opening = 2.3 meters) of 534 m2. All crabs caught in the trawl were

measured, sexed, and assessed for molt condition (degree of shell softness)

and reproductive condition (females with or without eggs) and returned to the

water. Catches of shrimp and fish from the beam trawls were preserved for

later processing in the laboratory.

Otter Trawl

Bottomfish and shrimp were sampled with a 7.6 m otter trawl (Figure 1,

bottom) designed for the Southern California Coastal Water Research Project

(Mearns and Allen 1978). The otter trawl was towed approximately 370 m (i/snautical mile) at a target ground speed and time of 4.2 km/hr (2.5 knots) for

5.3 minutes which yielded an area swept by the net (opening = 6 m) of 2,220

m2. Incidentally caught crabs were processed on board as described above and

returned to the water. Bottomfish and shrimp were identified and counted on

board ship, and then at the end of the day, frozen for later processing in the

laboratory. Laboratory processing for shrimp included identification to

3

Figure 1. Diagrams of the beam trawl (top) and otter trawl (bottom)used in this study~

he4drope (wEth i’Yomts)

cI~w)

brWles1~flg

Ie9 Is.nes

4

species (commercial species only), measurement of carapace lengths and

assessments of reproductive condition (shrimp with or without eggs).

Bottomfish processing included identification to species, measurements for

length and weight, and a check for obvious external abnormalities or parasites

(primarily flatfish). See Part II for further discussion of the bottomfish

methods and results.

Sample Sites and Stations

Beam trawls. Beam trawl sampling was conducted in and around seven

preliminary preferred or alternative disposal sites in four general areas of

Puget Sound (Figure 2). The Saratoga Passage site was surveyed in February

and June 1986, but not in September as the site was viewed as the second

alternative to the Port Gardner preferred site. The trawl stations in Saratoga

Passage consisted of three stations within the preliminary disposal site;

eight stations stratified by depth (10, 20, 40 and 80 m below mean lower low

water [MLLW]) along Transect 1 east and west of the disposal site; and three

stations along Transect 2 north of the disposal site (Figure 3). Transect 2

was sampled only in June.

Beam trawl sampling in Port Gardner was conducted during four seasons

(February, April, June and September 1986). Sampling was conducted at two

preliminary disposal sites and along seven north—south transects crossing Port

Gardner (sample depths from 10 to 165 m) (Figure 4). Three stations each were

sampled within PSDDA Sites 1 and 2 during each season. The boundaries of

PSDDA Site 1 were moved slightly eastward prior to sampling in September

(dashed circle, Figure 4). Thus, this new site included PSDDA 1 Stations 1

and 2, Transect 3 Station 130M and Station H (which was added in September to

provide better sampling coverage of the new site) (Figure 4). The Transect 1,

I-,.

0 fr~ 0 t”J

frh

00

~daQ

00

Ort

~c1

~0

00

0 (I)CI

)

H’

0 H’

rt0

CD H’

CD

CD

CD

H’

~i) 0 C) CD rt 0 CI) 0 r-t CD CD CD CD

Lu

6

PSDDA 2

SaratogaPassage

w-Q-E

WhidbeyIsland

2NM

Figure 3. Map of Saratoga Passage showing the beam trawl (.) and otter trawl (D)sampling stations in and around the preliminary Saratoga Passage(PSDDA 2) disposal site. The dash line indicates the area of theZone of Siting Feasibility (ZSF).

PO

RT

GA

RD

NE

R

-I

—

NO

RT

H

SO

UT

H

EV

ER

ET

T

Fig

ure

4.B

eam

tra

wl

(0)

and

ott

er

tra

wl

(0)

sam

ple

sta

tio

ns

inP

ort

Ga

rdn

er.

De

pth

sin

me

ters

.N

=N

ort

h,

MM

idd

le,

S=

So

uth

.

8

40N Station was dropped from the sampling array after February 1986 due to

repeated gear damage at this location. Some additional beam trawl stations

(A—H; Figure 4) were also added in June and September to provide better

sampling coverage of the inner portion of the bay in relation to another

proposed (NAVY) disposal site (Dinnel et al. 1987).

Beam trawl sampling in Elliott Bay was conducted during three seasons

(February, June and September 1986) at two preliminary disposal sites and

along two nearshore transects (10, 20, 40 and 80 m depths) (Figure 5). Three

stations were sampled within PSDDA Site 1 in inner Elliott Bay during February

and June. The location of PSDDA Site 1 was relocated slightly eastward prior

to the September trawls; hence, two additional stations (Stations 4 and 5;

Figure 5) were added to better characterize the resources within the new site.

Three stations were trawled within PSDDA Site 2 (off Fourmile Rock) during all

three seasons. The 40 m depth along Transect 2 off Fourmile Rock was deleted

due to a rough bottom and repeated gear damage.

Beam trawl sampling in Commencement Bay was conducted during three

seasons (February, June and September 1986) at two preliminary PSDDA disposal

sites and along two transects stratified by depth (10, 20, 40, 80 and 120 m

below MLLW) (Figure 6). Three trawis each were made in PSDDA Sites 1 and 2 in

February. By June, PSDDA Site 2 had been relocated to the north of PSDDA Site

1 based on information about relative deposition/erosion potential from the

depositional analysis procedure. This new site (called PSDDA Site 2B in this

report, Figure 5) was trawled together with PSDDA Site 1 in June. PSDDA Sites

1 and 2B were again sampled in September except that a slight shift of PSDDA

Site 1 eastward resulted in the addition of one new station (PSDDA Site 1,

Station 4; Figure 6) during this season. As was the case in Elliott Bay,

rough bottom conditions resulted in the deletion of the 80 m station on the

EL

LIO

TT

BA

Y

Fig

ure

5.M

aps

of

Ellio

ttB

aysh

ow

ing

the

beam

tra

wl

(•)

and

ott

er

tra

wl

(a)

sa

mp

ling

sta

tio

ns

byse

aso

nin

and

aro

un

dth

etw

op

rop

ose

dD

isp

osa

lS

ite

sin

Ellio

ttB

ay.

The

da

she

dlin

ea

rea

sin

dic

ate

the

Zon

eso

fS

itin

gF

ea

sib

ility

(ZS

F).

The

do

tte

dlin

ew

ith

inPS

DD

AS

ite

1(S

ep

tem

be

r)w

asth

esite

orig

ina

lly

se

lecte

dw

ith

inZS

F1

and

the

so

lid

lin

ea

rea

show

sth

ea

pp

roxim

ate

loca

tio

no

fth

en

ew

lylo

ca

ted

site

.

10

COMMENCEMENT BAY

Figure 6. Maps of Commencement Bayshowing the beam trawl (•)and otter trawl (D) samplingstations by season in andaround the two preliminarydisposal sites in CommencementBay. The dashed line areashows the Zone of SitingFeasibility (ZSF). The dottedline areas show areas ofprior locations for thepreliminary disposal sitesand the solid lines show thepresent disposal site locations.

11

west end of Transect 1.

The exact locations, depths and trawl directions for all trawl stations

are recorded in Appendix Tables 1-4.

Otter trawls. Otter trawling was conducted at selected beam trawl

stations in Saratoga Passage in June, in Port Gardner in February, April, June

and September, and in Elliott and Commencement bays in June and September.

See Part II of this report for further discussion of the otter trawl work and

the results of the beam trawl fish catches.

RESULTS

Dungeness Crab

Saratoga Passage. The overall average beam trawl catches of Dungeness

crab in Saratoga Passage were 1.2 and 42.8 crab/hectare (ha) for the February

and June cruises, respectively. Dungeness crab were never caught within the

preliminary PSDDA disposal site or at the deep Transect 2 stations north of

the PSDDA site (Figure 7). All Dungeness crab were caught along Transect 1 on

both east and west sides of Saratoga Passage. Station 1OE (10 meters deep —

east side) had the highest catches of Dungeness crab in both February and

June. Only one Dungeness crab was caught by the otter trawl in June, this

being at Station 20E on the Camano Island side of Saratoga Passage. Dungeness

crab catches for both trawl types and for both seasons are summarized in

Appendix Table 5.

Histograms of Dungeness crab carapace width-frequencies show that about

95% of the crabs caught were mature animals ranging in size from 100 to 165 mm

carapace width (cw) (Figure 8). Only two juvenile (20—30 mm) crabs were

caught, this being in February. These two individuals undoubtedly belonged to

SA

RA

TOG

AP

AS

SA

GE

Map

so

fS

ara

tog

aP

ass

ag

esh

ow

ing

the

ap

pro

xim

ate

beam

tra

wl

Du

ng

en

ess

cra

ban

dsh

rim

pca

tch

es

at

ea

chsa

mp

lesta

tio

nb

yse

aso

n.

Fig

ure

7.

13

20 - Saratoga Passage% Male = 62 February% Female 38

% Soft = 0

Gravid Females = 1/2

N=810

5>~0

ci) I, ___ _______________ ___

D ~ — i i i i I --—r0~ci)

U10- June

% Male = 24I.

% Female = 76

- % Soft = 12

Gravid Females = 1/24

N = 33

5-

2.5

0 I I I I I I —

0 20 40 60 80 100 120 140 160

Carapace Width(mm)

Figure 8. Dungeness crab size—frequency histograms by season for crabscaught in Saratoga Passage.

14

the 1985 year group. Soft crabs (indicative of recent molting) were only

found in June (12%) and only one gravid female was caught during each season.

Occasional rock crabs (Cancer productus and C. gracilis) were also caught

by beam trawl in Saratoga Passage. Once again, these crabs were all caught at

the inshore stations of Transect 1.

Port Gardner. Up to 65 stations in Port Gardner were sampled by beam

trawl in February (n = 56), April (n = 55), June (n = 55) and September (n

65) 1986. The overall average numbers of Dungeness crab caught/ha (÷ 1

standard deviation) in Port Gardner and within each of the preliminary

disposal sites (n = 5 in each case) for each season were (Figure 9):

Average # Crab/ha + 1 Standard Deviation

Season Port Gardner NAVY PSDDA 2 PSDDA 1

February 126 ± 150 225 ± 98 6 ± 11 0 ± 0

April 85±127 588±141 19+19 0±0

June 114 ± 178 502 ± 105 19 ± 52 0 ± 0

September 100 ± 119 76 ± 51 11 ÷ 0 25 ± 29

Average 106.2 ÷ 145.5 297.8 ÷ 105.3 15.8 ± 19.4 6.2 ÷ 14.5

Dungeness crab were also sampled by otter trawl (incidental to fish

catches) at selected stations in Port Gardner and at each disposal site (n

3) during each season. Crab catches by otter trawl were usually less than the

beam trawl (based on equivalent area trawled) and substantially less at the

NAVY disposal site (Figure 9). The average numbers (+ 1 standard deviation)

15

0

NAVY DISPOSALSITE

BEAMTRAWL _____

SITE 2 SITE 1

Figure 9. Comparative average densities of Dungeness crab atthe Navy Disposal Site and the two control sites inPort Gardner by season and by trawl type.

DUNGENESS CRAB

400

OTTERTRAWL______

300

200

100

0

400

300

200

100

0

400

300

200

100

0

400

300

200

100

uJ

C.)ILlI

LU0~

>-

C’,zUi

UiCD

Ui>

PSDDA PSDDA

16

of Dungeness crab caught/ha at the three disposal sites for each season were:

Average # Crab/ha ± 1 Standard Deviation

Season NAVY PSDDA 2 PSDDA 1

February 21 ± 10 0 ± 0 2 ± 3

April 15÷6 0±0 0±0

June 9±9 1±2 0±0

September 2 ± 3 20 ± 7 12 ± 4

Average 11.8 ± 7.5 5.2 ± 3.6 3.5 ± 2.5

Dungeness crab catches for the beam trawl in Port Gardner are summarized in

Appendix Tables 8-12. The otter trawl crab catches are summarized in Appendix

Tables 14, 15 and 17.

Based on the beam trawl sampling efforts, the average annual abundance of

Dungeness crabs in Port Gardner was estimated to be 106 crab/ha within our

sampling grid. The estimated relative abundances at PSDDA Sites 1 (= Control

1) and 3 (= Control 2) were only 13% and 6%, respectively, of this average

annual abundance for Port Gardner.

The distributions of male and female crabs illustrated in Figures 10 and

11 show that males were relatively scarce. Typically the males accounted for

only 10% of the total crab catch and were generally found at the shallower

stations (averge depth of capture for males 29 m). Female crabs were much

more plentiful, were found in abundance at deeper depths (down to~—’100 m;

average capture depth = 63 m), and especially preferred the “nearshore slope”

area of Port Gardner instead of the deep, flat areas in the middle of the bay.

Dis

trib

utio

no

fm

ale

Du

ng

en

ess

cra

bs

cau

gh

tb

ybe

amtr

aw

lin

Po

rtG

ard

ne

rd

urin

gth

ese

aso

na

lsa

mp

ling

in1

98

6.

—4

Fig

ure

10.

FE

MA

LEC

RA

B

PS

OD

A1

1

Po

rt-~

.~

Ga

rdn

er

o0

0•

~.~

150

25

0

~1

” ••-.••

••.f

lo••

•.•

•••~

:°

••~

.

•‘o

•

•0

•~•

psp

P~

-~’

•to

•.~

it0

•~~

•—

AP

RIL

Po

rtG

ard

ne

r

PS

DD

A1

xxi

NO

RT

H

•

.

JUN

E

c$•~-”~

0

NO

RT

H

SE

PT

EM

BE

R0

0

Po

rtG

ard

ne

ro

PS

OD

A1

0

0

Fig

ure

11

.D

istr

ibu

tio

no

ffe

ma

leD

un

ge

ne

sscra

bs

flau

gh

tb

ybe

amtr

aw

lin

Po

rtG

ard

ne

rd

urin

gth

ese

aso

na

lsa

mp

ling

in1

98

6.

19

This pattern is illustrated in greater detail in Figure 12 which shows that

the highest abundances of Dungeness crab occur above 100 m depth. Rarely were

crabs found at stations in the middle of the bay except during September when

crabs may have “spread out” to deeper areas while foraging for food.

The average carapace width of all Dungeness crabs caught in Port Gardner

was 125 ± 13 mm with little difference between males (132 ÷ 21 mm) and females

(124 ± 11 nun). Histograms of carapace width—frequencies (Figure 13) show,

however, that the size range for male crabs was greater (80-180 mm) than that

for the females (100—150 mm) and that crabs less than about 2 years of age

were not caught in the trawls.

Size—frequency histograms (Figure 14) for male crabs by season show a

gradual increase in average size from 131 mm in February to 139 mm in

September with no recruitment of young crabs (i.e., < 100 m ow) after April.

These histograms also suggest that larger legal—sized (> 160 mm) crabs

disappear from the population, probably due to removal by the commercial/sport

fishery.

Female size—frequency histograms (Figure is) show a pattern different

than the males. Essentially no growth is evident based on the seasonal

average sizes (123—126 mm ow), but close inspection of the histograms

indicates that growth of the smaller females (ow 100-130 mm) is occuring but

that this growth is counteracted by a decline in the proportion of females >

130 mm OW, possibly due to natural mortality or emigration to areas outside

our sampling grid.

Elliott Bay. Only four Dungeness crab were caught by beam trawl in

Elliott Bay during all three sample seasons. Hone were caught by the otter

trawl. Two Dungeness crabs (both mature, non—gravid females) were caught in

20

ci)0)

I)>

I.

C)

I

ct3

(-)

Figure 12. Distribution by depth of all Dungeness crabs caught bybeam trawl in Port Gardner during seasonal sampling in1986.

CRAB—PORT GARDNER

200

100

a

FEB 1986

20

300

40 80 100 110 115 120 130 135 140 145 165

200

100

0

APRIL

10 20 40 80 100 1 115 120

300

200

100

JUNE

300

80 90 100 105 110 115 20 130 135 140 145 165

200

100

0

SEPT

10 20 40

Depth (meters)

PORT

GARD

NER

DUNG

ENES

SCR

ARW

IDTH

S

MA

LE6

~z1

32

PERC

ENT~ ~

020

4060

8010

012

014

016

018

0

CARA

PACE

WID

TH(m

m)

9

FE

MA

LE6

~z1

24

PERC

ENT~ 0

iiiill!illill

iiill!

Iiiiiiiiill

i1

11

11

IIiiii

iiit

i::

::::

::::

~tiii

ill

11

11

11

11

II

020

4060

8010

012

014

016

018

0

CARA

PACE

WID

TH(m

m)

Fig

ure

13

.C

ara

pa

cew

idth

—fr

eq

ue

ncy

his

tog

ram

so

fa

llm

ale

and

fem

ale

Du

ng

en

ess

cra

bs

cau

gh

tb

ybe

amtr

aw

lin

Po

rtG

ard

ne

rd

urin

gse

aso

na

lsa

mp

ling

in1

98

6.

10

5

050 60

Carapace width—frequencyDungeness crab caught byduring seasonal sampling in 1986.

140 150 160 170 180

PERCENTFEBx:131

22

MALE OUNGENESS CRAB SIZES

iLIiijImI...iIa70 80 90 100 110 120

APRILX : 127

130 140 150 160 170 180

II_I I ~1.i, .i

C

liiiUI

10

PERCENT s

0

10

PERCENT 5

a

10

PERCENT s

a

0 60 70 80 90 100 110

JUNE)( :134

1..1.1.1

120 130 140 150 160 170 180

LII.IiIIii. [.1100 110 120 13060 70 80 90

SEPT~:139

50 60 70

Figure 14.

80 90 100 110CARAPACE

IL+÷120 130 140 150

WIDTH (mm)160 170 180

histograms for all malebeam trawl in Port Gardner

23

10~FEMALE OUNGENESS CRAB SIZES

10

FEB~:125

tmIt1Il1t1,JIHIIM~p

60 70 80 90 100

I

110 120 130

I—

— liii

PERCENT sAPRIL)( :126

10~

PERCENT 5

JUNEX :~23

SEPT~:123

Ittilil!

Figure 15.

120 130 140 150 160 170 180WIOTH (mm)

Carapace width—frequency histograms for all female~ungeness crabs caught by beam trawl in Port Gardnerduring seasonal sampling in 1986.

PERCENT 5 I I

140 150 160 170 180

0 60 70 80 90 100-I

110 120 130 140 1 5~0

10

PERCENT 5

n

160 170 180

50 60 70I ~1

L ......

18080 90 100 110 120 130 140 150 160 170

II

II

IiLWjJj1~j1b1I IlIItIttIIItIt

60 70 80 90 100 110

CARAPACE

24

June at the 10 and 20 m stations of Transect 1 off Duwamish Head (Figure 16).

Two mature males were caught in September, also at the 10 m station of

Transect 1. Occasional rock crabs were also caught at the shallower stations

of Transects 1 and 2.

Commencement Bay. No Dungeness crab were caught in Commencement Bay by

either trawl gear. Occasional rock crabs were again caught at the shallower

stations (10—40 m) in all three seasons.

Shrimp

Catch data were collected on six species of Pandalid shrimp of commercial

or sport value. These six species were: Spot prawn, Pandalus platyceros;

Coonstripe shrimp, P. danae; Humpback shrimp, P. hypsinotus; Flexed shrimp, P.

goniurus; Pink shrimp, P. borealis; Smooth pink shrimp, P. jordani; and

sidestripe shrimp, Pandalopsis dispar (Butler 1980).

Saratoga Passage. The overall average number of shrimp caught by the

beam trawl in Saratoga Passage in February and June was essentially the same

at 51.9 and 56.2 shrimp/ha, respectively. The largest shrimp catch (300

shrimp/ha) was at the 80 m station on the Whidbey Island side of Transect 1 in

June, followed by the 100 m station (243 shrimp/ha) on Transect 2, also in

June (Figure 7). The two most abundant shrimp species were the pink and

smooth pink shrimps (Appendix Table 6).

The average shrimp catch within the preliminary PSDDA disposal site was

46.8 and 68.7 shrimp/ha for February and June, respectively. Sidestripe

shrimp were the most abundant shrimp species caught by the beam trawl in the

disposal site in February, while pink shrimp were the most common species in

EL

LIO

TT

BA

Y

Fig

ure

16

.M

aps

of

Ellio

ttB

aysh

ow

ing

se

aso

na

lbe

amtr

aw

lca

tch

es

of

Du

ng

en

ess

cra

ban

dsh

rim

pa

tth

esa

mp

ling

sta

tio

ns

inE

llio

ttB

ay.

U,

26

June (Figure 9; Appendix Table 6). However, the otter trawl in June caught

more sidestripe shrimp than pink shrimp and caught a total of 126.2 shrimp/ha

for all species combined (Figure 17; Appendix Table 7).

Port Gardner. The overall average number of shrimp/ha (÷ 1 standard

deviation; all species combined) caught by the beam and otter trawls in the 3

preliminary Port Gardner disposal sites during each season of 1986 were (n = 3

in each case):

Average # Shrimp/ha ± 1 Standard Deviation

Season NAVY PSDDA 2 PSDDA 1

Beam Trawl:

February 687 ± 518 81 ± 11 0 ± 0

April 0+0 12+11 56+19

June 8+13 0±0 6±11

September 293 ± 249 6 ÷ 11 31 ± 11

Average 446.9 + 381.6 25.0 ÷ 35.1 23.4 ± 25.4

Otter Trawl:

February 188 + 170 354 ÷ 184 135 ± 43

April 113±21 52±21 30±5

June 5±5 117±149 80±44

September 443 ± 81 86 ± 20 101 ± 18

Average 186.9 ± 175.0 147.1 ± 164.2 86.3 ÷ 48.4

27

Figure 17. Beam and otter trawl shrimp catches by species and byseason for PSDDA Site 2 in Saratoga Passage. The barsare the average catches for the thred stations withinthe Disposal Site. N.S. = not sampled.

SARATOGA PASSAGE

Ucc

0UizI0

00~

ccI(I)

10080604020

0

1601208040

01601208040

0

400300200100

0

20151050

400

300

200

100

0

BeamTrawl

Li0 He r

Trawl

. Spot Prawn

. 0 0 0

. Sidestripe

:_

. Smooth Pink

0 0. Pink

— — II

. Humpback

~ 0 0 0All Shrimp

BNS •fl NSNS

FEB JUNE

1986

SEPT

28

For each gear type, the pattern of relative shrimp densities between the three

disposal sites (based on annual averages) was NAVY > PSODA 2 > PSDDA 1.

Plots of overañ shrimp abundances in Port Gardner by season (Figure 18)

show that the nearshore slope area (including the NAVY site) was the most

important area for shrimp with relatively few shrimp occurring at the deeper

stations in the middle of the bay. Figure 18 also shows that there was a

distinct seasonality in general shrimp abundances with the highest densities

being present during the fall-winter period. The fate of these shrimp during

spring and summer is presently unknown.

Plots of shrimp distributions by depth (Figure 19) reinforce the finding

that the nearshore slope area provides the most important habitat. The great

majority of all shrimp caught by beam trawl in 1986 were found at stations

between about 20 and 100 m depth with the exception of April when very few

shrimp were caught (Figure 19).

The shrimp species most commonly caught within the three disposal sites

in Port Gardner varied with site, season and trawl gear (Figure 20).

Generally, sidestripe and pink shrimp were the most abundant at the disposal

sites with the smooth pink shrimp being fairly abundant at the NAVY site.

Coonstripe shrimp were never caught at these relatively deep stations and

humpback and flexed shrimp were scarce. The reasons for the sometimes extreme

differences in shrimp density estimates between the two trawl gears are

presently unknown and may be, in part, species-dependent. The Port Gardner

shrimp catch data for both trawls are summarized in Appendix Tables 13, 14 and

16—18.

Elliott Bay. The overall average beam trawl catches of shrimp in Elliott

SH

RIM

P—

PO

RT

GA

RD

NE

R

Fig

ure

18,

Dis

trib

utio

no

fsh

rim

p(a

llsp

ecie

sco

mb

ine

d)

cau

gh

tb

ybe

amtr

aw

lin

Po

rtG

ard

ne

rd

urin

gse

aso

na

lsa

mp

ling

in1

98

6.

500400300200100

0

600500400300200100

0~

600L. 500

400l)~3 300

II-~ 600

500400300200100

0

Depth (meters)

Figure 19, Distribution by depth of shrimp (all species combined)caught by beam trawl in Port Gardner during seasonalsampling in 1986.

30

SHRIMP—PORT GARDNER

FEB 1986

10 20 40 80 100 110 115 120 130 135 140 145 165

APRIL

10 20 40 80 100 110 115 120 130 135 140 145 165

JUNE

823

10 20 40 80 90 100 105 110 115 120 130 135 140 145 165

SEPT

10 20 40 80 90 100 105 110 115 120 130 135 140 145 165

PO

RT

GA

RD

NE

R

w I () LU I I C.)

C.)

0~ cc I C,)

20PS

DD

A2

Sp

ot

Pra

wn

10

00

00

00

400

Sid

estr

ip.

30

02

00 0

—O

fl

-~

100

soo

Sm

oo

thP

ink

60

04

00

20

0A

LL

O0

20

0P

ink

150~

100 0

—ri

0r—

~0

,0

r—~

8H

um

pb

ack

6 4 2A

LL

00 8

Fle

xe

d6 4 2 0

01

10

11

00

00

All

Sh

rim

p8

00

60

0

40

0

20

0 0—

—pii

—

PSD

DA

1S

po

tP

raw

n

5.

AL

LO

0S

ide

str

ipe

0r1

0—

_r—

ir1

Sm

oo

thP

ink

AL

LO

CP

ink

O0

r—~

——

08

Hu

mp

ba

ck

AL

L0

8F

lexe

d6 4

.

co

Fi

00

00

All

Sh

rim

p

o~

ri—

——

FEB

AP

RIL

JUN

ES

EP

T

Num

ber

of

shrim

pca

ug

ht

pe

rh

ecta

reb

yb

oth

beam

and

ott

er

tra

wis

inth

eth

ree

pre

lim

ina

ryd

isp

osa

lsite

sin

Po

rtG

ard

ne

rd

urin

gse

aso

na

lsa

mp

ling

in1

98

6.

20 15 10

400

30

020

010

0 0

80

06

00

40

02

00

20

0

150

100

FEB

AP

RIL

JUN

ES

EP

T

80

0

60

0

40

0

20

0

FEB

AP

RIL

JUN

ES

EP

T

Bea

mT

raw

l[~

]Ott

er

Tra

wl

Fig

ure

20

.

32

Bay during February, June and September were 131, 66 and 135 shrimp/ha,

respectively. Generally, shrimp were most abundant in the beam trawis at

PSDDA Site 1 (inner Elliott Bay) in February; most abundant at PSDDA Site 2

(Fourmile Rock) in June; and substantially more abundant at PSDDA Site 1 in

September (Figures 8 and 10; Appendix Table 6) Sidestripe and pink shrimp

were the most abundant shrimp in the beam trawls at the Fourmile Rock site

while pink and smooth pink shrimp were most common at the inner bay disposal

site (Figure 21). The otter trawl catches generally showed the same pattern

of shrimp distribution except that spot prawn catches were higher than for the

beam trawl at the inner bay site in September and a few humpback shrimp were

caught in June and September (Figure 21; Appendix Table 7).

A “t”—test was conducted to compare the mean catches of shrimp between

the two Elliott Bay disposal sites (data from all seasons combined; Log~Q

transformation of catches). The results showed that shrimp were caught in

significantly higher numbers (p = 0.0009) when data from both types of trawl

gear were combined, but that the level of significance was marginal for each

gear type alone (p = 0.054 for the beam trawl shrimp catches and p = 0.050 for

the otter trawl catches).

Commencement Bay. The overall average beam trawl shrimp catches in

Commencement Bay in February, June and September 1986 were 49, 29 and 128

shrimp/ha, respectively. The two largest shrimp catches in Commencement Bay

were off Browns Point in September where coonstripe shrimp were plentiful

(1,067/ha) at the 10 m station and pink shrimp were relatively abundant

(502/ha) at the 80 m station (Figure 22; Appendix Table 6).

Shrimp catches were very low at the PSDDA 2 site which was only sampled

in February (Figure 23). Beam trawl and otter trawl shrimp catches at PSDDA

Sites 1 and 2B were almost identical in both June and September with pink and

EL

LIO

TT

BA

Y

Fig

ure

21

.B

eam

and

ott

er

tra

wl

shrim

pca

tch

es

by

site

,b

ysp

ecie

san

db

yp

rop

ose

dd

isp

osa

lsite

s.

The

ba

rsa

rea

vera

ge

ca

tch

es

for

the

ea

chd

isp

osa

lsite

.N

.S.

=n

ot

sam

ple

d.

sea

son

for

the

two

Ellio

ttB

ayth

ree

tofive

sta

tio

ns

with

in

FOU

RM

ILE

RO

CK

INN

ER

ELL

IOTT

BAY

Ui cr 1— 0 UiI I 0 I 0 I C

l)

100

100

80S

po

tP

raw

n80

60-

6040

4020

200

00

—0

016

0.

Sid

ea

trip

e16

012

0-

120

8080

40-

400

Br—

i0

160

.S

mo

oth

Pin

k16

012

0-

120

80.

8040

-40

00

00

0o

400

Pin

k40

03

00

30

020

020

010

010

00

—--

0

20H

um

pb

ack

2015

1510

-10

5-

.50

00

00

0A

llS

hrim

p40

040

0

300

300

200

200

io:-

•N

SIn

10:

Be

am

Tra

wl

LI Ott

er

Tra

wl

Sp

ot

Pra

wn

~o

liii

.S

ido

str

ip.

—0

~—

,

,S

mo

oth

Pin

k

•1

—-

.P

ink

j~1

~~

~iiI

riH

um

pb

ack

ao

no

fl

All

Sh

rim

p89

1

LN

SL

LFE

BJU

NE

SEPT

FEB

JUN

E

1986

1986

SEPT

34

COMMENCEMENT BAY

Figure 22. Maps of Commencement Bayshowing beam trawl catches of shrimpat the sampling stations inCommencement Bay.

CO

MM

EN

CE

ME

NT

BA

Y 100

60 60 40 20 0

160

120

80 40 016

012

08

040

040

03

00

200

100 0 20 15 10 5 0

40

0

30

0

200

100 0

Fig

ure

23

.B

eam

and

ott

er

tra

wl

shrim

psite

sin

Com

men

cem

ent

Ba

y.w

ith

ine

ach

dis

po

sa

lsite

.

ca

tch

es

by

site

,b

ysp

ecie

san

db

yse

aso

nfo

rth

ep

rop

ose

dd

isp

osa

lT

heb

ars

are

the

ave

rag

eca

tch

es

for

the

thre

eto

fou

rsta

tio

ns

N.S

.=

no

tsa

mp

led

.

PS

DD

A1

PS

DD

A2

PS

DD

A2B

LU cr 0 LU z z 0 I— () a cc z C’)

100 80 60 40 20 0

160

120

80 40 016

012

08

0 40 0

40

030

020

010

0 0

20 15 10 5 0

40

0

300

200

100 0

100

80 60 40 20 0

160

120

80 40 016

012

08

040 0

400

30

020

010

0 0

20 15 10 5 0

40

0

Sp

ot

Pra

wn

pS

ld.s

trip

.

Sm

oo

thP

ink

0

Pin

k

Hu

mp

ba

ck

0

All

Sh

rim

p

•M

SM

SN

SN

SN

5

FEB

JUNE

SEPT

1986

S Bea

mT

raw

l

Ott

er

Tra

wl

.S

po

tP

raw

n

.0

0.

Sid

.str

lp.

.0

~.

Sm

oo

thP

ink

~0

00

0~

Pin

k

Hu

mp

ba

ck

00

00

All

Sh

rim

p

~N

SN

SI.

_

300

200

FEB

JUNE

1986

100 0

SEPT

FEB

JUNE

1986

SEPT

36

sidestripe shrimp being the dominant species (Figure 23; Appendix Table 5 and

6). “T”—tests conducted to compare the mean shrimp catches (data from all

seasons combined; Log10 transformation of catches) between PSDDA Sites 1 and

2B showed that there were no significant differences (p = 0.05) in mean

catches regardless of trawl type.

Shrimp distributions by depth. The combination of all shrimp data from

all seasons and areas (except Port Gardner) shows that the different species

have specific depth preferences. Coonstripe shrimp preferred the shallowest

depths (<30 m), often being associated with eelgrass (Zostera marina) and

various algas (Figure 24). The mid-depths (50—100 m) were generally preferred

by spot prawns and pink, smooth pink and humpback shrimp. Sidestripe and some

pink shrimp were found at the deepest (100—iso m) depths.

Shrimp size distributions. Coonstripe shrimp was the smallest species

caught with carapaces lengths (CL) between about 9 to 12 mm and were generally

larger with increasing depth (Figure 25). Both species of pink shrimp were

small to moderate in size, averaging 13 to 18 mm CL with no trend in size with

depth. Sidestripe and humpback shrimp were moderately large in size (18 - 24

mm average CL), trending to smaller sizes with increasing depth. The largest

shrimp, the spot prawn, averaged 26 to 34 mm CL and also trended to smaller

sizes at depth.

Shrimp length-frequencies. Spot prawn were not caught in any of the

PSDDA sites in February but showed indications of a slightly bimodal length—

frequency during June and September with one size group from about 25 to 38 mm

and a second size group from about 40 to 45 mm (Figure 26). Sidestripe shrimp

37

60

40

20

0

60

40

20

0

(‘34-a0‘1)I 20

-~ 00

4-a

c~ 60C)

40

c~ 20

0 0>

60

40

20

0

300

200

100

0 125 150

Depth (m)

Figure 24. Distribution by depth and by species for all beam trawl—caught shrimp, all areas (except Port Gardner) and seasonscombined.

80

025 50 75 100

38

D)Ca)-J

a)C-)

a

C’,

0a)0)C’,

a)>

I I I I

0-15 16-30 31-70 71-95

Depth Range (m)

Figure 25, Average carapace lengths by species and by depth ranges for allshrimp caught, all seasons and areas (except Port Gardner)

35

30

25

20

15

10

5

p

96-120 >120

combined.

FEb

SE

pT

Ca

rap

ace

Leng

thF

igu

re2

6.

Le

ng

th—

fre

qu

en

cyh

isto

gra

ms

for

all

sp

ot

pra

wn

and

sid

estr

ipe

shrim

pca

ug

ht

du

rin

gth

eth

ree

sam

plin

gse

aso

ns,

all

are

as

(exce

pt

Po

rtG

ard

ne

r)co

mb

ine

d.

Sid

est

ripe

IIII

>~ ci ci) a) LL ci) > ci)

40-

Spo

tP

raw

n3

0-

20 10

~

0N

on

eC

au

gh

tI

•I

I‘

40

~

30

20JU

NE

10

~

0,

IIIII

1111

1

40

30

~

20

-

1~

.•

•IIiIW

kII~1

1Iilij

•I

~•

-I

IIi,Iih

11

..

~Ii.

II

II

010

20

30

40

0I

I

20

30

10 (mm

)

40

40

length-frequency patterns suggested a single size group in both February (16.-

24 mm) and June (20-30 mm) and a bimodal pattern in September (10—15 mm and

20—30 nun) (Figure 26). Coonstripe shrimp length—frequencies suggested a

single size group in February and June (8-18 nun) and recruitment of young

shrimp in September (6-10 mm) (Figure 27). Relatively few humpback shrimp

were caught, but those that were suggested a single size group with sizes

between 22—30 mm (Figure 27). Smooth pink shrimp length-frequencies gave a

suggestion of a slight bimodal size distribution with size groups from 10—14

mm and 17-20 mm in February with both groups growing progressively larger in

June and September (Figure 28). Pink shrimp also showed a bimodal size

distribution with size groups from 9-13 mm and 15-20 mm in February. The

distinction of the two apparent size groups was less clear in June and

September but there was a hint of new recruitment in September with shrimp

between 9—12 mm (Figure 28).

DISCUSSION AND CONCLUSIONS

Dungeness Crab

Dungeness crab were completely absent from all trawls conducted in

Commencement Bay in 1986. Although an occasional Dungeness crab has been

caught in other trawling operations in the shallow waterways (C. Eaton, pers.

comm.), it is clear that this species will not be a factor in siting a

disposal site in Commencement Bay.

Only four Dungeness crab were caught in Elliott Bay, all by beam trawl at

the shallow stations on the Duwamish Head Transect (Figure 16). Commercial

crabbing operations were also observed in shallow water areas between Fourmile

Rock and West Point. However, the scarcity of Dungeness crabs in the trawls

and the total lack of crabs in the trawls from the preliminary disposal sites

>~ C.)

C ci) D ci) U- ci) > ci) Ix

40

-

30

-

20 10 0

20 10 0

Hum

pbac

k

Non

eC

au

qh

t

Fig

ure

27

.L

en

gth

—fr

eq

ue

ncy

his

tog

ram

sfo

ra

llco

on

str

ipe

and

hum

pbac

ksa

mp

ling

sea

son

s,a

lla

rea

s(e

xce

pt

Po

rtG

ard

ne

r)co

mb

ine

d.

shrim

pca

ug

ht

du

rin

gth

eth

ree

Coo

nstr

ipe

III•—

r—•

•‘

40 30

20 10 0

40

30

I-

—-.

II

•~

FEb

JUN

E

SE

pT

Ca

rap

ace

010

203

04

00

1020

30

40

Leng

th(m

m)

0 c ci) ci) LI ci) >

__

__

__

__

__

__

__

_

ci) ci:

Ca

rap

ace

Leng

th(m

m)

Fig

ure

28

.L

en

gth

~-f

req

ue

ncy

his

tog

ram

sfo

ra

llsm

oo

thp

ink

and

pin

ksh

rim

pca

ug

ht

du

rin

gth

eth

ree

sa

mp

ling

sea

son

s,a

lla

rea

s(e

xce

pt

Po

rtG

ard

ne

r)co

mb

ine

d.

Sm

ooth

Pin

k

IId

Pin

k

II>~

I—

II

•I

40

-

30

-

20 10

-

0.

40 30

20 10 0

40

30

-

20

-

10 0

FEb

JUN

E

SEpT

I.1

fflui

h11.

I1~

I•

I•

I

010

2030

400

1020

30

40

43

indicates that this species is also not a factor in selecting a final Elliott

Bay disposal site location.

Dungeness crab were moderately abundant in the beam trawl catches from

Saratoga Passage (Figure 7); however, all crabs were caught along the

shoreward slope areas at depths <80 m. No crabs were caught in the

preliminary disposal area nor in the deeper water areas north of the disposal

site. Hence, the present location of the proposed disposal site is probably

in the best location for avoiding impacts to crab. However, evidence from the

trawls in Port Gardner suggests that some Dungeness crab move into deep water

(i.e., 100-150 m) during the late summer to early fall period. Trawls were

not made in Saratoga Pass during this period. Thus, care should be exercised

about any assumptions that crab are absent year-round.

Dungeness crab were found to be a very important resource in Port

Gardner, consistently averging about 100 crab/ha for all seasons sampled. Of

the crabs caught in the trawls, almost 90% were mature females, 78% of which

were gravid during the February sampling. Thus, Port Gardner appears to be an

important habitat area for the mature females.

The most important area of Port Gardner for the females is the nearshore

slope area with few crabs being found in the deeper mid-portion of the Bay

(Figures 9—12). Figure 9 shows that, unlike the NAVY disposal site, the two

preliminary PSDDA sites contain relatively few crabs. Of these two sites, the

PSDDA 1 site in the middle of the bay is farthest from the

nearshore crab aggregations. A possible exception to this rule may be during

summer—early fall when crabs appear to “spread out” into the deeper areas, but

still at densities far less than the “slope” area.

44

Shrimp

Commercially important species of shrimp were caught in all of the

preliminary PSDDA disposal sites. Summaries of the shrimp catches within the

disposal sites of each area (Table 1) show that the average shrimp catches by

weight for the combined otter trawl catches were 0.56, 0.06, 1.69 and 1.22

kg/ha for Saratoga Passage, Port Gardner, Elliott Bay and Commencement Bay,

respectively.

Historically, shrimp have been the basis of a viable trawl fishery in

Puget Sound and Hood Canal. Annual landings of shrimp exceeded 400,000 pounds

during several years between 1904 and 1915 and averaged about 50,000 pounds

during the 1920’s and 1950’s (Smith 1957). The averge landings between 1955

and 1982 have been highly variable (s to 144 thousand pounds) and averaged

58,000 pounds (Figure 29).

The historical shrimping grounds fished from the late 1800’s through the

1930’s included each of the areas in which a PSDDA disposal site is proposed

(Figure 30). Saratoga Passage and Elliott Bay are shown as historical spot

prawn shrimping areas while Commencement Bay was trawled for smooth pink

shrimp. Our present trawls caught spot prawn in Elliott and Commencement Bays

but no spot prawn in Saratoga Passage. Relatively few smooth pink shrimp were

caught but the closely related pink shrimp was caught in small numbers in

Saratoga Passage and in moderate numbers in Elliott and Commencement Bays

(Table 1).

Some perspective on the relative importance of shrimp resources in the

preliminary disposal sites can be attained by comparing the average otter

trawl catches in these sites with otter trawl catches of shrimp in other areas

of Puget Sound and Hood Canal. Chew (unpublished data) conducted shrimp

surveys during the winter of each year from 1967 to 1979 at about ten sites in

45

Table 1. Average shrimp catches, lengths and weights (wet biomass) for all shrimpcaught by otter trawl in the proposed PSDDA disposal sites in SaratogaPassage, Port Gardner, Elliott Bay and Commencement Bay during allsample months (combined), 1986.

PortSpecies Saratoga Passage Gardner Elliott. Bay Commencement Bay

Spot Prawn

Ave. #/Ha 0 0.8 23.2 1.5Ave. carapace length (mm) — 19.0 33.6 26.8Ave. weight/shrimp (g) 0 5.0 23.0 12.0Total weight/Ha (kg) 0 0.00 0.53 0.02

Sidestripe

Ave. lI/Ha 54.1 6.2 23.2 53.3Ave. carapace length (mm) 23.2 18.0 23.0 15.3Ave. weight/shrimp (g) 6.2 5.0 6.0 1.9Total weight/Ha (kg) 0.33 0.02 0.14 0.10

Smooth Pink

Ave. lI/Ha 0 0 23.9 0.8Ave. carapace length (mm) — — 16.9 16.5Ave. weight/shrimp (g) 0 0 3.4 3.1Total weight/Ha (kg) 0 0 0.08 0.00

Pink

Ave. lI/Ha 72.1 17.2 260.6 306.4Ave. carapace length (mm) 16.5 14.4 16.8 17.2Ave. weight/shrimp (g) 3.2 2.5 3.5 3.6Total weight/Ha (kg) 0,23 0.04 0.91 1.10

Humpback

Ave. lI/Ha 0 0 2,4 0Ave. carapace length (mm) — — 26.4 —

Ave. weight/shrimp (g) 0 0 12.0 0Total weight/Ha (kg) 0 0 0.03 0

All Species Combined

126.2 24.2 333.3 362.0Ave. lIHa

Total weight/Ha (kg) 0,56 0.06 1.69 1.22

THOU

SAND

POUN

DS

PUCE

TSO

UND

SHRI

MP

LAND

INOS

YEAR

Fig

ure

29

.

I 140

120

100 80.

60

3540

5560

6570

7580

An

nu

al

com

me

rcia

lsh

rim

pla

nd

ing

fro

mP

ug

et

Sou

nd(in

clu

din

gH

ood

Ca

na

l)fr

om

1935

to1

98

2.

Da

tafr

om

WD

F(1

97

4an

d1

98

2).

47

_______ Spot

Figure 30. Map of Western Washington showing areas of commercial shrimpproduction from late 1800ts to mid—1930!s. The Humpback shrimp

VANCOUVER I5LAND

CLALLA~1

J~rrER5oN

MA5ON

GRAV5HARBORCOUNTY COUNTY

Humpback [II I I I I I~ Smooth Pink

is Pandalus hypsinotus. From Smith (1937).

48

Hood Canal and Puget Sound. Summaries of Chew’s data show that the average

shrimp catches/ha in four areas of Hood Canal and three areas of Puget Sound

all (except Seabeck, Hood Canal) exceeded the average catches in any of the

preliminary PSDDA disposal sites (Tables 1 and 2). The one disposal site that

appeared to have a potential for commercial shrimp harvesting was the inner

Elliott Bay site where spot prawn, sidestripe, pink and smooth pink shrimp

were caught in reasonable numbers (Figure 10, Appendix Table 7). Table 3

provides a breakdown of relative shrimp densities within the two preliminary

PSDDA sites in Elliott Bay and shows that the inner Elliott Bay site contained

about 3 to 7 times the density of shrimp that were caught at the Four—mile

Rock site (data from June and September otter trawis). However, this area is

also severely impacted by Indian salmon fishing, ship navigation lanes and

anchorage areas as well as toxic contaminants in the sediments of the nearby

Duwamish Waterways; hence, the value of these shrimp to a fishery is suspect.

The potential value of the reproductive capacity of these stocks for supplying

new recruits to other productive areas of Puget Sound is not presently known

but cannot be ruled out in the decision making process. The Commencement Bay

disposal sites also contained some sidestripe and pink shrimp, but both sites

contained essentially equal populations, hence, not affording a choice between

these two sites based on this factor.

49

Table 2. Estimated average shrimp catches/Ha from otter trawisconducted in selected areas of Hood Canal and Puget Soundfrom 1967 to 1979. These estimates are derived fromunpublished data collected and summarized by Dr. KennethChew, School of Fisheries, University of Washington.

Location/Depth (m) Number of trawis Catch/Ha (kg)

Dabob Bay

20—45 33 2.945—70 26 2.770 — 125 24 3.5

Pleasant Harbor

35-65 5 2.965-90 8 10.0

Seabeck

45-SO 3 0.8

Potlatch

70-90 4 6.8

Port Susan

25 — 70 9 12.880 — 120 7 5.7

Tulalip

50—80 3 13.580- 120 4 11.8

Carr Inlet

45 — 80 4 15.180 — 135 3 2.4

50

Table 3. Shrimp weights/ha from the Elliott Bay preliminary disposal sites asestimated from the otter trawl catches in June and September 1986.Shrimp weights for each species were calculated from length—weightregressions developed from data collected by K. Chew (unpublished).

Estimated Total Weight (Kg)/ha

• June SeptemberSpecies Four—Mile Rock Inner Elliott Four—Mile Rock Inner Elliott

Spot Prawn 0.016 0.210 0.107 1.641

Sidestripe Shrimp 0,108 0.060 0.388 0.064

Smooth Pink Shrimp 0 0,362 0 0.010

Pink Shrimp 0.265 0.470 0.141 3.004

Humpback Shrimp 0 0.034 0 0.064

Total Weight 0.389 1.136 0.636 4.783

51

Part II

Demersal Fish Studies

by

Robert F. Donnelly, Bruce S. Miller, Robert R. Lauth, and Shelley C. Clarke

INTRODUCTION

This study investigated fish assemblages at preliminary Puget Sound

Dredge Disposal Analysis (PSDDA) disposal sites in the main basin of Puget

Sound and evaluates these assemblages prior to actual disposal of dredged

materials. Information obtained will be used in the final site selection

process and can be used as baseline data to monitor changes in fish

assemblages following disposal activities.

Disposal of dredged materials can affect fish habitats in many ways.

Sediment type has been shown to be particularly important for spawning (Morton

1977). Alteration of substrate particle composition by dredged materials and

consequent alteration of spawning grounds could be detrimental to the

abundance of certain fish species.

Dredged materials can alter the species composition of fish at

disposal sites by causing changes in the benthic community upon which the fish

feed (Lunz and Kendall 1982). A Rhode Island dredged materials disposal study

(Saila et al. 1972) suggests that covering the bottom with a uniform sediment

type would decrease the diversity of prey organisms and possibly decrease the

diversity of fish species. Desbruyeres et al. [1980, in Thistle (1981)] found

five times greater benthic faunal density six months after a disturbance at

2160 m; however, the fauna in the disturbed area was taxonomically different

from the surrounding fauna. At a deepwater disposal site in Puget Sound,

Bingham (1978) showed a similar effect. Nine months after disposal, diversity

52

of prey organisms was greater at the disposal site than at reference areas.

A disturbance caused by an oil spill in shallow water actually resulted in a

biomass increase six months to one year after the spill (Orensanz and Gallucci

1982). The work of Grassle (1977), however, cautions that the recovery after

disturbance may be depth dependent. Grassle’s study found that a deepsea site

(1760 m deep) had a colonization rate two orders of magnitude lower than a

comparable intertidal site.

Although it is important to be aware of the potential changes, it is

difficult to accurately predict what impact the disposal of dredged materials

will have on fish assemblages due to the individual nature of each disposal

site. Therefore, it is important to identify areas where fish resource

conservation is essential from a commercial or ecological perspective before a

decision is made regarding disposal. This report documents the benthic fish

assemblages of the preliminary main basin PSSDA disposal sites and adjacent

reference areas, and can aid in selection of sites where disposal of dredged

materials will have a minimal impact.

MATERIALS AND METHODS

Bottomfish (benthic and demersal fishes) were sampled in Commencement

Bay, Elliott Bay, Saratoga Passage and Port Gardner during 1986. Commencement

Bay was sampled on June 13 and September 8; Elliott Bay was sampled on July 3

and September 9; and Saratoga Passage was sampled once on July 1. Port Gardner

was sampled during four seasons on February 12 and 13, April 18 and 21, June

30 and July 2, and September 11 and 15. Marine environmental data (salinity,

dissolved oxygen, water temperature and water clarity) were also collected.

A 7.6—rn, single wire otter trawl (Mearns and Allen 1978) was the primary

53

sampling gear for bottomfish. The body of the net was made of 3.5 mm stretch

mesh and the cod end of 0.5 cm stretch mesh covered with 2.5 mm stretch mesh

to prevent chafing. The net was deployed from the 16—m research vessel

Kittiwake. The effective fishing width of the otter trawl was 6 m. Each

sample consisted of one otter trawl haul towed for a distance of 370 m at a

target ground speed of 4.2 km per hour. The total area swept (sampled) was

2,220 m2. Fish were also collected incidentally by the beam trawl used to

sample crabs (see Appendix A). The beam trawl is described elsewhere in the

crab and shrimp section (Part I) of this report.

Sampling was conducted both inside and outside of each preliminary PSDDA

site (Figures 1—4). Three replicate samples were collected inside each PSDDA

site and NAVY site (Port Gardner only) during each sampling cruise. One

sample was taken from each station outside of the PSDDA sites and at each of

the U.S. Army Corps of Engineerst established reference stations (Clarke 1986)

during each cruise. Reference stations were not sampled in Commencement Bay.

PSDDA sites and reference stations were the only locations sampled in Elliott

Bay (Fig. 2). In Commencement Bay, Saratoga Passage and Port Gardner the

PSDDA site(s), the reference station(s) and several additional stations

stratified by depth were sampled.

Each trawl catch was brought onboard and fish were sorted by species and

life history stage (adult or juvenile), counted and recorded; miscellaneous

observations (e.g., spawning condition) were also recorded. The catch was

then placed into plastic bags, labeled, put into ice chests and covered with

ice. The samples were transported to the University of Washington and placed

into a 0°C freezer until processed.

Surface water temperature, salinity and dissolved oxygen samples were

taken from a bucket of water collected from the surface waters. Bottom water

54

Figure 1. Map of Commencement Bay showing locations sampled forbottomfish (~) on June 13 (summer) and September 8(autumn). The large area enclosed by the dashed lineis the zone of siting feasibility (zSF). Circular areasenclosed by solid lines are the preliminary PSDDA sites.

55

Figure 2. Map of Elliott Bay showing locations sampledon July 3 (summer) and September 9 (autumn).by the dashed lines are the ZSFs. The solidpreliminary PSDDA sites.

for bottomfish (~)The areas enclosed

lines enclose the

56

Figure 3. Map of Saratoga Passage showing locations sampled for bottom—fish (~) on July 1 (sunimer). The area enclosed by thedashed line is the ZSF. The rectangular area enclosed bythe solid line is the PSDDA site.

Fig

ure

4~M

apo

fP

ort

Gar

dner

show

ing

the

sta

tio

nsa

mpl

edfo

rb

ott

om

fish

(S).

The

irre

gu

lar

dash

edlin

ere

pres

ents

the

20ni

con

tou

r.Th

ere

cta

ng

ula

rar

eas

en

clo

sed

byda

shed

line

sar

ePS

DDA

site

san

dth

ecircu

lar

are

ae

nclo

se

db

ya

cla

sh

ed

lin

eis

bh

eN

AV

Ysib

e.

Sta

bio

ns

Gan

dH

are

refe

ren

cest

atio

ns

1an

d2,

resp

ect

ive

ly.

Tl,

T2,

and

T~4

are

tra

nse

cts

1,

2,

an

d~

,re

sp

ective

ly.

U,

58

temperature, salinity and dissolved oxygen samples were taken from water

collected approximately I m above the bottom with a Scott—Richards water

bottle. Water temperatures were measured with a hand—held thermometer and

recorded in the field. Water samples for salinity determination were placed

into bottles for later measurement in the laboratory. Water samples for

dissolved oxygen determination were placed into acid washed glass bottles,

fixative added, the bottles glass stoppered, and the contents later processed

in the laboratory. Water clarity measurements were taken with a Secchi disc

from the lee side of the vessel and recorded in the field.

Laboratory Processing of Fish

Fish samples were removed from the freezer and thawed. The contents were

separated by species and life history stage. Total length (mm) of each fish

and total weight (grams) of each life history stage were recorded on data

forms and then entered into electronic storage. Since the tips of ratfish

tails were often missing, a length from the snout to the posterior end of the

second dorsal fin, as well as total length (when possible), was recorded for

this species. Adult flatfish and ratfish were sexed.

Flatfish Diseases

Marine flatfishes in Puget Sound are known to be infected by blood worms

(the nematode,Philometra), skin tumors, liver tumors and fin erosion (Amish

1976; Angell et al. 1975; Miller and Wellings 1971; Wellings et al. 1976;

Malins et al. 1982).

Blood worms are clearly visible and typically located in the subcutaneous

areas near or at the base of the dorsal and anal fins (Amish 1976). Skin

tumors (Angell et al. 1975; McArn et al. 1968; Miller and Wellings 1971) are

59

found as two main stages: angioepithelial nodules and epidermal papillomas.

All flatfish were externally inspected for blood worms and skin tumors.

Liver tumors are thought to be indicators of pollution (Malins et al.

1962). In the advanced stage, liver tumors are characterized by small nodules

visible on the external surface of the liver. English sole caught in polluted

areas have often been shown to have liver tumors (Malins et al. 1982; Tetra

Tech 1985). Gross examination (non—microscopic) of the external surface of

the livers from about 20% of all flatfish caught was done in the laboratory.

Another disease associated with flatfish in polluted areas is fin

erosion. Fin erosion typically affects the dorsal and anal fins and is

characterized by partial destruction of the fin(s) in question. The severity

ranges from minor defects to extensive destruction of the fin(s) (Wellings et

al. 1976). All flatfish were examined in the field for fin erosion.

Environmental Measurements

Salinity samples were processed by the University of Washington, School

of Oceanography Technical Services group, by conductivity bridge (Paquette

1958). Dissolved oxygen samples were processed by the University of

Washington, School of Fisheries Environmental Laboratory, using titration

techniques described in Standard Methods (1980).

Data Analysis

Species richness, defined as the total number of species present at each

sample site or station, was determined for all stations in Commencement Bay,

Elliott Bay and Saratoga Passage.

Species diversity was calculated using the Shannon—Wiener species

diversity index H’ (Pielou 1978) as follows:

60

H’ = - Z P~ ~

where ~j was the proportion of the total sample that belonged to the ~th

species and n = the number of species. As a consequence of the formula, H’

increases with an increase in the number of species and/or as the individuals

caught become more evenly distributed across all species present.

Abundance and biomass averages were calculated for the combined PSDDA

site samples. “Replicate” samples were taken only at the proposed PSDDA sites

and the NAVY site in Port Gardner; all other stations were sampled once per

season. Length-frequency histograms were constructed for the most abundant

species from the Elliott Bay, Commencement Bay and Saratoga Passage otter

trawl data.

The number of flatfish caught per hectare was calculated for each site by

multiplying the abundance estimates for each flatfish species by the constant

4.5 [which is equal to 10,000 m2 (one hectare) divided by 2,220 m2 (the total

area swept by the otter trawl during each sample)]. Similarly, the reader can

also convert to biomass caught per hectare, or number caught per hectare, for

the remaining fish species by multiplying the given biomass or abundance

values by the constant 4.5.

RESULTS

A total of 55 species of fish were collected by otter trawl during this

study (Table 1). Common names are used throughout this report, although Table

1 lists both the common and scientific names of all fish caught. The

following results are from the otter trawl data only, since beam trawl results

(Appendix A) did not add significant additional information for the purpose of

61

Table 1. List of bottomfish species caught by otter trawl during this study.Species are listed in alphabetical order according to their common name.

Fish SpeciesCommon Name Scientific Name

American shad Alosa sapidissimaarrowtooth flounder Atheresthes stomiasblack eelpout Lycodes diapterusblackbelly eelpout Lycodopsis pacificablackfin starsnout poacher Bathyagonus nigripinnisblacktip poacher Xeneretmus latifronsbluebarred prickleback Plectobranchus evidesbluespotted poacher Xeneretmus triacanthuscanary rockfish Sebastes pinnigerC-O sole Pleuronichthys coenosuscopper rockfish Sebastes caurinusDover sole Microstomus pacificusEnglish sole Parophyrs vetlusflathead sole Hippoglossoides elassodonlingcod Ophiodon elongatuslongfin smelt Spirinchus thaleichthyslongnose skate Raja rhinanorthern ronquil Ronqullus jordan!northern spearnose poacher Agonopsis vulsaPacific cod Gadus macrocephalusPacific hake Merluccius productusPacific herring Clupea harengus pallasiPacific lamprey Lampetra tridentataPacific sanddab Citharichthys sordidusPacific staghorn sculpin Leptocottus armatusPacific tomcod Microgadus proximuspallid eelpout Lycodapus mandibularispile perch Rhacochllus vaccaplainf in midshipman Porichthys notatusquillback rockfish Sebastes maligerratfish Hydrolagus collieired brotula Brosmophycis marginatarex sole Glyptocephalus zachirusrock sole Lepidopsetta bilineatarockfish UID Sebastes sp.roughback sculpin Chitonotus pugetensissablefish Anoplopoma fimbriasailfin sculpin Nautichthys oculofasciatussand sole Psettichthys melanostictussculpin UID Artediussp.shiner perch Cymatogasteraggregatashortfin eelpout Lycodes brevipesslender sole Lyopsetta exilisslim sculpin Radulinus asprellussnailfish UID Cyclopteridaesnake prickleback Lumpenus sagittasoft sculpin Gilbertidia sigalutesspeckled sanddab Citharichthys stigmaeusspiny dogfish Squalus acanthiasspinyhead sculpin Dasycottus setigersplitnose rockfish Sebastes diploproastarry flounder Platichthys stellatussturgeon poacher Agonus acipenserinustadpole sculpin Psychrolutes paradoxuswalleye pollock Theragra chalcogramma

62

final site selections. Abundance, biomass, species richness and species

diversity were used to characterize the fish assemblage at each PSDDA

location.

Flatfish caught per hectare was calculated (Appendix E) at the request of

the U.S. Army Corps of Engineers because 6 flatfish/hectare was recommended as

a preliminary criterion by Washington Department of Fisheries (WDF) as a

minimum number of flatfish needed to support a commercial fishery (WDF 1987).

However, it must be understood that the 7.6 m research otter trawl used in

this study, and by other research groups (e.g., Southern California Coastal

Water Research Project), is selective (as is all sampling gear) and it is

unknown how the 7.6 m trawl catches compare to the actual abundance of

flatfish present, or how the catches compare to the catches used by WDF to

compute the 6 flatfish/hectare criterion. For example, the 7.6 m research

trawl probably catches relatively more juveniles than adults compared to a

commercial trawl.

Commencement Bay

Abundance and biomass. Total abundance and biomass values showed

seasonal and depth differences between many of the catches (Figures 5 and 6).

The summer values were lower than the autumn values. The deeper stations,

which included the PSDDA sites, had the lowest values regardless of season.

Total abundance and biomass values were highest at 40 in then declined at 20 in

(Figures 5 and 6). English sole, Dover sole and ratfish were found in most

samples and generally the PSDDA sites contained the lowest abundance and

biomass of these three species when compared to the samples collected outside

the PSDDA sites. English sole abundance and biomass values were greatest at

40 m in both early summer and autumn, while at the deeper stations, including

the PSDDA sites, the English sole abundance and biomass values were lower than

63

Figure 5. Number (abundance) of fish caught in Commencement Bay, shown bystation and season. The values are based on a single sample,except for the PSDDA sites, which are the average of threesamples. NS,= n~ sampleth

40000

30000

• SUMMER

D AUTUMN

by station and season. The valuessample, except for the PSDDA sites,of three samples. NS not sampled.

• SUMMER

D AUTUMN

N0

0

f

F

SH

400

300

200

100

0

PSDDA 1(165 m)

PSDDA 2(165 m)

156 m

NS Li40 m 20 m

BIgOrM a 20000AmSsS

10000

0

Figure 6.

JL~~zi.156 m

NS~40 m 20 m

PSODA 1 PSDDA 2(165 m) (165 m)

Biomass (in grams) of fish caught in Commencement Bay, shownare based on a singlewhich are the average

64

those of the Dover sole and ratfish (Figures 7, A and B and 8, A and B). The

abundance and biomass of ratfish was greater at the deeper stations than at

either 20 m or 40 m.

Species richness. The values for species richness varied by season and

depth (Figure 9). Summer samples had lower values than the autumn period.

The deeper locations, which included the PSDDA sites, had the lowest values

for species richness.

Species diversity. Values for species diversity, H’, were similar

throughout Commencement Bay, except for the 156 m station (Figure 10). The

156 m station had a much lower value during the summer than the autumn.

Length-frequency. A significant proportion of English sole caught during

the summer at 40 m were less than 205 mm (Figure 11, A and B). These fish

were entirely missing from the autumn samples at the same station (Figure 11,

A and B). English sole caught at 20 m during autumn sampling were larger than

the fish caught at 40 m (Figure 12 and 11B).

Fish health. English sole, Dover sole, rex sole and rock sole all showed

indications of blood worm infections. Incidences ranged from 0~ to 100~

(Table 2). English sole had consistently high infection rates, often as high

as 100~, although the sample sizes associated with the highest incidence rates

were less than 5 fish each. Incidence of skin tumors and fin erosion were all

0%. Gross examination of flatfish livers did not reveal any evidence of liver

tumors.

Environmental measurements. Water temperature showed an inverse relation

to depth (Table 3). Water temperatures were higher at the surface than at

depth, while salinities were lower at the surface and higher at depth. The

Secchi disc measurements were similar at all recording sites.

65

N0

0

F

SH

I -~

20 m

Figure 7. Number (abundance) of English sole, Dover sole and ratfishcaught in Coniinenceinent Bay during summer (A) and autumn (B),shown by station. The values are based on a single sample,except for the PSDDA sites, which are the average of threesamples. NS not sampled.

• English sole Summer

0 Dover sole Summer

[El Ratfish Summer

A

PSDDA 1(165 m)

200

150

100

50

0~—

200

150

100

50

0—

PSDDA 2(165 m)

NS

20 m

• English sole Autum~]

El Dover sole Autumn

El Ratfish Autumn

P ~~( Vt

156 m

N0

0

F

SH

B

PSDDA 1(165 m)

PSDDA 2(165 m)

156 m

66

Figure 8. BiomasscaughtValuessites,

(in grams) of English sole, Dover sole and ratfishin Commencement Bay during summer (A) and autumn (B).are based on a single sample, except for the PSDDAwhich are the average of three samples. NS not sampled.

S English sole Summer

0 Dover sole Summer

0 Ratfish Summer

A

NS

20 mPSDDA 1(165 m)

BIgOrMaAmSsS

BIgOrMaAmSsS

30000

25000

20000

15000

10000

5000

0

30000

25000

20000

1 5000

10000

5000

0

PSDDA 2(165 m)

156 m 40 m

S English sole Autumn

0 Dover sole Autumn

~ Ratfish Autumn

B

20 mPSDDA 1 PSODA 2(165 m) (165 m)

156m 40m

67

SpEC

ES

R

CHNESS

Figure 9.

SpEC

ES

2.0

PSDDA 1(165 m)

Figure 10.

Species richness (total number of species) of fish caught inCommencement Bay, shown by station and season. NS = not sampled.

PSDDA 2(165 m)

Species diversity (H’) of fish caught in Commencement Bay,shown by station and season. NS = not sampled.

16

14

12

10

8

6

4

2

0

m SUMMER

D AUTUMN

PSODA 1 PSDDA 2(165 m) (165 m)

-~L

156m 40m 20m

1.8

1.6

1.4

1.2

1.0

0.8

0.6

0.4

0.2

0.0

D

VEBS

TY

• SUMMER

D AUTUMN

156 mNS

40 m 20 m

68

7

S English sole - male A6 EJ Englsih sole - female

5FREQUENCY

~ ;~ ‘ .. ]~~I_~

125 145 165 185 205 225 245 265 285 305 325 345 365 385 405LENGTH mm

16

S English sole - male B14 D English sole - female

12

FR 10EQU 8ENC 6Y

~~Ji I ii. I ~B, , , , , , ,

125 145 165 185 205 225 245 265 285 305 325 345 365 385 405LENGTH

Figure 11. Length frequency of English sole, shown by sex, caught inCommencement Bay during summer (A) and autumn (B) at 40 m.

69

4

S English sole - female

EJ English sole - female

3

FRE0U 2ENCY ~ — ~ ~I- ~ 1r_,__~__r__

125 145 165 185 205 225 245 265 285 305 325 345 365 385 405

LENGTH mm

Figure 12. Length freq~uency of English sole, shown by sex, caught inCommencement Bay during autumn at 20 m.

Ta

ble

2.

Pe

rce

nt

incid

en

ce

and

sam

ple

siz

e(in

pa

ren

the

se

s)

of

blo

od

wor

m(P

hilo

me

tra

sp

.)in

fectio

nin

fla

tfis

hca

ug

ht

inC

omm

ence

men

tB

ay,

show

nb

ysp

ecie

s,

sta

tio

nan

dse

aso

n.

S=

sum

mer

,A

au

tum

n.

PSD

DA

1PS

DD

A2

(16

5m

)(1

65

m)

156

m40

m20

mS

AS

AS

AS

AS

A

En

glis

hso

le0(

2.)

10

0(1

)1

00

(4)

33

(3)

43

(40

)4

8(1

15

)5

8(1

9)

Do

ver

so

le0

(4)

0(3

)0

(5)

0(1

0)

0(6

)9

(11

)0

(5)

35

(5)

Sle

nd

er

so

le0

(3)

0(1

3)

0(2

)0

(4)

0(3

)0

(12

)

Roc

kso

le0

(2)

20

(10

)3

1(1

6)

Rex

so

le1

00

(1)

0(1

)0

(3)

0(5

)0

(7)

Arr

ow

too

thflo

un

de

r0

(1)

C—0

so

le0

(1)

0(1

)S

pe

ckle

dsa

nd

da

b0

(3)

C

71

Table 3. Environmental measurements of temperature, salinity and waterclarity in Commencement Bay during autumn by station.

Surface Depth

Temperature, OC

PSDDA 1 (165 m) 15.1 11.3PSDDA 2 (165 m) 14.3 11.240 m 13.5 12.0

Salinity, O/oo

PSDDA 1 (165 m) 26.8 30.9PSDDA 2 (165 m) 28.6 30.940 m 28.7 30.3

Secchi, m

PSDDA 1 (165 m) 5.0PSDDA 2 (165 m) 6.040m 6.0

72

Elliott Bay

Abundance and biomass. The abundance and biomass values varied by

station and by season (Figures 13 and 14). Abundance and biomass values were

higher for autumn than for summer at all stations except PSDDA 1 Reference

station.

Six species of fish dominated the catches in Elliott Bay: English sole,

Dover sole, Pacific hake, slender sole, ratfish and blackbelly eelpout,

although not every species was found at each site (Figures 15, A and B and 16,

A and B). The PSDDA 1 site (inner Elliott Bay) was the shallowest and had the

largest abundance and biomass of Pacific hake, slender sole and blackbelly

eelpout. The PSDDA 2 site Fourmile Rock) had a greater abundance and biomass

of English sole, Dover sole and ratfish than the PSDDA 1 site. The PSDDA 2

site had lower abundance and biomass values compared with the values found at

the adjacent reference stations. Generally, abundance and biomass values

increased from the summer to the autumn sampling; specifically, English sole,

Dover sole, and ratfish. The shallower PSDDA 1 area had greater numbers of

the smaller fishes such as blackbelly eelpouts and slender sole in contrast to

the deeper PSDDA 2 area where the larger species dominated.

Species richness. The values for species richness varied by season and

depth (Figure 17). Species richness was generally lower during the summer

than the autumn, except for the PSDDA 1 reference station, where values were

the same. The PSDDA 1 site and the PSDDA 1 reference station generally had

larger values than the PSDDA 2 site and the PSDDA 2 reference stations, except

for the PSDDA 2 site during the autumn.

Species diversity. The values for species diversity, H’, generally

diminished from the inner bay PSDDA 1 site and reference station to the

Fourmile Rock PSDDA 2 site and reference station (Figure 18) irrespective of

73

Figure 15. Number (abundance) of fish caught in Elliott Bay shown bystation and season. The values at the PSDDA ref. sites arebased on a single sample, while the values at the PSODA sitesare averages of three samples. NS = not sampled.

Biomass (in grams) of fish caught in Elliott Bay,station and season. The values at the PSDDA ref.based on a single sample, while the values at theare averages of three samples. NS not sampled.

250

200

150

100

50

N0

0

F

SI-I

• SUMMER

D AUTUMN

NS0 I

PSDDA 1(75 m)

.1PSDDA 1 PSDDA 2 PSDDA 2 PSODA 2

REF (172 m) REF1 REF2(90 m) (180 m) (175 m)

25000

20000

r g 15000

MaAm

s 10000

S

5000

Figure 14.

•EJ AUTUMN

NS0

PSDDA 1(75 m)

PSDDA 1 PSDDA 2 PSDDA 2 PSDDA 2REF (172 m) REF 1 REF 2

(90 m) (180 m) (175 m)

shownsitesPSDDA

byaresites

74

100 A• English sole Summer

900 Dover sole Summer

80 ~ Slender sole Summer

N 70 D Ratfish Summero 0 BB Eelpout Summer

600

50

F40

SH 30

20

1: r,_1~.LJ1 __~1~~1~] r~ NS

PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 PSDDA 2(75 m) REF (172 m) REF 1 REF 2

(90 m) (180 m) (175 m)

100B

90 English sole Autumn

80 0 Dover sole Autumn

L~ Slender sole AutumnN 70o D Ratfish Autumn

60 0 BB Eelpout Autumn

50 R Pacific hake Autumn

F40

SH 30

~ ~ ~PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 PSDDA 2

(75 m) REF (172 m) REF1 REF2

(90 m) (180 m) (175 m)

Figure 15. Number (abundance) of English sole, Dover sole, slender sole,ratfish and blackbelly eelpout (BB) caught in Elliott Bayduring summer (A) and autumn (B), shown by station. The valuesat the PSDDA ref. sites are based on a sin~le sample, while thevalues at the PSDDA sites are averages of three samples.NS not sampled.

75.

Figure 16. Biomass (in grams) of English sole, Dover sole, slender sole,ratfish and blackbelly eelpout (BB) caught in Elliott Bayduring summer (A) and autumn (B), shown by station. The valuesat the PSDDA ref. sites are based on a single sample, while thevalues at the PSDDA sites are averages of three samples.

S English sole Summer

0 Dover sole Summer

~ Slender sole Summer

0 Ratfish Summer

D BB Eelpout Summer

A

B1g

MaAmssS

B

MaAmssS

8000

7000

6000

5000

4000

3000

2000

1000

0

8000

7000

6000

5000

4000

3000

2000

1000

0

PSDDA 1 PSODA 1 PSDDA 2 PSDDA 2 PSDDA 2(75 m) REF (172 m) REF 1 REF 2

(90 m) (180 m) (175 m)

S English sole Autumn

0 Dover sole Autumn

~ Slender sole Autumn

0 Ratfish Autumn

D BB Eelpout Autumn

B

PSDDA 1(75 m)

PSDDA 1REF

(90 m)

PSDDA2 PSDDA2 PSDDA2(172 m) REF1 REF2

(180 m) (175 m)

NS not sampled

76

D

VERS

TY

• SUJIMER

D AUT~~j

• SUMMER

D AUTUMN

J

20

18

S16

EC 14

E 12S

R 10

c 8HN 6ES ~S

2

0

PSDDA 1 PSDDA 1(75 m) REF

(90 m)

Figure 17. Species richness (total number of species) of fish caughtin Elliott Bay, shown by station and season. NS = not sampled.

Sp ______________

EC

ES

PSDDA 2 PSDDA 2 PSDDA 2(172 m) REF 1 REF 2

(180 m) (175 m)

2.0

1.8

1.6

1.4

1.2

1.0

0.8

0.6

0.4

0.2

0.0PSODA 1(75 m)

PSDDA 1REF

(90 m)

PSDDA 2(172 m)

NS

PSDDA 2REF 1

(180 m)

Figure 18. Species diversity (H’) of fish caught in Elliott Bay,shown by station and season. NS = not sampled.

PSDDA 2REF 2

(175 m)

77

season.

Length-frequency. English sole data from the PSDDA 2 Reference Station 1

during autumn suggested a bimodal distribution of adults with no juveniles

present (Figure 19).

Fish health. English sole, Dover sole and flathead sole showed evidence

of blood worm infections (Table 4). Incidence in these three species ranged

from 0% to 42%. The PSDDA 2 area had the highest incidence of blood worm

infection in English sole and Dover sole with flathead sole showing only a

minor incidence. There were no indications of skin tumors or fin erosion.

Gross examination of flatfish livers did not show any indication of liver

tumors.

Environmental measurements. Temperature and dissolved oxygen values were

higher at the surface than at depth (Table 5), while salinity was lower at the

surface than at depth. Dissolved oxygen and Secchi disc measurements showed a

seasonal pattern: dissolved oxygen was slightly lower in autumn than in summer

and Secchi disc values were slightly higher in autumn than in summer.

Saratoga Passage

Abundance and biomass. Only one sample cruise on July 1 was conducted in

Saratoga Passage. Abundance and biomass showed variation by depth and by

station (Figures 20 and 21). Abundance relative to biomass was greater for

all stations except for the PSDDA site. The PSDDA site had an intermediate

abundance value and had the highest biomass value. The dominant species

included ratfish, English sole, Dover sole, slender sole and adult Pacific

hake. Pacific hake were found in the PSDDA site and the reference station,

while English sole were only found at the shallower locations. Dover sole

were confined to the 40 m west station. Ratfish and slender sole occurred at

78

8

7

6

F S English sole - maleR D English solo - femaleEaLJ4EN

Y

2

125 145 165 185 205 225 245 265 285 305 325 345 365 385

LENGTH

Figure 19. Length frequency of English sole, shown by sex, caught atPSDDA 2, reference 1, during autumn in Elliott Bay.

Ta

ble

4.

Pe

rce

nt

incid

en

ce

and

sam

ple

siz

e(in

pa

ren

the

se

s)

of

blo

od

wor

m(P

hilo

me

tra

sp

.)in

fectio

nin

fla

tfis

hca

ug

ht

inE

llio

ttB

ay,

show

nb

ysp

ecie

s,

sta

tio

nan

dse

aso

n.

Ssu

mm

er,

A=

au

tum

n.

PSD

DA

1PS

DD

A1

Re

fPS

DD

A2

PSD

DA

2R

ef

1PS

DD

A2

Re

f2

SA

SA

SA

SA

SA

(75

m)

(90

in)

(17

2m

)(1

80

in)

(17

5in

)

En

glis

hso

le0

(1)

0(2

)3

5(5

5)

0(1

)1

4(2

8)

42

(33

)D

ove

rso

le0

(13

)0

(7)

0(1

)0

(26

)5

(20

)0

(9)

0(2

)0

(3)

Sle

nd

er

so

le0

(80

)0

(14

8)

0(1

3)

0(6

2)

0(3

5)

0(2

8)

0(3

)0

(8)

0(9

)F

lath

ea

dso

le6

(54

)3

(73

)0

(15

)0

(17

)0

(5)

Roc

kso

le0

(5)

0(1

)0

(1)

Rex

so

le0

(9)

0(9

)0

(1)

80

Table 5. Environmental measurements of water temperature, dissolvedoxygen, salinity and water clarity in Elliott Bay, by stationand season.

Summer AutumnSurface Bottom Surface Bottom

Temperature, 0C

PSDDA 1 (75 m), 13.5 10.3 18.5 11.9PSDDA 2 (172 m) 13.9 11.0 12.9 —

Dissolved Oxygen, mg/l (% saturation)

PSDDA 1 (75 m) 9.72 (110%) 7.79 (84%) 8.36 (105%) 7.67 (92%)PSDDA 2 (172 m) 9.77 (113%) 7.65 (84%) 6.34 (73%) -—

Salinity, o/oo

PSDDA 1 (75 m) 28.1 29.9 28.9 30.7PSDDA 2 (172 m) 28.8 30.2 30.1

Secchi, m

Summer Autumn

PSDDA 1 (75 m) 4.0 7.0PSDDA 2 (172 m) 4.5

BIgOrMaAmSsS

20 T18

16

N 140

120

F8

SI-f

40 m W

81

6-

4-.

2-

0-

Figure 20. Number (abundance) of fish caught in Saratoga Passage duringsummer, shown by station. The values are based on a singlesample, except for the PSDDA site, which is the average ofthree samples.

4000

3000

2000

1000

0 - -

Figure 21. Biomass (in grams) of fish caught in Saratoga Passage duringsuraner, shown by station. The values are based on a singlesample, except for the PSDDA site, which is the average ofthree samples.

PSDDA(108 m)

82

the deeper (PSDDA) stations and intermediate depths (Figures 22 and 23).

Species richness. Values for species richness fluctuated by depth; the

highest values occurred at the PSDDA site reference station (Figure 24). All

other species richness values were lower and showed no discernible pattern.

Species diversity. Values for species diversity, H’, varied by depth,

the deeper stations, including PSDDA, had the highest values (Figure 25). No

pattern was apparent among the shallower stations.

Fish health. Incidence of blood worms, skin tumors and fin erosion were

all 0% (Table 6). No evidence of liver tumors was found based on gross

examination of flatfish livers.

Environmental measurements. No environmental measurements were collected

in Saratoga Passage because of weather conditions that forced an early

curtailment of sampling.

Port Gardner

Abundance and biomass. Abundance and biomass fluctuated by time of year,

depth, and station. The NAVY site generally had the largest number and

biornass of fish throughout the year. During the winter the 40 m depth had

numbers of fish comparable to the NAVY site; however, the biomass values were

lower. (Figures 26 and 27). PSDDA 1 and PSDDA 2 sites had low values for

abundance and biomass for all seasons except winter when abundance was at its

highest (compared with other seasons) and biomass values were second only to

the NAVY site (Figures 26 and 27). Five of the locations that were sampled

throughout the year were situated at depths of 100 m or more; these included:

PSDDA 1, PSDDA 2, lOOm M, llOm S and 145m S. The hOrn S and 145m S stations

had the lowest abundance and biomass values of the 5 deep locations.

Species diversity. Species diversity, H’, values showed seasonal and

depth differences between many stations, but showed no discernable pattern

83

Number (abundance) of English sole, Dover sole, slender sole,Pacific hake and ratfish caught in Saratoga Passage duringsummer, shown by station. The values are based on a singlesample, except for the PSDDA site, which is the average ofthree samples.

Biomass (in grams) of English sole, Dover sole, slender sole,Pacific hake and ratfish caught in Saratoga Passage duringsummer, shown by station. The values are based on a singlesample, except for the PSDDA site, which is the average ofthree samples.

ii IPSDDA

REF(100 m)

I80 m E20 m E 40 m E 40 m W

7,

• English sole Summer6

0 Dover sole Summer

N 5 0 Slender sole Summero ~ Pacific hake Summer

o 4 0 Raffish Summer

F 3

PSODA(108 m)

Figure 22.

900

800

700

8 600IgOr 500MaA m 400SaS 300

200

100

0

Figure 23.

S English sole Summer

0 Dover sole Summer

~ Slender sole Summer

~ Pacific hake Summer

0 Raffish Summer

PSDDA PSDDA(108 m) Ref

(100 m)

80 m E

SPEC

ES

R

CHNESS

SPEC

ES

D

VERS

TY

Species diversity (H’) of fish caughtduring summer, shown by station.

in Saratoga Passage

84

6

5

4

3

2

PSDDA(108 m) REF

(100 m)

Figure 24. Species richness (total number of species) of fish caught in

Saratoga Passage during summer, shown by station.

1.8

1.6

1.4

1.2

1.0

0.8

0.6

0.4

0.2

0.0

PSDDA(108 m)

Figure 25.

40 m E 40 m W

85

Table 6. Percent incidence and sample size (in parentheses) ofbloodworm (Philometra sp.) infection in flatfish caught atSaratoga Passage during summer, shown by station and species.

PSDDA 8Oiu E 40m B 40m W 20m B(108 m)

English sole 0(7) 0(7) 0(2) 0(4)Slender sole 0(17) 0(4)Dover sole 0(1)Rock sole 0(i) 0(6) 0(2)

Fig

ure

26

.N

umbe

r(a

bu

nd

an

ce)

of

fish

cau

gh

tin

Po

rtG

ard

ne

r,sh

own

by

sta

tio

nan

dse

aso

n.

The

va

lue

sa

reb

ase

don

asin

gle

sam

ple

,e

xce

pt

for

the

NAV

Yan

dPS

DD

Asite

s,

wh

ich

are

the

ave

rag

eo

fth

ree

sam

ple

s.NS

=n

ot

sam

ple

d.

400

1Rw8

6N

jDSP

86

030

0

~S

~u

se

0 f2

00

~1~I_~k9,I

~j~

NS

~N

S

f i ~10

0

h

~N

S0

NAV

YPS

DD

A2

PSD

DA

1R

ef1

Ref

2St

eE

(80

m(

(120

m(

(137

m)

(132

m)

(128

m)

(105

m(

200

ave

00

88

6

~SU

86

N 0 0 f f 1 S h N 0 0 f f 1 S h

4Om

S10

0mM

Tran

sect

1

N 0 0 f f 1 S h

30

0

200 00

• 088

86

~SU

86

&3A

U

L,J

302

200 00

20m

S

008

86

~20

86

AU86

4Om

STr

anse

ct2

11

0m

S4

Om

S1

45

mS

Tran

sect

4

40

00

0

B ig 0r

ma

aa

SS

S

B ig

0r

ma

am

SS

S

30

00

0B 1 0 m a S S

g r a m S

20

00

0

•W

86

OS

P88

~S

U86

~A

UB

O

10

00

0

(80

•w

as spas

~SU

S6

Ro

t(1

32

2S

taE

m)

(10

5m

)2

Om

S4

Om

ST

ran

se

ct

1

4000

0

4000

0

3000

0

0000

0

0000

40

00

0

30

00

0

20

00

0

1000

0

3000

0

B ig

0T

aa

aa

SS

1000

0

UW

OS

~SP

86

~SU

86

~A

U~

2O

mS

•W

86

0S

P8S

~SU

BS

£SA

U86

4Om

ST

ran

se

ct

21

lOm

S

Fig

ure

27

.

2Om

S4O

mS

14

5m

ST

ran

se

ct

4

Bio

ma

ss(in

gra

ms)

of

fish

cau

gh

tin

Po

rtG

ard

ne

r,sh

own

bysta

tio

nan

dse

aso

n.

The

va

lue

sa

reb

ase

don

asin

gle

sam

ple

,e

xce

pt

for

the

NAV

Yan

dPS

DD

Asite

s,

wh

ich

are

the

ave

rag

eo

fth

ree

sam

ple

s.NS

=n

ot

sam

p1ed

~

88

(Figure 28).

Fish health. English sole, Dover sole, flathead sole, rex sole and rock

sole all showed indications of blood worm infections (Table 7). Incidence

varied between species, seasons, depth and station but did not show a

discernable pattern. One skin tumor was noted on a slender sole caught at

Station 100 mN. Incidence of fin errosion was 0%. Gross examination of

flatfish livers did not reveal any evidence of liver tumors.

Environmental measurements. Water temperatures during winter and autumn

were higher at the bottom than the surface (Table 8). Spring and summer water

temperatures were the reverse with the surface warmer than the bottom. In

general, salinities were lower at the surface than the bottom. Secchi disc

measurements showed that the best water clarity (higher Secchi disc

measurement) occurred in the winter while there were no differences between

the other seasons.

DISCUSSION AND CONCLUSIONS

The Research Otter Trawl For Documenting Fish Assemblage

The 7.6 m otter trawl has been the dominant sampling gear in Puget Sound

demersal fish research for about the last decade. This net is widely used by

many groups for similar research in other areas of the country.

Standardization of gear reduces the problems associated with comparing results

between studies. In addition, the small size of the net allows for ease of

use from a range of vessel sizes starting at about 6 m.

The 7.6 m otter trawl has limitations. The net is not fished

commercially and due to size, shape and other differences, catches are not

directly comparable to commercial otter trawl catches. Other limitations

S

peC

e8

V

e

S

V

S

peC

eS

DH

V

e

S

Y

n0

PSODA 2 PSODA 1 NAVY Ref 1 Ref 2 Sta E(120 m) (137 m) (80 m) (132 m) (128 m) (105 m)

2.5

2.0

1.5

1.0

0.5

0.0

2.5

2.0

1.5

1.0

0.5

0.0

Figure 28. Species diversity of fish (H1) caught in Port Gardner,shown by station and season. NS = not sampled.

205

Transect 1 Transect 2 Transect 4

Ta

ble

7.

Pe

rce

nt

incid

en

ce

an

dsa

mp

lesiz

e(in

pa

ren

the

se

s)

of

blo

od

wo

rm(P

hilo

me

tra

sp

.)

infe

ctio

nin

fla

tfis

h,

show

nb

ysp

ecie

s,

sta

tio

nan

dse

aso

na

tP

ort

Ga

rdn

er.

W=

Win

ter,

SP=

Sp

rin

g,

SUS

umm

er,

AU—

Au

tum

n,

Fia

tfish

Spe

cies

NAV

YP

SD

DA

2P

SD

DA

1W

SP

SLJ

AUW

SPSU

AUW

SP

SUAU

Rex

sole

50

(2)

0(1

8)

0(1

)0

(2)

0(1

)0

(3)

Flat

head

sole

3(7

7)

0(1

7)

0(7

)0

(3)

0(1

)R

ock

sole

0(1

)1

00

(1)

Sle

nder

sole

1(11

8)4

(56

)0

(15

)0

(66

)0

(29

)0

(14

)0

(6)

0(2

3)

0(7

8)

0(2

6)

0(9

)0

(23

)D

over

sole

5(2

2)

0(9

)0

(2)

0(1

0)

0(1

8)

0(2

1)

0(5

)0

(11

)0

(2)

0(1

4)

0(5

)E

nglis

hso

le1

0(3

35

)8

(91

)2

(86

)6

(16

9)

0(7

)1

3(8

)0

(37

)1

0(1

0)

5(2

1)

50

(2)

33

(1)

13

(8)

Tran

120

STr

an1

40S

Tran

1lO

OM

WS

PSU

AUW

SPSU

AUW

SPSU

AUR

exso

le0

(1)

0(3

)0

(2)

0(2

)F

lath

ead

sole

0(1

)0

(5)

0(2

)R

ock

sole

0(4

)0

(18

)0

(1)

17

(6)

0(7

)S

lend

erso

le0

(1)

0(5

)1

00

(2)

0(1

5)

33

(3)

Dov

erso

le0

(12

)0

(2)

0(2

)0

(1)

0(2

)E

nglis

hso

le0

(2)

0(1

)0

(5)

0(1

74

)0

(2)

0(1

5)

6(1

7)

8(2

6)

10

(61

)0

(28

)3

3(3

)

Tran

220

STr

an2

40S

Tran

211

OS

WS

PSU

AUW

SPS

liAU

WSP

SUAU

Rex

sole

0(1

)0

(3)

0(1

)0

(1)

Fla

thea

dso

le0

(1)

Roc

kso

le0

(3)

10

0(1

)0

(2)

0(1

)2

8(1

8)

20

(5)

0(5

)0

(3)

Sle

nder

sole

0(2

)0

(6)

0(2

)0

(3)

0(1

)0

(2)

0(4

)D

over

sole

0(1

)0

(1)

8(5

2)

0(1

1)

0(1

)0

(3)

0(7

)E

nglis

hso

le0

(2)

0(3

)2

(12

1)

66

(3)

0(2

2)

0(5

4)

25

(4)

0(6

)

Tran

420

STr

an4

405

Tran

414

5SW

SP

SUAU

WSP

SUAU

WS

PSU

AUR

exso

le0

(3)

Fla

thea

dso

le0

(4)

0(1

)R

ock

sole

10

0(1

)0

(2)

0(5

)2

6(3

1)

38

(13

)S

lend

erso

le0

(4)

0(1

4)

0(1

)0

(1)

0(1

)D

over

sole

0(1

1)

0(1

)0

(7)

0(2

8)

0(5

)E

nglis

hso

le0

(8)

0(6

)0

(14

)2

(14

0)

21

(19

)1

7(6

)

Re

fiR

ef2

Sta

ESU

AUSU

AUAU

Rex

sole

Flat

head

sole

Roc

kso

leS

lend

erso

le0

(2)

0(3

)0

(15

)0

(6)

Dov

erso

le0

(6)

0(2

)0

(4)

0(1

)0

(2)

Eng

lish

sole

0(5

)0

(4)

0(5

)1

4(7

)0

(3)

91

Table 8. Measurements of temperature, salinity and water clarity bystation and season at Port Gardner. W = Winter, SP = Spring,SU = Summer, AU = Autumn3 NS = not sampled.

Site Surface BottomTemperature ~C

W SP SU AU W SP SU AUNAVY 7.0 10.5 11.9 14.0 7.5 9.5 11.0 12.0PSDDA 2 6.5 10.0 15.2 15.0 8.0 9.0 11.0 13.0PSDDA 1 6.0 10.2 15.0 14.4 7.5 9.3 9.9 12.0Tran 1 2OmS NS 10.8 18.1 15.0 NS 9.5 11.5 13.0Tran 2 4OmS 6.5 10.5 10.5 14.0 7.5 NS 11.5 12.0Tran 4 l45mS 6.0 10.5 12.0 NS 8.0 9.0 12.5 NS

Salinity 0/00NAVY 29.85 16.79 22.34 28.73 NS 29.73 NS 30.58PSDDA 2 21.23 18.53 23.58 NS NS 29.67 29.81 30.81PSDDA 1 NS 22.98 24.99 28.23 NS 29.85 24.99 30.56Tran 1 2OmS NS NS 19.58 28.32 NS 29.16 29.58 30.07Tran 2 4OmS NS NS 29.81 NS NS 29.53 29.77 30.33Tran 4 145mS 22.01 NS 23.58 NS NS 30.10 29.70 NS

Secchi Disc mNAVY 3.50 3.00 3.00 5.00PSDDA 2 5.50 4.25 4.50 4.00PSDDA 1 8.50 7.25 4.50 NSTran 1 2OmS NS 3.50 3.00 5.50Tran 2 40m5 3.50 3.00 3.00 5.00Tran 4 145mS 4.50 NS 3.50 NS

92

include: 1) current and wind conditions at the time of sampling, 2) gear

selectivity relative to each fish species and life history stage, and 5)

variability in area swept.

Commencement Bay

Several trawling studies have previously been conducted in Commencement

Bay. These studies concentrated their efforts in the nearshore areas (Becker

1984; Weitcamp and Schadt 1981; Tetra Tech 1985) and in the inner part of

Commencement Bay (e.g., the old flood channels of the Puyallup River (Malins

et al. 1982; Tetra Tech 1985; Weitkamp and Schadt 1981). Becker (1984) and

Tetra Tech (1985) used the same otter trawl as the present study; however,

sampling depths only reached 52 meters in contrast to the 175 meter depths of

the proposed PSDDA sites. Weitkamp and Schadt (1981) used a different smaller

otter trawl and again only sampled the nearshore shallow portions of

Commencement Bay.

Data from Commencement Bay indicate that three of the four indices of

site utilization by fish varied inversely with depth. As depth increased,

species richness, total abundance and total biomass decreased. No correlation

between depth and species diversity was evident. However, Tetra Tech (1985)

found the species diversity in the inner harbor waterways to be much higher

(3.5) compared to study. Results of this study suggest higher catches

occurring in deeper water during autumn, while Weitkamp and Schadt (1981)

found abundance in shallower areas was highest in summer and lower during

other seasons of the year. Becker (1984) found that Dover sole were located

at deeper stations while English sole were typically found in shallower

waters, similar to the findings of the present PSDDA study.

Fish health was generally good for the flatfish caught in Commencement

Bay during this study. The only disease found was blood worm infection in

93

English sole. Incidence for this disease reached 100% at some stations, but

the sample sizes were small (less than five individuals per sample) for

locations with high incidence rates. Low incidences (< 1%) of fin erosion and

skin tumors have been previously found in flatfish inhabiting the inner

(shallow) portions of Commencement Bay in areas known to be contaminated with

industrial wastes (Malins et al. 1982, Tetra Tech 1985). Histopathological

analyses of livers from flatfish found inhabiting these areas also revealed a

low incidence of liver tumors (Malins et al. 1982, Tetra Tech 1985).

Environmental measurements were only available for the autumn period.

Dissolved oxygen, temperature, salinity and Secchi disc measurements were all

within the ranges found in other parts of Puget Sound (Stober and Chew 1984).

The preliminary PSDDA sites in Commencement Bay had the greatest depth

and the lowest abundance, biomass and species richness measures of the

stations sampled during this study. The number of adult and juvenile flatfish

captured in the preliminary PSDDA sites was low in absolute terms and small

when compared to stations outside of the PSDDA sites.

Elliott Bay

Several studies have been conducted within and adjacent to Elliott Bay

(Bingham 1978; Miller et al. 1974; Miller et al. 1977; Miller 1980; Maims et

al. 1982; Stober and Chew 1984). Donnelly et al. (1984) and Bingham (1978)

were the only studies that sampled the same areas as the present PSDDA study.

The results of fish sampling in Elliott Bay revealed that the PSDDA sites

generally had higher values of species richness than their corresponding

reference stations. The PSDDA 2 (Fourmile Rock) site and the adjacent

reference stations exhibited a pattern much like PSDDA 1 (inner Elliott Bay):

summer season abundance, biomass and species richness figures were comparable,

94

whereas the autumn values of biomass and species richness for the PSDDA 2 site

exceeded the reference stations. Species diversity did not show any clear

pattern.

PSDDA 2 samples were taken in close proximity to samples collected for

the Renton Sewage Treatment Plant Project (Donnelly et al. 1984). Donnelly et

al. found biomass values higher at the deeper sites in contrast to this study

where the PSDDA 2 site had higher biomass values than the deeper reference

stations. Donnelly et al. (1984) also indicated that. species richness and

species diversity increased with depth to 50 m and then decreased. This would

suggest that the PSDDA 2 site would have higher species richness and species

diversity values than the PSDDA 2 reference stations. Indeed, PSDDA 2 values

were either comparable to, or exceeded, the reference station values for

species richness and species diversity. Neither study was conducted during

all four seasons, thus, results from this study should not be considered

indicative of conditions at the sample sites throughout the year.

The PSDDA 1 site was compared with a report on the effects of dredged

materials disposal on benthic invertebrates in inner Elliott Bay (Bingham

1978). The 1978 report found no substantial difference in infaunal

(invertebrate) species richness or biomass, but did find that the shallower

stations had the greatest species richness and biomass. The same observation

was made at the PSDDA 1 sites (for fish) where the deeper reference station

had lower species richness and biomass. Neither Bingham (1978) nor this study

found any clear trend in species diversity versus depth.

Fish health was generally good. Blood worm infection in English sole was

the only disease noted. The incidence was somewhat lower than the findings of

Donnelly et al. (1984). Donnelly et al. also noted the presence of other

diseases: skin tumors and fin erosion. Fin erosion, skin tumors and liver

95

tumors have been found in Elliott Bay, but typically near the inner shore and

the Duwamish River (Malins et al. 1982). The present study sites were located

in the deeper regions away from the shore of Elliott Bay and may explain why

the disease incidence was found to be lower than at previous inshore sampling

locations.

Environmental measurements were available for both seasons. The

dissolved oxygen values were all near saturation. Dissolved oxygen,

temperature, salinity and Secchi disc measurements were all within the values

found elsewhere in Puget Sound (Miller et al. 1977; Donnelly et al. 1984).

The preliminary PSDDA sites generally had the highest values for

abundance, biomass and species diversity of the locations sampled in Elliott

Bay. Dover sole and English sole, two commercially important flatfish, were

found in higher numbers and biomass at the PSDDA 2 site and adjacent reference

stations than at PSDDA 1. These results suggest that disposal of dredged

materials would have less impact on commercial flatfish at the PSDDA 1 site

than at PSDDA 2 site.

Saratoga Passage

The Saratoga Passage PSDDA site ecological measures were generally higher than

the adjacent, shallower stations. A tagging study done by Day (1976) suggests

that the deeper area of Saratoga Passage was a residence or spawning area for

English sole. Day captured English sole in the deep area of Saratoga Passage

where they were tagged and transported to other areas of Puget Sound. The

majority of tag recoveries were within the deep area of Saratoga Passage,

suggesting a homing ability to a preferred feeding location or possibly a

spawning area. No English sole were caught at the PSDDA site during the

sampling cruise; however, English sole were caught nearby at 80 m. Sampling

was too limited to make any concluding statement about the Saratoga Passage

96

PSDDA site.

No diseases were found in any flatfish.

Port Gardner

English (1979) and Washington Department of Ecology (1976) sampled bottom

fish at several depths along a transect line very close to Transect 2 of the

PSDDA study. During the PSDDA study the catches of English sole less than

150 mm were lower than those found by English (1979). Each study used

different capture gear; the earlier study used a 3 m beam trawl, while study

used the 7.6 m otter trawl. Other explanations for the abundance differences

may be biological or environmental in origin. Shephard et al. (1984)

indicated three major reasons for variation in stock size from one year to the

next: 1) annual changes in environmental conditions, 2) ecological

interactions during early life history, and 3) variations in adult spawning

abundance.

The results of the fish sampling in Port Gardner showed that the PSDDA

sites had low abundance and biomass values when compared to other stations for

spring, summer and autumn. In contrast, winter sampling at the PSDDA sites

produced abundance values higher than most stations except for the NAVY and

the 40 m depth. Furthermore, biomass values at the two PSDDA sites during the

winter were second only to the NAVY site. These data suggest that the larger,

older fish move into the preliminary PSDDA sites during the winter months and

disperse during the rest of year.

Species diversity did not show any trends that could be related to

differences between stations or sites.

Fish health was generally good. Malins et al (1982), using microscopic

examination of flatfish livers, found only low levels of nonspecific

degenerative/necrotic lesions and intracellular storage disorders in Port

97

Susan (adjacent to Port Gardner). Gross examination of livers for this study

showed no evidence of tumors.

Washington Department of Fisheries (WDF) allows an annual commercial

flatfish harvest during the spring and early summer in Port Gardner. The

fishing area includes not only Port Gardner, but the adjacent main basin and

the area adjoining Port Gardner to Saratoga Passage. Catches from the

combined area contribute about 9% to the total annual Puget Sound flatfish

harvest. Catches within Port Gardner itself contribute only a small portion

of the 9% (WDF personal communication).

The preliminary PSDDA sites may be subjected to low levels of commercial

trawling since they are located in the deeper portions of Port Gardner and,

for most of the year, had low values for abundance and biomass relative to

other stations. In contrast, winter abundance and biomass values at the

preliminary PSDDA sites were much higher and second only to the NAVY site.

The levels of abundance and biomass were very similar between the two PSDDA

sites during every season, thus equal consideration (relative to fish

resources) should be given to both preliminary PSDDA sites for disposal of

dredged materials. However, data from Transect 4, Station 145 5, which is

deeper and located more near the mouth of Port Gardner, show much lower

abundance and biomass values than the PSDDA sites. The fish data from Port

Gardner has not been fully compiled and analyzed, but when completed, will be

issued as part of the Port Gardner report to the U. S. Army Corps of Engineers.

98

LITERATURE CITED

Amish, R. 1976. Infestation of some Puget Sound demersal fishes by blood

worm, Philometra americana. M.S. Thesis, Univ. Washington, Seattle. 41

pp.

Angell, C. L., B. S. Miller, and S. R. Wellings. 1975. Epizootiology of

tumors in a population of juvenile English sole (Parophyrs vetulus) from

Puget Sound, Washington. J. Fish. Res. Board Canada 52:1723—1732.

Becker, D. 5. 1984. Resource partitioning by small—mouthed pleuronectids in

Puget Sound, Washington. Ph.D. Dissertation, Univ. Washington, Seattle.

139 pp.

Bingham, C. R. 1978. Aquatic disposal field investigations, Duwamish

Waterway disposal site, Puget Sound, Washington; Appendix 0: Benthic

community structural changes resulting from dredged material disposal,

Elliott Bay disposal site. U.S. Army Eng. Waterways Exp. Stn. Vicksburg,

MI, Tech. Rep. D—77—24.

Butler, T. H. 1980. Shrimps of the Pacific Coast of Canada. Can. Bull.

Fish. Aquatic Sci. 202:280 pp.

Chew, K. K. Unpublished. Shrimp trawl data collected from Hood Canal and

Puget Sound from 1967 to 1979. School of Fisheries, University of

Washington, Seattle.

Clarke, D. 0. 1986. Benthic resources assessment technique evaluation of

disposal sites in Puget Sound and adjacent waters. Environmental Lab,

U.S. Army Corps Engineers, Vicksburg, Mississippi. Unpublished

manuscript.

99

Day, D. 1976. Homing behavior and population stratification in Central Puget

Sound English sole (Parophrys vetulus). J. Fish. Res. Board Can. 33:278-

282.

Desbruyers, D., J. Y. Bervas, and A. Khripovnoff. 1980. Un cas de

colonisation rapide d’un sediment profond. Oceonol. Acta 3:285—291.

Dinnel, P. A., D. A. Armstrong, and C. Dungan. 1985a. Initiation of a

Dungeness crab, Cancer magister, habitat study in North Puget Sound. Pp.

327-337 In: Proceedings of the Symposium on Dungeness Crab Biology and

Management. Univ. Alaska, Alaska Sea Grant Rpt. No. 85-3.

Dinnel, P. A., D. A. Armstrong, and R.O. McMillan. 1985b. Survey of

Dungeness crab, Cancer magister, in Oak Harbor, Washington. Final Rpt.

to the Seattle District, U.S. Army Corps of Engineers by School of

Fisheries, Univ. Washington, Seattle. 23 pp.

Dinnel, P. A., D. A. Armstrong, and R.O. McMillan. 1986. Dungeness crab,

Cancer magister, distribution, recruitment, growth and habitat use in

Lummi Bay, Washington, Final Report to Lummi Indian Tribe by Fish. Res.

Institute, University of Washington, Seattle. FRI—Ui—8612. 61 pp.

Dinnel, P. A., D. A. Armstrong, B. S. Miller, and R. F. Donnelly. 1987. U. S.

Navy homeport disposal site investigations. Final Report to Seattle

District, U.S. Army Corps of Engineers by Fish. Res. Institute,

University of Washington, Seattle. (In preparation).

Donnelly, R., B. Miller, R. Lauth, and J. Shriner. 1984. Fish ecology. Vol.

VI, Section 7 In: Renton Sewage Treatment Plant Project: Seahurst

Baseline Study (Q. 3. Stober and K. K. Chew, Eds.). Final report by

University of Washington, Fish. Res. Institute to METRO. FRI—UW—8413.

276 pp.

100

Eaton, C. Skipper of Research Vessel Kittiwake, Seattle, Washington (personal

communication).

English, T. S. 1979. Biological systems acoustical assessments in Port

Gardner and adjacent waters. 1 Sept. 1976 to 30 June 1979. Annual Rpt.

to WDOE, Rep. A79-13. Univ. Washington, Dept. Oceanography. 86 pp.

Grassle, 3. F. 1977. Slow recolonization of deep-sea sediment. Nature,

London. 265:618—619.

Gunderson, D. R., and I. E. Ellis. 1986. Development of a plumb staff beam

trawl for sampling demersal fauna. Fisheries Research 4:35-41.

Lunz, J. D. and D. R.Kendall. 1982. Benthic resources assessment technique,

a method for quantifying the effects of benthic community changes of fish

resources. P. 1021—1027 In Conference Proceedings, Oceans 82.

Malins, D. C., B. B. McCain, D. W. Brown, A. K. Sparks, H. 0. Hodgins, and S.

L. Chan. 1982. Chemical contaminates and abnormalities in fish and

invertebrates from Puget Sound. NOAA Tech. Memo. OMPA—19, 168 pp.

McArn, 0. B., R. 0. Chuinard, B. S. Miller, R. E. Brooks, and S. R. Wellings.

1968. Pathology of skin tumors found on English sole and starry flounder

of Puget Sound, Washington. 3. Nat. Cancer Inst. 41:229—242.

Mearns, A. J., and M. 3. Allen. 1978. Use of small otter trawls in coastal

biological surveys. Contribution No. 66, South. Calif. Water Res.

Project. EPA-600/3-78-083.

Miller, B. S. 1980. Survey of resident marine fishes at Terminals 91 and 73

(Elliott Bay, Seattle, Washington). Univ. of Wash. Coil. of Fish. Tech.

Rept. FRI-UW-8014, 29 pp.

Miller, B. S.., B. B. McCain, R. C. Wingert, S. F. Borton, K. V. Pierce, and D.

T. Griggs. 1977. Ecological and disease studies of demersal fishes in

Puget Sound near METRO-operated sewage treatment plants and in the

101

Duwamish River. Puget Sound Interim Studies Rpt., Univ. of Wash. Coil.

of Fish. FRI-UW-7721, 164 pp.

Miller, B. S., and S. R. Wellings. 1971. Epizootiology of tumors on flathead

sole (Hippoglossoides elassodon) in East Sound, Orcas Island, Washington.

Trans. Am. Fish. Soc. 100:247—266.

Miller, B. S., R. C. Wingert, and S. F. Borton. 1974. Ecological survey of

demersal fishes in the Duwamish River and at West Point. Univ. of Wash.

Coil. of Fish. Tech. Rpt. FRI—UW—7509, 35 pp.

Morton, J. W. 1977. Ecological effects of dredging and dredge spoil

disposal: a literature review. U.S. Fish and Wildlife Serv., Washington,

D.C. Tech. Paper #94.

Orensanz, J. N. and V. F. Gallucci. 1982. Post catastrophic benthic

recovery: process and management implications. Monografias Biologicas

(2): 181—198.

Paquette, R. 0. 1958. A modification of the Wenrie—Smith—Soule salinity

bridge for the determination of salinity in seawater. Tech. Rpt. 61,

Dept. Ocean., Univ. Wash., Seattle.

Pielou, E. C. 1978. Population ecology and community ecology: principles and

methods. Gordon and Breach Sci. Pub., New York, 424 pp.

Saila, S. C., S. D. Pratt, and T. T. Polgar. 1972. Dredge spoil disposal in

Rhode Island Sound. Univ. Rhode Island. Mar. Tech. Rep. 2, 48 pp.

Shepherd, J. C., J. 0. Pope, and R. D. Cousens. 1984. Variations in fish

stocks and hypotheses concerning their links with climate. Rapp. P.—v.

Reun. Cons. mt. Explor. Mer. 185: 255—267.

Smith, R. T. 1937. Observations on the shrimp fishery in Puget Sound.

Washington Department of Fisheries Biol. Rpt. No. 36D, 11+ pp.

Standard Methods. 1980. Standard methods for the examination of water and

102

wastewater, 15th edition. APHA, AWWA, WPCF, Wash. D.C., 1,034 pp.

Stober, Q. J., and K. K. Chew. 1984. Renton Sewage Treatment Plant Project:

Duwamish Head. Final Report by Fisheries Research Institute to the

Municipality of Metropolitan Seattle, WA. Tech. Rpt. FRI—UW-8417, 370

pp.

Tetra Tech, Inc. 1985. Commencement Bay nearshore/tideflats remedial

investigation. Vol. 1. Washington State Department of Ecology and U.S.

Environmental Protection Agency.

Thistle, D. 1981. Natural physical disturbances and communities of marine

soft bottoms. Marine Ecology, Progress Series 6: 223—228.

Washington Department of Ecology. 1976. Ecological baseline and monitoring

study for Port Gardner and adjacent waters. A summary report for the

years 1972 through 1975. WDOE Olympia, WA., 545 pp.

WDF (Washington Department of Fisheries). 1974. 1974 Fisheries Statistical

Report. Olympia, WA., p. 63.

WDF (Washington Department of Fisheries). 1982. 1982 Fisheries Statistical

Report. Olympia, WA., p. 60.

WDF (Washington Department of Fisheries). 1987. Letter from Joseph Blum

(WDF) to Frank Urabeck (Army Corps of Engineers) dated 9 January 1987.

Weitkamp, D. E., D. McEntee, and R. Whitman. 1986. Dungeness crab survey of

Everett Harbor and vicinity, 1984-1985. Final Rpt. to U.S. Navy by

Parametrix, Inc., Bellevue, Washington, 29+ pp.

Weitkamp, D. E., and T. H. Schadt. 1981. Commencement Bay studies technical

studies: Volume III, fish and wetlands. Report to the U.S. Army Corps of

Engineers, Seattle District. Appendices, 75 pp.

Wellings, S. R., C. E. Alpers, B. B. McCain, and B. S. Miller. 1976. Fin

erosion disease of starry flounder (Platichthys stellatus) and English

.99~-LL~:ccP~~Tfl3pieo~sea4~Ta~uo~uzqse~

‘9flW0S‘JGATHT~l9TUiUAflC~inJOi2JetflSGGinUT(snlnq.GAsLxi~doiua)etos

EOT

Ifliva

xia~~aav

~UT

Ap

pe

nd

ixT

ab

le1

.S

ara

tog

aP

ass

ag

ebe

amtr

aw

lsta

tio

nlo

ca

tio

nd

ata

.T

hetr

aw

ld

ep

ths

are

ave

rag

es

fro

mth

ose

reco

rde

dfr

om

the

Fe

bru

ary

and

Jun

e,

1986

cru

ise

s.

All

tra

wls

we

re1

/8n

au

tica

lm

iles

(NM

)lo

ng

at

ap

pro

xim

ate

ly1

.5kn

ots

gro

un

dsp

ee

d.

F-’

C Li’

STA

RT

SET

OF

NET

TRAW

LD

EPTH

(m)

Ap

pro

xim

ate

De

pth

com

pass

Sta

tio

n(m

)R

ad

ar

Ran

ges

(NM

)an

dM

ark

ers

be

arin

gS

tart

tow

End

tow

PSD

DA

Site

:

Sta

tio

n1

113

2.4

Lo

we

llP

t./1

.25

Sa

rato

ga

Pt.

2000

108

106

Sta

tio

n2

106

1.4

E.

Pt.

/1.2

5S

.E

dge

Pt.

N.

of

Mab

ana

25

°11

011

8S

tatio

n3

109

1.1

E.

Pt.

/1.2

Pt.

of

Cam

ano

N.

of

Mab

ana

300

113

125

Tra

nse

ct

#1

:

low

ii1.

565

.E

dge

E.

Pt.

2900

1017

20W

201.

56S

.E

dge

E.

Pt.

2900

2418

40W

391

.55

E.

Pt.

2900

3746

80W

791.

55E

.P

t.28

5°80

6380

E80

2.25

S.

Edg

eL

ow

ell

Pt.

/l.6

3S

ara

tog

aP

t.28

0°80

7940

E39

2.2

05.

Edg

eL

ow

ell

Pt.

/1.7

1S

ara

tog

aP

t.28

0°45

4620

E21

2.25

S.

Edg

eL

ow

ell

Pt.

/l.8

1S

ara

tog

aP

t.26

0°20

391O

E11

2.13

Lo

we

llP

t.29

0°14

11

Tra

nse

ct#

2:

lOO

N10

21

.25.

Edg

eL

ow

ell

Pt.

/0.5

5P

t.N

.o

fM

aban

a26

0°10

210

011

5N11

71.

65L

ow

ell

Pt.

/1.1

5E

.P

t.24

0°12

112

112

0N12

11.

1S

.E

dge

Lo

we

llP

t./l.2

E.

Pt.

265°

122

116

Ap

pe

nd

ixT

ab

le2

.S

um

ma

ryo

fP

ort

Ga

rdn

er

beam

tra

wl

sta

tio

nlo

ca

tio

nd

ata

,A

llbe

amtr

aw

lto

ws

we

re1

/8n

au

tica

lm

ile

(NM

)lo

ng

at

ap

pro

xim

ate

ly1

.4kn

ots

gro

un

dsp

ee

d.

Th

eo

tte

rtr

aw

lsw

ere

fish

ed

for

1/5

NM

at

asu

bse

to

fth

ese

sta

tio

ns.

Tra

nse

ct

sta

tio

ns

are

ide

ntifie

db

yth

ea

pp

roxim

ate

de

pth

inm

ete

rsa

nd

are

ao

fth

eb

ay

wh

ere

:N

=n

ort

h,

Sso

uth

an

dN

=m

idd

le.

Nav

yC

ADS

ite

(80

m):

Sta

tio

n1

Sta

tio

n2

Sta

tio

n3

Sta

rtE

ndto

wto

w

Com

pass

he

ad

ing

(de

gre

es

ma

gn

etic)

1100

1100

1100

Sta

rtS

tatio

n#

se

to

fn

et

De

pth

(m)

Ran

gem

ark

ers

and

dis

tan

ce

(NM

)

(at

sta

rto

fn

et

se

t)

8480

800

.65

8280

850

.60

8175

830

.54

PS

DD

AS

ite

2

Site

(liO

n):

ma

rke

rm

ark

er

ma

rke

r

4/0

.72

4/0

.72

4/0

.70

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

Ap

pro

xim

ate

wire

ou

t(ft

.)

105

110

112

0.9

no

tch

W.

Pie

r/i

.0ed

geS

.P

ier

113

117

118

0.9

2n

otc

hW

.o

fS

.P

ier/

1.2

5SW

.co

rne

rS

.P

ier

124

127

116

0.8

5bu

mp

W.

of

sho

ren

otc

h/i

.5SW

.co

rne

rS

.P

ier

(A)

105

108

110

0.9

2SW

.co

rne

rS

.P

ier/

0.7

5sh

ore

(5)

103

105

107

0.8

5SW

.co

rne

rS

.P

ier/

0.7

5no

bon

sho

re(I

)11

012

212

91

.25

SW.

co

rne

rS

.P

ier/

i.1

5sh

ore

at

no

tch

(J)

119

114

115

1.4

0SW

.co

rne

rS

.P

ier/

1.2

5sh

ore

at

no

tch

NAV

YR

ADC

AD(S

tatio

n1

~S

tatio

n2

(~S

tatio

n3

Sta

tio

n4

Sta

tio

n5

Sta

tio

n6

Sta

tio

n7

PS

DD

AS

ite

1S

tatio

n1

Sta

tio

n2

Sta

tio

n3

Tra

nse

ct

#1lo

s20

S40

S60

S80

S1O

OM 80N

60N

40N

130

133

2~

I013

413

52

.30

139

140

2.5

0

Ma

rke

r4

/1M

ark

er

4/i

Ma

rke

r4

/1

(13

0rn

):12

813

313

6 12 21 41 60 83 101 82 62

.50

Ed

ge

wa

ter

.50

Ed

ge

wa

ter

.50

Ed

ge

wa

ter

110

115

130

145

145

145

305

135

305

140

145

145

145

200 40 35 35 40

250

80 75

C

1212

0.5

0Sw

.co

rne

rS

.23

220

.70

SW.

co

rne

rS

.45

420

.80

SW.

co

rne

rS

.62

600

.82

SW.

co

rne

rS

.82

780

.88

SW.

co

rne

rS

.10

210

30

.83

Ma

rke

r4

/0.4

580

760

.94

Ma

rke

r4

/1.5

060

600

.92

Ma

rke

r4

/1.5

0S

tatio

nd

rop

pe

ddu

eto

rep

ea

ted

ne

tda

mag

e.

1150

1150

1200

1150

1100

1200

1150

1350

1350

1380 250

350

600

800

1000

1100

1050

80

0

Pie

rP

ier

Pie

rP

ier

Pie

rSW

.co

rne

rS

.P

ier

Da

rlin

gto

nD

arlin

gto

n

Ap

pe

nd

ixT

ab

le2

(co

nt.

)

De

pth

Cm

)S

tart

tow

Ap

pro

xim

ate

wire

ou

t(it.)

Tra

nse

ct

#3

:10

820

S40

S60

380

S11

OS

130M

130N

Tra

nse

ct

1/4:

105

205

403

603

80S

11O

S14

5S13

5N

Tra

nse

ct

#5

:20

340

560

380

311

0316

5314

5M

S.

edge

S.

Pie

rSW

.co

rne

rS

.P

ier

S.

edge

S.

Pie

rSW

.co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

S.

ed

ge

S.

Pie

rM

ark

er

4/1

.35

Ed

ge

wa

ter

Ma

rke

r4

/2.0

8E

dg

ew

ate

r

At

tip

àf

fue

ld

ock

3.1

1SW

.co

rne

rS

.P

ier

3.1

33

.12

3.1

91

.92

1.8

3

65 70 75 70 75 65 145

145 40 40 35 35 40 45

210

140 60 40 45 45 45

280

125

250

350

600

800

1050

1200

1300

1350 250

350

600

800

1050

1175

1350

1500

350

600

80

010

5012

0016

0015

00

Sta

rtS

tatio

n#

se

to

fn

et

End tow

(at

sta

rto

fn

et

se

t)

Ran

gem

ark

ers

and

dis

tan

ce

(NM

)C

ompa

ssh

ea

din

g

Tra

nse

ct

#2

:10

520

S40

S60

$80

$lb

SIi

ON

130N

1OO

N

(de

gre

es

ma

gn

etic)

1.2

5SW

.co

rne

rS

.P

ier

1.2

5SW

.co

rne

rS

.P

ier

1.2

0SW

.co

rne

rS

.P

ier

1.2

0SW

.co

rne

rS.

Pie

r1.

10SW

.co

rne

rS

.P

ier

1.14

SW.

co

rne

rS.

Pie

r1

.35

Ma

rke

r4

/1.3

8D

arlin

gto

n3

.5R

an

da

ll/i

.6S

E.

Ge

dn

ey/

2.O

NWS

.P

ier

1.3

5G

ed

ne

y/2

.OM

ark

er

4/1

.0“R

BN

”

260

250

250

250

240

230

155

120

11.0

250

350

600

800

1000

1150

1150

1300

1050

11 22 40 62 82 112

111

136

102 12 23 41 61 80 112

129

137 11 22 41 62 79 113

145

140 21 42 61 81 119

171

151

12 20 40 65 81 110

110

134 95 12 22 42 60 82 116

132

131 12 22 41 58 85 112

147

138 24 38 64 80 118

169

152

12 40 52 70 80 110

111

125 84 10 21 46 67 82 116

134

130 11 21 42 62 83 123

145

139 22 52 43 82 106

170

151

1.8

81

.87

1.8

51

.89

1.8

61

.88

2.0

92

.08

2.5

72

.57

2.6

32

.65

2.6

82

.70

2.4

52

.26

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

Ra

nd

all

Pt.

/2.2

0E

dg

ew

ate

r/0

.85

“RB

N’

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

SW.

co

rne

rS

.P

ier

ne

are

st

sho

reW

hid

be

y/0

.63

tip

fue

ld

ock

Ed

ge

wa

ter/

1.8

3sh

ore

Clin

ton

Doc

k

Ap

pe

nd

ixT

ab

le2

(co

nt.

)

Dep

thCm

)S

tart

tow

Com

pass

he

ad

ing

(de

gre

es

ma

gn

etic

)

Ap

pro

xim

ate

wire

ou

t(f

t.)

Tra

nse

ct_

#6

:80

S80

M60

N40

N20

NiO

N

Tra

nse

ct#7

:10

05IO

OMlO

ON

8ON

6ON

40N

2ON

iON

165

280

260

260

250

260

220

145

155

120

130

320

300

310

1050

1050 800

600

350

250

1100

1100

1100

1050 800

600

350

250

Sta

rtS

tatio

n#

set

of

ne

tEn

dto

w

Ran

gem

arke

rsan

dd

ista

nce

CNN)

(at

sta

rto

fn

et

se

t)

7484

640

.33

SW.

tip

S.

Pie

r80

8071

0.5

0M

ark

er

4/0

.50

SW.

corn

er

S.P

ier

6361

560

.40

Ma

rke

r4

4042

420

.35

Ma

rke

r4

/0.7

0SW

.co

rne

rS

.P

ier

2121

210

.37

Ma

rke

r4

1211

110

.35

Ma

rke

r4

102

9889

0.8

0SW

.co

rne

rS

.P

ier

101

102

105

1.2

0M

ark

er

4/1

.26

SW.

corn

er

S.

Pie

r10

210

110

11.

72~

RB

N’/0

.33

Ma

rke

r3

8081

641

.5‘R

BN

”/0

.53

Ma

rke

r3

5662

691

.45

‘RB

N”

4243

331

.5M

ark

er

4/1

.50

“RB

N”

2229

161

.60

“RB

N’

1114

121

.60

‘RBN

~’~

No

n-T

ran

sect

Sta

tio

ns:

C

B10

510

611

41.

25M

arke

r4

/1.1

0n

ea

rest

sho

reE

.o

fn

otc

h31

011

00C

9291

921

.00

Sw.

corn

er

S.

Pie

r/i

.34

sho

reD

arlin

gto

n80

1050

D10

410

310

11

.30

SW.

corn

er

S.

Pie

r/i

.46

Da

rlin

gto

n33

511

00F

110

109

113

0.6

5SW

.co

rne

rS.

Pie

r/0

.40

sho

rea

tT

ran

sect

125

011

00C

138

128

127

1.7

0M

arke

r4

/1.5

0sh

ore

at

Edg

ewat

er14

513

50Fl

130

128

128

1.9

0M

ark

er

4/1

.50

sho

rea

tE

dgew

ater

145

1300

Ap

pe

nd

ixT

ab

le3

.E

llio

ttB

aybe

amtr

aw

lsta

tio

nlo

ca

tio

nd

ata

.T

hetr

aw

ld

ep

ths

are

ave

rag

es

fro

mth

ose

reco

rde

dfr

om

the

Fe

bru

ary

,Ju

ne

and

Se

pte

mb

er,

1986

cru

ise

s.

All

tra

wls

we

re1

/8n

au

tica

lm

iles

(NM

)lo

ng

at

ap

pro

xim

ate

ly1.

5kn

ots

gro

un

dsp

ee

d.

Q

STAR

TSE

TO

FN

ETTR

AWL

DEP

TH(m

)

Ap

pro

xim

ate

De

pth

com

pass

Sta

tio

n(m

)R

ad

ar

Ran

ges

(NM

)an

dM

ark

ers

be

arin

gS

tart

tow

End

tow

Fo

urm

ileR

ock

:

Sta

tio

n1

178

1.0

5S

ho

re/i

.57

SWT

ipP

ier

917

90

168

160

Sta

tio

n2

168

0.8

1S

ho

re/1

.1SW

Tip

Pie

r91

23

5°

178

181

Sta

tio

n3

163

0.8

5S

ho

re/i

.0SW

Tip

Pie

r91

22

0°

185

188

Inn

er

Ellio

ttB

ay:

Sta

tio

n1

910

.7S

ho

reD

.H

ea

d/i

.2NW

Co

rne

r#4

61

80

°91

90S

tatio

n2

680

.84

Sh

ore

D.

He

ad

/i.0

NWC

orn

er

#46

16

0°

6262

Sta

tio

n3

780

.98

Sh

ore

D.

He

ad

/0.8

3NW

Co

rne

r#4

61

30

°78

73S

tatio

n4

660

.25

Tip

Tod

dD

ryd

ock/0

.84

NWC

orn

er

#46

65

°64

60S

tatio

n5

811

.6SW

Pie

r9

1/0

.95

Duw

.H

ead

(ne

are

st)

31

0°

9112

4

Tra

nse

ct

#1

lOS

120

.5N

otc

h/i

.28

Alk

i1

85

°12

1320

S21

0.5

No

tch

/i.2

3A

lki

19

0°

2120

40S

390

.5N

otc

h/i

.34

Alk

i1

85

°43

3780

S82

1.3

Alk

iP

t.1

75

°80

82

Tra

nse

ct

#2

:

iON

121.

1SW

Tip

Pie

r91

1000

20N

210

.9SW

Tip

Pie

r91

28

5°

2121

SON

870

.95

SWT

ipP

ier

9129

0~72

68

Ap

pe

nd

ixT

ab

le4

.C

omm

ence

men

tB

aybe

amtr

aw

lsta

tio

nlo

ca

tio

nd

ata

.T

hetr

aw

ld

ep

ths

are

ave

rag

es

fro

mth

ose

reco

rde

dfr

om

the

Fe

bru

ary

,Ju

ne

and

Se

pte

mb

er,

1986

cru

ise

s.

All

tra

wls

we

re1

/8n

au

tica

lm

iles

(NM

)lo

ng

at

ap

pro

xim

ate

ly1.

5kn

ots

gro

un

dsp

ee

d.

STA

RT

SET

OF

NET

TRAW

LD

EPTH

(m)

Ap

pro

xim

ate

De

pth

com

pass

Sta

tio

n(m

)R

ad

ar

Ran

ges

(NM

)an

dM

ark

ers

be

arin

gS

tart

tow

End

tow

PSD

DA

Site

1:

Sta

tio

n1

168

0.9

8A

sarc

oC

orn

er/

i.8

2N

eill/i

.35

Br.

Pt.

12

5°

169

166

Sta

tio

n2

169

1.1

8A

sarc

oC

orn

er/

2.O

Ne

ill/1

.15

Br.

Pt.

1400

169

168

Sta

tio

n3

173

1.4

Asa

rco

Co

rne

r/i

.8N

eill/1

.OB

r.P

t.14

0016

816

8S

tatio

n4

171

1.5

8A

sarc

oC

orn

er/

i.8

2N

eill/O

.82

Br.

Pt.

1300

169

167

PSD

DA

Site

2:

Sta

tio

n1

163

1.3

5A

sarc

oN

otc

h/O

.65

Sh

ore

30

5°

167

170

Sta

tio

n2

159

1.5

Asa

rco

No

tch

/2.6

3N

eill/1

.i5

Br.

Pt.

30

5°

166

166

Sta

tio

n3

158

1.6

5A

sarc

oN

otc

h/2

.62

Ne

ill/0

.93

Br.

Pt.

30

5°

166

166

PSD

DA

Site

2B:

Sta

tio

n1

175

1.2

5A

sarc

oC

orn

er/

i.5

Ne

ill/1

.3B

r.P

t.85

~17

417

3S

tatio

n2

175

1.5

Asa

rco

Co

rne

r/i.i2

Br.

Pt.

1000

175

174

Sta

tio

n3

175

1.7

4A

sarc

oC

orn

er/

i.0

Br.

Pt.

10

5°

175

173

Tra

nse

ct

#1

:

iOE

120

.08

Br.

Pt.

16

5°

ii15

20E

220

.09

Br.

Pt.

16

5°

2023

40E

400

.09

Abe

amB

r.P

t.1

65

°38

4880

E81

0.2

2A

beam

Br.

Pt.

18

0°

8675

40W

410

.5A

sarc

oC

orn

er

30

0°

6040

20W

210

.47

Asa

rco

Co

rne

r3

00

°22

20lo

w10

0.4

5A

sarc

oC

orn

er

29

0°

ii10

Ap

pe

nd

ixT

ab

le4

(Co

nt.

)

STAR

TSE

TO

FN

ETTR

AWL

DEP

TH(in

)

Ap

pro

xim

ate

De

pth

com

pass

Sta

tio

n(m

)R

ad

ar

Ran

ges

(NM

)an

dM

ark

ers

be

arin

gS

tart

tow

End

tow

Tra

nse

ct

#2

:

lOS

140

.48

End

of

Sitc

um

Wa

terw

ay

1200

1315

20S

220

.28

Co

rne

rP

ier

540

025

2040

S42

0.3

8C

orn

er

Pie

r5

600

3336

8OS

821

.0N

.S

ho

re(E

.o

fB

r.P

t.)/

O.9

5S

.S

ho

re2

15

°89

9812

0S12

01

.0S

ho

reO

ldT

aco

ma

/i.0

Fr

“B”

(Hyle

bo

s)

20

5°

120

120

112

Appendix Table 5. Dungeness crab catches/hectare in Saratoga Passage duringFebruary and June, 1986. The averages listed in the tableare means +1 standard deviation. The station numbers forthe transects indicate approximate trawl depth in metersand location where N = north, E = east and W = west.N.S. = not sampled.

Dungeness Crab Catch/Hectare

February JuneStation Beam Trawl Beam Trawl Otter Trawl

PSDDA Site 2 (11OM):

Station 1 0 0 0Station 2 0 0 0Station 3 0 0 0Station 4 0 N.S. N.S.

Average 0 0 0

Transect #1:

1OE 11.2 449.4 N.S.20E 0 0 4.540E 0 56.2 080E 0 37.5 080W 1.9 18.7 N.S.40W 0 18.7 020W 0 0 N.S.lOW 1.9 18.7 N.S.

Average 1.9 ± 3.9 74.9 ± 152.5 1.1 ± 2.2

Transect #2:

lOON N.S. 0 N.S.120N N.S. 0 N.S.115N N.S. 0 N.S.

Average 0

Saratoga Pass Average 1.2 ± 5.2 42.8 ± 118.3 0.6 ± 1.7

Ap

pe

nd

ixT

ab

le6

.B

eam

tra

wl

shrim

pca

tch

es/h

ecta

rein

Sa

rato

ga

Pa

ssa

ge

du

rin

gF

eb

rua

ryan

dJu

ne

,1

98

6,

and

inE

llio

ttan

dC

omm

ence

men

tb

ays

du

rin

gF

eb

rua

ry,

Jun

ean

dS

ep

tem

be

r,1

98

6.

The

ave

rag

es

liste

din

the

tab

lea

rem

eans

±1

sta

nd

ard

de

via

tio

n.

The

sta

tio

nn

um

be

rsfo

rth

etr

an

se

cts

ind

ica

tea

pp

roxim

ate

de

pth

sin

me

ters

and

loca

tio

nw

he

reN

No

rth

,E

=E

ast,

W=

Wes

tan

dS

=S

ou

th.

N,S

.=

no

tsa

mp

led

.

FEBR

UAR

YJU

NE

Sm

ooth

Sm

ooth

Aro

a/S

tntio

nS

po

tS

ide

atr

ipe

00

0n

atr

ipe

pin

kP

ink

Rom

pkao

kA

llS

po

tS

ide

atr

ipe

00

0n

atr

ipe

pin

kP

ink

llnm

pkao

kA

ll

SAR

ATO

GA

PASS

AGE

PSSS

AS

ite

2(lIO

N):

Sta

tio

nI

01

8.7

09

.41

8.7

04

6.8

Sta

tio

n2

03

7.5

00

00

37

.5S

tatio

n3

03

7.5

00

18

.70

56

.2

Ave

rag

e0

31

.2•

10

.90

3.1

•5

.41

2.5

+1

0.8

04

6.8

+9

.4

Tra

na

eo

t#

1:

IOE

00

00

00

020

E0

00

00

00

40E

00

09

3.6

56

.20

14

9.8

80

k0

00

00

00

ROW

00

00

00

040

W0

00

93

.61

87

.30

28

0.9

20W

00

00

00

0lO

W0

00

00

00

00

01

8.7

74

.90

93

.60

37

.50

07

4.9

01

12

.40

00

00

0-

0

01

2.5

•2

1.7

06

.2•

10

.84

9.9

•4

3.2

06

8.7

•6

0.2

Tra

nn

eo

t#

2:

lOO

NN

.S.

1208

8.5

.ll5

N8

.5.

Sa

rato

ga

Ray

AV

erag

e0

8.5

•1

5.4

ELL

IOT

TRA

TF

oo

rnila

Roo

k(1

7014

):S

tatio

nI

00

Sta

tio

n2

05

6.2

Sta

tio

n3

01

8.7

Ave

rag

e0

25

.0+

28

.6

00

37

.50

00

00

00

00

00

00

00

00

37

.50

05

6.2

20

6.0

00

00

00

00

00

00

00

0

03

7.5

00

00

O0

02

99

.70

00

00

0

O3.

70

02

54

.70

24

.34

00

00

00

00

00

00

0~

01

7.9

+3

7.5

25

.5•

56

.40

51

.9+

88

.63

.4•

11

.34

.0•

10

.82

.7•

10

.05

.4•

15

.54

1.5

78

.50

56

.2+

99.1

0 0 0 0

00

00

05

6.2

01

8.7

32

.4

N.S

.

Inn

er

Ellio

ttB

ay(7

514)

:S

tatio

n1

0S

tatio

n2

0S

tatio

n3

0S

tatio

n4

11.5

.S

tatio

n5

~A

vera

g,

0

18

.70 0

6.2

•1

0.8

O3

7.5

02

62

.20

37

.5

01

12

.41

29

.7

00

05

6.2

07

4.9

04

3.7

~3

9.0

24

3.5

26

2.2

39

3.3

30

0.0

+8

1.6

0 0 0 0

18

7.3

00

03

55

.80

18

1.0

+1

78

.00

07

4.9

01

68

.5

2_1

87

.3

04

3.6

60

.2

O1

8.7

07

4.9

00

03

1.2

03

9.0

00

03

7.5

00

18

.70

0N

.S.

N.S

.

18

.7+

18

.80

Tra

nm

eo

tN

I10

520

S40

S80

S

Tra

nn

ao

t#

2:

ION

208

808

SEPT

EMBE

R

Sm

ooth

Sp

ot

Sid

en

trip

eC

oo

na

trip

ep

An

tP

ink

Rom

pkao

kAS

S

09

3.6

02

43

.40

18

7.3

01

74

.8+

75

.7

0 0 08

.5.

00

00

R.S

.

03

7.5

00

03

7.5

O2

5.0

02

1.7

00

00

00

00

00

01

8.7

00

0

31

8.4

00

03

18

.47

4.9

00

07

4.9

o9

3.6

09

3.6

18

.70

37

.50

07

5.0

00

18

.73

7.5

07

4.9

05

6.2

56

.20

__

__

_0

0

01

8.7

+1

8.8

43

.7+

47

.10

81

.2•

10

.81

5.0

•2

4.4

11

.2•

25

.1

00

00

00

00

00

00

00

00

18

.70

00

00

00

00

18

.70

18

.70

37

.4

00

18

.70

00

_9_

-~

37

.4

00

18

.71

8.7

00

11

2.4

01

8.7

00

07

8A

.50

18

.74

49

.40

07

4.9

07

.5•

10

.22

84

.63

29

.3

Ellio

ttB

ayA

vera

ge

08

.5•

17

.53

7.5

09

5.9

54

.5•

11

5.0

013

1.1

a1

44

.88

.6o

18.1

1.4

05

.24

4.7

*6

6.9

06

6.2

a7

6.7

—3

0.7

•7

8.2

——

—7

.2•

21

.0—

4.3

•11

.2—

—

05

6.2

00

07

4.9

04

3.7

+3

9.0

013

1.1

01

8.7

08

42

.71

8.7

54

3.0

07

4.9

3.7

+8

.4 32

2.0

+3

56

.9

00

00

00

00

00

00

00

00

00

55

.90

00

00

55

.95

6.2

0

8.7

•1

9.9

00

56

.20

00

56

.20

09

3.6

00

09

3.6

00

00

00

00

00

01

8.7

01

8.7

00

18

.70

00

11

.2+

27

.22

.5•

6.6

00

00

01

8.7

37

.40

93

.6

10

2.4

•2

21

.21

34

.8+

23

6.9

8.7

•1

8.6

1.7

•5

.0

00000000jooo

oo0~000

0000

0

I,

0

—

~

00

0000

0

CD

x

CD

ON

C

CDCD-.

0jooooo0oo~.~no~.000

j’~’0~00000000~0

I~00000010000000

1+joo~oo00000ooo0000

0

~oo~0

~P~0

g00000coo0000

~

I.I,I.

~P~-.0

~0000000000000Ieee~I~~

~I°°°°°00000000coo0000

~00000oooooo~0eec0I000

I:~00000000boo0000

~-‘000000000000-~‘~~D-~0

‘-.0

~1°°°°°000000000000000

I~~~

~

P000o000000000000o0ooj...o0000

0ft.0

000000000000‘“o~o—oo~0II0P

—.0~0

‘~ooo~~o~ooo~~0000010000

~o~~o00000

I~~I~~

~000000000000010000

-.‘_,4

I.~~°~°Is-.~

~7TT

Ap

pe

nd

ixT

ab

le7

.N

umbe

ro

fsh

rim

pca

ug

ht/

he

cta

reb

yo

tte

rtr

aw

lin

Sa

rato

ga

Pa

ssa

ge

inJu

ne

and

Ellio

ttan

dC

omm

ence

men

tB

ays

inJu

ne

and

Se

pte

mb

er

19

86

.T

hetr

an

se

ct

sta

tio

nn

um

be

rsin

dic

ate

de

pth

inm

ete

rsan

dlo

ca

tio

nw

he

reE

=E

ast

and

W=

We

st.

N~

.N

ot

Sa

mp

led

.

Sp

ot

Sid

estr

ipe

Co

on

str

ipe

Sm

ooth

Pin

kH

umpb

ack

Fle

xe

dS

tatio

nP

raw

nS

hrim

pS

hrim

pP

ink

Sh

rim

pS

hrim

pS

hrim

pS

hrim

p

Sa

rato

ga

Pa

ssa

ge

,Ju

ne

19

86

:

Ellio

ttB

ay,

Jun

e1

98

6:

PSD

DA

1,

Sta

.1

PSD

DA

1,

Sta

.2

PSD

DA

1,

Sta

,3

PSD

DA

2,

Sta

.1

PSD

DA

2,

Sta

.2

PSD

DA

2,

Sta

.3

Ave

rag

e(±

1S

.D.)

12

6.1

4.5

4.5

26

1.3

63

.14

0.5

11

7.1

PSD

DA

2,

Sta

.1

05

4.0

00

63

.10

0PS

DD

A2

,S

ta.

20

18

.00

06

3.1

00

PSD

DA

2,

Sta

.3

09

0.1

00

90

.10

0T

ran

se

ct

1,

40W

00

00

00

0T

ran

se

ct

1,

20E

00

00

00

0T

ran

se

ct

1,

40E

00

00

4,5

00

Tra

nse

ct

1,

80E

04

.50

09

9.1

00

Ave

rag

e(±

1S

.D.)

02

3.8

±3

5.2

00

45

.7±

43

.40

36

.00

03

01

.80

18

.00

00

00

04

.52

7.0

00

02

2,5

00

09

.0-~

6.8

±1

4.4

12

.8±

11

.50

50

,3±

12

3.2

Li’

0 0 0 0 0 0 0 0

0 0 0 0 0

Ellio

ttB

ay,

Se

pte

mb

er

19

86

:

PSD

DA

1,

Sta

.1

40

.51

3.5

PSD

DA

1,

Sta

.2

N.S

.N

.S.

PSD

DA

1,

Sta

.3

00

PSD

DA

1,

Sta

,4

19

3.7

00

PSD

DA

1,

Sta

.5

18

.01

8.0

0PS

DD

A2

,S

ta.

19

.01

3.5

0

10

2.1

±9

0.5

0.8

±1

.8

00

25

2.3

9.0

0N

.S.

N.S

.N

.S.

N.S

.N

.S.

00

56

7.6

00

01

,74

7.7

22

.50

9.0

68

0.2

00

00

00

Ap

pe

nd

ixT

ab

le7

(co

ntin

ue

d): S

po

tS

ide

str

ipe

Co

on

str

ipe

Sm

ooth

Pin

kH

umpb

ack

Fle

xe

dS

tatio

nP

raw

nS

hrim

pS

hrim

pP

ink

Sh

rim

pS

hrim

pS

hrim

pS

hrim

p

PSD

DA

2,

Sta

.2

PSD

DA

2,

Sta

.3

Ave

rag

e(±

1S

.D.)

Com

men

cem

ent

Ba

y,Ju

ne

19

86

:

o0

o0

o1

.3±

3.4

45

.00

31

.50

47

4.9

±6

22

.84

.5±

8.6

Com

men

cem

ent

Ba

y,S

ep

tem

be

r1

98

6:

04

.51

03

.67

6.6

38

.0±

70

.13

2.2

±4

0.9

PSD

DA

1,

Sta

.1

4.5

9.0

04

.52

20

.70

PSD

DA

1,

Sta

.2

04

.50

03

42

.30

PSD

DA

1,

Sta

.3

00

04

.52

43

.20

PSD

DA

2B,

Sta

.1

04

.50

01

30

.60

PSD

DA

2B,

Sta

.2

02

2.5

00

10

8.1

0PS

DD

A2B

,S

ta.

30

13

.50

04

36

.90

Tra

nse

ct

1,

20E

00

00

00

Tra

nse

ct

1,

40E

00

00

00

Ave

rag

e0

.6±

1.6

6.8

+8

.0

0 0 0 0 0 0 0 0 0 0 0 00

1.1

±2

.11

85

.2±

15

6.0

0

PSD

DA

1,

Sta

.1

N.S

.N

.S.

N.S

.N

.S.

N.S

.N

.S.

N.S

.PS

DD

A1

,S

ta.

20

76

.60

03

10

.80

0PS

DD

A1

,S

ta.

39

,08

5.6

00

37

8.4

00

PSD

DA

1,

Sta

.4

09

4.6

00

41

4.4

00

PSD

DA

2B,

Sta

.1

09

9.1

00

38

7.4

00

PSD

DA

2B,

Sta

.2

4.5

67

.60

02

97

.30

0PS

DD

A2B

,S

ta.

30

16

2.2

00

40

9.9

00

Tra

nse

ct

1,

20E

00

00

00

0T

ran

se

ct

1,

40E

00

00

00

0

Ave

rag

e(±

1S

.D.)

1.7

±~

74

.1±

52

.10

02

74

.8±

17

4.8

00

App

endi

xT

able

8.Be

amtr

aw

lca

tche

so

fD

unge

ness

crab

(per

l000

m2)

from

Por

tG

ardn

erdu

ring

win

ter

(Feb

ruar

y4

-7,

1986

).

Dun

gene

ssC

rab/

1000

m2

~Fe

mal

esFe

mal

esS

tatio

n*

with

eggs

with

ou

teg

gsM

ales

All

crab

sCo

mm

ents

Nav

yD

ispo

sal

Site

:(8

0m)

Sta

tion

19.

41.

90

11.2

60g

al.

mud

,w

ood,

de

bris

Sta

tion

228

.10

028

.150

ga

l.m

ud,

woo

d,tr

ash

Sta

tion

324

.31.

91.

928

.130

ga

l.w

ood,

tra

sh

Ave

rage

20.6

±.9•

9+1

.3~

1.1

0.6

±1.

122

.5~

9.8

PSDD

AS

ite2

:(1

20m

)

Sta

tion

10

00

05

ga

l.w

ood,

de

bris

Sta

tion

20

00

02

ga

l.fish

,d

etr

itu

sS

tatio

n3

1.9

00

1.9

2g

al.

fish

,d

etr

itu

s

Ave

rage

0.6

±1.

10.

00.

00.

6±

1.1

PSDD

AS

ite1

:(1

60m

)

Sta

tion

10

00

01

ga

l.w

ood,

de

tritu

sS

tatio

n2

00

00

1g

al.

worm

tube

s,d

etr

itu

sS

tatio

n3

00

00

1g

al.

worm

tube

s

Ave

rage

0.0

0.0

0.0

0.0

App

endi

xT

able

8(c

on

tinu

ed

)

2.4

±3

5

+2.

1-

3.1

1.9

±1.

9

Dun

gene

ssC

rab/

l000

m2

Fem

ales

Fem

ales

Sta

tion

*w

itheg

gsw

itho

ut

eggs

Mal

esA

llcr

abs

Com

men

ts

Tra

nsec

t#1

10-s

20-S

40-S

80-S

100-

M80

-N40

-N

Ave

rage

Tra

nsec

t#2

:

0 1.9

11.2

20.6

1.9

7.5

11.8

7.8

i:7.

3

1.9 0 3.7 0 0 1.9 0

1.1

±1.

5

13.1 5.6 0 0 0 0 0

15.0

7.5

15.0

20.6

1.9

9.4

11.8

alga

e,d

etr

itu

sal

gae,

de

tritu

sal

gae,

wood

alg

ae

,w

ood,

tra

shal

gae,

woo

d,d

etr

itu

sw

ood,

de

tritu

sfish

;be

ntbe

am,

torn

net

crab

s10

—S

1.9

7.5

20-S

5.6

5.6

40

-s1.

90

80—

S31

.83.

712

0-S

00

l20-

M0

0l5

0-N

1.9

010

0-N

1.9

0

1g

al.

2ga

l15

ga

l.20

ga

l.2

ga

l.3

ga

l.2

ga

l.

4g

al.

25g

al.

15g

al.

25g

al.

25g

al.

2g

al.

2g

al.

3g

al.

20.6

5.6 0 0 0 0 0

30.0

16.8

1.9

35.5 0 0 1.9

1.9

Ave

rage

Tra

nsec

t#3

:

20-S

40

-s80

-5l2

0-~

l50-

MiS

O-N

H H cc

2.7

±5.

011

.6±

6.1

alg

ae

,ee

lgra

ss~

rock

,sh

ell

woo

d,cr

abs

mud

,w

ood,

de

tritu

swo

rmtu

bes,

de

tritu

sw

ood,

de

tritu

s,

she

llw

ood,

de

tritu

s+

++

5.6

-10

.73.

3-

7.3

11

.0-

14.6

03.

77.

511

.28

ga

l.a

lga

e,

ee

lgra

ss,

she

ll0

1.9

01.

920

ga

l.w

ood,

tra

sh3.

70

03.

725

ga

l.w

ood,

sta

rfis

h,

bo

ttle

s3.

73.

70

7.4

7g

al.

woo

d,sta

rfis

h,

bo

ttle

s9.

43.

71.

915

.030

ga

l.cl

ay

ba

lls,

rock

,wo

od0

00

02

ga

l.w

ood,

worm

tube

s,0

00

01

ga

l.wo

rmtu

bes

1.3

i:2.

85.

6±

5.8

Ave

rage

App

endi

xT

able

8(c

on

tinu

ed

)

Tra

nsec

t#5

:

wood

chip

sw

ood,

alga

ew

ood,

sta

rfis

h,

tra

shd

etr

itu

s,

fish

worm

tube

s,d

etr

itu

swo

rmtu

bes,

he

art

urc

hin

s

~D

unge

ness

Cra

b/l0

00ni

2Fe

mal

esFe

mal

esS

tatio

n*

with

eggs

with

ou

teg

gsM

ales

All

crab

sCo

mm

ents

Tra

nsec

t#4

:

10-S

20-S

40-S

80-5

120—

S17

5—s

160-

N

Ave

rage

030

.0 1.9

16.9 5.6 0 1.9

011

.2 0 0 0 0 0..

1.9

13.1 1.9

1.9 0 0 0

1.0

54.3 3.8

18.8 5.6 0 1.9.

2ga

l5

ga

l.2

ga

l.5

gal

5g

al.

2g

al.

1g

al.

8.0

±11

.31.

6±

4.2

2.7

±4.

712

.3±

19.5

rock

,al

gae,

and

larg

etire

alga

e,w

ood,

she

llw

ood,

sta

rfis

h,

bo

ttle

sg

rave

l,w

ood,

bo

ttle

sg

rave

l,d

etr

itu

swo

rmtu

bes,

de

tritu

swo

rmtu

bes

20-S

01.

95.

67.

540

-S1.

90

3.7

5.6

80-S

5.6

1.9

07.

512

0—S

9.4

1.9

011

.320

0-S

00

00

l75-

M0

00

0

Ave

rage

2.8

~3.

9

Tra

nsec

t#6

:

100 30 40 4 2 2

ga

l.g

al.

ga

l.g

al.

ga

l.g

al.

1.0

j1.

01.

6+

1.5

5.3

+4.

5

80-5

18.7

18.7

027

.480

_rvi

45.0

3.7

048

.740

-N11

.213

.19.

433

.720

-N3.

70

5.6

9.3

lU-N

3.7

9.4

7.5

20.6

Ave

rage

16.5

±7.

19.

0±

7.4

4.5

±4.

329

.9±

15.3

50g

al.

10g

al.

25ga

l.2

gal.

2ga

l.

woo

d,w

ood,

woo

d,w

ood,

wood

,

tra

shC

aflS

tra

shd

eb

ris

bo

ttle

s

App

endi

xT

able

8(c

on

tinu

ed

)

Dun

gene

ssC

rab/

1000

m2

Fem

ales

Fem

ales

Sta

tion

*w

itheg

gsw

itho

ut

eggs

Mal

esA

llcr

abs

Com

men

ts

Tra

nsec

t#7

:

100-

S9.

40

09.

415

ga

l.w

ood,

de

tritu

s,

cans

lOO

-M1.

90

01.

93

ga

l.w

orm

tube

s,w

ood

100-

N5.

60

05.

62

ga

l.w

ood,

de

bris

80-N

8.5

00

8.5

2g

al.

gra

vel,

crab

s40

-N54

.33.

71.

959

.920

ga

l.w

ood,

crab

s20

-N43

.17.

51.

952

.510

ga

l.w

ood,

crab

s10

—N

7.5

13.1

1.9

22.5

2g

al.

crab

s

Ave

rage

18.6

~20

.93.

5~

5.1

0.8

±1

.022

.9±

23.7

*S

tatio

nnu

mbe

rsfo

rth

etr

an

sect

sin

dic

ate

whe

reN

=nor

th,

S=s

outh

,an

dM

~mid

dle

dept

hin

met

ers

plus

loca

tion

r”J C

+Mea

nj

1st

anda

rdd

evi

atio

n.

App

endi

xT

able

9.D

unge

ness

Cra

bD

en

sitie

sPe

rH

ecta

reC

alcu

late

dFr

omBe

amTr

awl

Cat

ches

InPo

rt~G

ardn

erD

urin

gA

pril

,19

86.

Sta

tion

1Fe

mal

esM

ales

All

Cra

bsS

ubst

rate

Com

men

ts

Nav

yD

ispo

sal

Site

(80m

)S

tatio

n1

477

1949

620

gal

.W

ood,

De

bris

,B

ott

les

Sta

tion

243

90

439

20g

al.

Woo

d,B

ark,

Cans

Sta

tion

322

90

229

10g

al.

Woo

d,S

he

ll,D

ebris

Ave

rage

382

~13

426

1138

814

1

PSDD

AS

ite2

(ilO

m)

Sta

tion

138

038

5ga

l.

Woo

d,D

etr

itus

Sta

tion

219

019

30ga

lW

ood

Sta

tion

30

00

3g

al.

Woo

dC

hips

Ave

rage

1919

0.0

1919

PSDD

AS

ite1

(130

m)

Sta

tion

10

00

2g

al.

Woo

d,D

etr

itus,

Wor

mTu

bes

Sta

tion

20

00

1g

al.

De

tritu

s

Sta

tion

30

00

1ga

l.

Wor

mTu

bes

Ave

rage

0.0

0.0

0.0

Tab

leA

ppen

dix

9.D

unge

ness

Cra

bD

en

sitie

sP

erH

ecta

reC

alcu

late

dFr

omBe

amTr

awl

Cat

ches

(co

ntin

ue

d)

InP

ort

Gar

dner

Dur

ing

Ap

ril,

1986

.

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Com

men

ts

Tra

nsec

t#1

1g

al.

ULVA

,W

ood

Chi

ps

1g

al.

iJULP

~~W

ood

Chi

ps

6g

al.

iJ.L~

(~.

Woo

dC

hips

10g

al.~

Woo

d,B

ott

les

12g

al.

Woo

d,M

ud2

ga

l.D

etr

itus,

Woo

dC

hips

40-N

__

__

__

__

__

__

__

__

__

__

__

__

__

Ave

rage

Tra

nsec

t#2

10-S

5g

al.

Zos

tera

,UL

VA

20-S

20ga

l.

ULVA

40-S

4ga

l.

ULVA

,

80-S

8g

al.

Woo

d,

110-

S5

ga

l.W

ood,

110-

M2

ga

l.D

etr

itus

130-

N2

ga

l.W

ood,

De

tritu

s,Ca

ns

100-

N_

__

__

__

__

__

__

__

__

__

__

1ga

l.

Woo

d

Ave

rage10

-S38

3876

20-S

1919

38

40-S

190

19

80-S

950

95

100-

M0

00

80-N

570

N.S

.3N

.S.

N.S

.

38

~3

41

0~

164

8~

36

5719

76

570

57

380

38

760

76

00

0

00

0

00

0

190

19

31~

3O2

t7

N.S

1’J

Woo

d,S

hell

Deb

ris,

Cans

De

trit

us

Tab

leA

ppen

dix

9.D

unge

ness

Cra

bD

en

sitie

sP

erH

écta

reC

alcu

late

dFr

omBe

amTr

awl

Cat

ches

(co

ntin

ue

d)

InP

ort

Gar

dner

Dur

ing

Ap

ril,

1986

.

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Com

men

ts

Tra

nsec

t#3

10-S

3819

576

ga

l.UL

VA,

Sh

ell,

Bo

ttle

s

20-S

380

3840

gal

.UL

VA,

Woo

d

40-S

115

011

530

ga

l.W

ood,

Roc

k,B

ott

les

80-S

950

957

ga

l.W

ood

110-

S0

00

20g

al.

Cla

yB

alls

,W

ood

130-

rio

00

1g

al.

De

tritu

s,W

orm

Tube

s

130-

N19

019

1ga

l.

Wor

mTu

bes

Ave

rage

44t

453

746

45

Tra

nsec

t#4

10-S

1938

572

ga

l.Z

oste

ra,

ULVA

,S

hell

20-5

00

0:

ga

l.~JJ

JL~,

Woo

d,S

hell

40-S

190

1950

ga

l.UL

VA,

Woo

d,B

ott

les

80-S

191

1921

03

ga

l.UL

VA,

Woo

d,Ca

ns

110-

S19

019

7g

al.

Roc

k,W

ood,

Gra

vel

145-

50

00

2g

al.

De

tritu

s,W

orm

Tube

s

135-

N0

—0

02

ga

l.W

ood,

De

tritu

s,W

orm

Tube

s

Ave

rage

35

t69

8~

15

44

~7

6

Tra

nsec

t#5

20—

S57

057

30g

al.

ULVA

,T

ire

,W

ood,

Sh

ell,

40-S

1919

3850

ga

l.W

ood

Chi

psG

rave

l

80-S

115

011

560

ga

l.R

ock,

Gra

vel,

Bo

ttle

s

110-

S0

00

3g

al.

Mud

,Pe

aG

rave

l,W

ood

165-

S0

00

2g

al.

Woo

dC

hips

,W

orm

Tube

s

145M

IL_

__

_2

ga

l.H

eart

Urc

hin

s,W

orm

Tube

sA

vera

ge32

±46

33

35±

46

Tab

leA

ppen

dix

9.D

unge

ness

Cra

bD

en

sitie

sPe

rH

ecta

reC

alcu

late

dFr

omBe

amTr

awl

Cat

ches

(co

ntin

ue

d)

InP

ort

Gar

dner

Dur

ing

Ap

ril,

1986

.

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Com

men

ts

Tra

nsec

t#6

80-S

191

019

160

ga

l.UL

VA,

Woo

d,S

hell

80-M

573

1959

225

ga

l.W

ood,

De

bris

,B

ott

les

40-N

248

5730

520

ga

l.W

ood,

Bo

ttle

s

20-N

210

1922

94

ga

l.W

ood,

De

tritu

s

10-N

229

1924

82

ga

l.W

ood

Chi

ps

Ave

rage

290

160

2321

313

161

Tra

nsec

t#7

100-

S57

057

4g

al.

ULVA

,W

ood

Chi

ps

100-

M38

038

2g

al.

De

tritu

s,W

orm

Tube

s

100-

N38

038

2g

al.

De

tritu

s

80-N

760

762

ga

l.D

etr

itus,

Woo

dC

hips

40-N

153

015

33

ga

l.W

ood

Chi

ps

20-N

114

011

44

ga

l.W

ood

10—

N19

1933

0.1

ga

l.D

etr

itus,

She

ll

Ave

rage

7147

.73

773

45

Gra

ndA

vera

ge79

122

612

8512

7’

1S

tatio

nnu

mbe

rsfo

rth

etr

an

s~ct

sin

dic

ate

dept

hin

met

ers

plus

loca

tion

whe

reN

=N

orth

,M

=M

iddl

e,S

Sou

th

2M

ean

1S

tand

ard

Div

iatio

n

N.S

.=

Not

Sam

pled

Ap

pe

nd

ixT

ab

le1

0.

Du

ng

en

ess

cra

bG

ard

ne

rd

urin

gd

en

sitie

sp

er

he

cta

reca

lcu

late

dfr

om

beam

tra

wl

ca

tch

es

inP

ort

Jun

e,

19

86

.

De

nsity/H

ecta

re1

Fe

ma

les

Ma

les

All

cra

bs

Su

bstr

ate

com

men

tsS

tatio

n

Na

vyD

isp

osa

lS

ite

(80

m)

Sta

tio

n1

Sta

tio

n2

Sta

tio

n3

375

543

431

245

0:~:

86

47 75 37

53÷

20

421

618

46

8

502

~10

3A

vera

ge

PSD

DA

Site

2(h

Orn

)

Sta

tio

n1

Sta

tio

n2

Sta

tio

n3

Ave

rag

e

40g

al.

50g

al.

10g

al.

1g

al.

1g

al.

1g

al.

woo

d,ca

ns,

cra

bs,

de

bris

woo

d,ca

ns,

cra

bs

woo

d,cra

bs,

de

bris

wo

od

,d

etr

itu

s

wor

mtu

be

s,

woo

d

woo

d,d

etr

itu

s

00

0

00

0

00

0

00

0

(13

0m

)PS

DD

AS

ite

1

Sta

tio

n1

Sta

tio

n2

Sta

tio

n3

Ave

rag

e

“3

0 0 0 0

0 0 0 0

0 0 0 0

2g

al.

wor

mtu

be

s,

de

trita

lke

lp

1g

al.

wor

mtu

be

s

2g

al.

wor

mtu

be

s

Ap

pe

nd

ixT

ab

le10

(Co

ntin

ue

d)

Fe

ma

les

Ma

les

All

Cra

bs

Com

men

ts

Tra

nse

ct

#1

10—

S0

5656

3g

al.

Ulv

a,

woo

d

20—

S19

3756

20g

al.

Ulv

a

40-S

190

1910

ga

l.w

ood,

Ulv

a

80-S

169

016

96

ga

l.w

ood,

Ulv

a

100-

M0

00

2g

al.

de

tritu

s

80—

N91

80

918

8g

al.

de

tritu

s

40-N

N.S

.3N

.S.

N.S

.

Ave

rag

e18

8÷

364

16+

2520

3+

355

Tra

nse

ct

#2

10

-519

019

15g

al.

Ulv

a,

woo

d

20

-S19

019

20g

al.

Ulv

a,

woo

d

40

-S19

019

5g

al.

woo

d,sh

ell,

Ulv

a

80—

594

094

20g

al.

wo

od

,g

rave

l

110—

511

20

112

10g

al.

pea

gra

ve

l,

110—

M19

019

2g

al.

IJlv

a,

woo

d,

130—

N37

037

2g

al.

woo

dch

ips,

10

0-N

560

561

ga

l.w

ood

ch

ips,

Ave

rag

e47

÷37

047

÷37

woo

d

wor

mtu

be

s

wor

mtu

be

s

de

tritu

s

Ap

pe

nd

ixT

ab

le10

(Co

ntin

ue

d)

Fe

ma

les

Tra

nse

ct

#3

Ma

les

All

cra

bs

Com

men

ts

10—

50

00

3g

al.

woo

d,U

lva

,sh

ell

20

-S37

037

25g

al.

woo

dch

ips,

Ulv

a

40—

S56

1975

15g

al.

woo

dch

ips

80—

S56

056

5g

al.

woo

dch

ips,

can

s,b

ott

les

110—

537

037

8g

al.

cla

yb

alls

130-

M0

00

1g

al.

woo

d,w

orm

tub

es

130—

N~0

00

2g

al.

wo

od

,w

orm

tub

es,

ke

lp

Ave

rag

e27

÷26

3+

729

±30

Tra

nse

ct

#4

10—

S19

3756

3g

al.

Ulv

a,

de

tritu

s

20—

S0

3737

15g

al.

woo

d,U

lva

,sh

ell

40—

S56

1975

30g

al.

woo

d,b

ott

les

80

-S75

075

4g

al.

woo

d,ca

ns

110—

S37

1956

2g

al.

pea

gra

ve

l,w

ood

145—

S0

00

2g

al.

wor

mtu

be

s,

woo

d

135—

N0

~0

01

ga

l.w

orm

tub

es,

he

art

urc

hin

s

Ave

rag

e27

±30

16+

1~1

43±

32

Ap

pe

nd

ixT

ab

le10

(Co

ntin

ue

d)

Fe

ma

les

Ma

les

All

cra

bs

Com

men

ts

Tra

nse

ct

#5

20—

S94

3713

1

40—

S75

075

80—

S56

056

110—

S19

019

16

5-S

00

0

20g

al.

pea

gra

ve

l,U

lva

40g

al.

woo

d,b

ott

les,

gra

ve

l,d

eb

ris

25g

al.

wo

od

,b

ott

les,

gra

ve

l

3g

al.

wor

mtu

be

s,

woo

d,pe

ag

rave

l

2g

al.

wor

mtu

be

s,

woo

d,h

ea

rtu

rch

ins

2g

al.

wor

mtu

be

s,

he

art

urc

hin

s14

5-M

00

0

Ave

rag

e41

+40

6÷

1547

±51

Tra

nse

ct

#6

80

-S56

1975

80—

M26

20

262

40

-N20

60

206

20

-N28

119

300

10

-N11

219

131

Ave

rag

e18

3±

9711

÷10

195

±92

50g

al.

15g

al.

20g

al.

4g

al.

3g

al.

woo

d

woo

d,ca

ns

woo

d

woo

dch

ips

bo

ttle

s,

woo

d

Ap

pe

nd

ixT

ab

le10

(Co

ntin

ue

d)

Tra

nse

ct

#7

100—

S10

452

156

10g

al.

wo

od

,ca

ns

100-

M0

00

2g

al.

wor

mtu

be

s,

woo

dch

ips

100—

N44

919

468

4g

al.

de

tritu

s

80—

N20

60

206

2g

al.

woo

dch

ips,

can

s

40—

N22

519

244

10g

al.

woo

dch

ips

20—

N33

70

337

7g

al.

woo

d

10—

N19

019

1/2

ga

l.d

etr

itu

s

Ave

rag

e19

1+

165

13+

1920

4+

167

GRAN

DA7

ERAG

E10

4+

171

10÷

1811

4+

178

1S

tatio

nn

um

be

rsfo

rth

etr

an

se

cts

ind

ica

ted

ep

thin

me

ters

plu

slo

ca

tio

nw

he

reN

no

rth

,M

=

mid

dle

,an

dS

so

uth

.

2M

ean

-~1

sta

nd

ard

de

via

tio

n.

-~N

.S.

=n

ot

sam

ple

d.

App

endi

xTa

ble

II.

Dunq

enes

scr

abde

nsiti

espe

rhe

ctar

eca

lcul

ated

from

beam

traw

lca

tche

sin

Por

tG

ardn

erdu

ring

Sept

embe

r,19

86.

Sta

tion

num

bers

for

the

tran

sect

sin

dica

tede

pth

inm

eter

spl

uslo

catio

nw

here

N=N

orth

,M

=Mid

dle,

and

S=So

uth.

The

aver

ages

are

mea

ns+

1st

anda

rdde

viat

ion.

Den

sity

/Hec

tare

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Sub

stra

teCo

,mne

nts

Nav

yD

ispo

sal

Site

(80m

)

Sta

tion

1S

tatio

n2

Sta

tion

3

00

570

190

9520

ga

l.w

ood,

debr

is11

510

ga

l.w

ood,

debr

is19

1g

al.

woo

d,de

bris

15g

al.

wood

1g

al.

worm

tube

swo

odch

ips

2g

al.

woo

d,sh

ell

1g

al.

worm

tube

s,sh

ell

1g

al.

worm

tube

s,sh

ell

0.5

ga

l.wo

rmtu

bes,

wood

Ave

rage

Tra

nsec

t#1

25±

290

25

±2

9

10-S

20-S

40-S

00-S

100-

M80

-N40

-N

Ave

rage

1938

5719

191

024

819

950

00

—no

tsa

mpl

ed

57 76 191

267 95 0

114

±9~

Sta

tion

1S

tatio

n2

Sta

tion

3

Ave

rage

95 115 19

0 0 0

PSDD

A2

(hO

rn)

Sta

tion

1S

tatio

n2

Sta

tion

3

Ave

rage

PSDD

A1.

(130

m)

76

+5

10

76

+5

1

•19

019

190

190

00

13+

110

13±

11

C

0 57 19

3g

al.

alga

e,w

ood,

de

tritu

s

30g

al.

alga

e,wo

od15

ga

l.w

ood,

alga

e20

ga

l.w

ood,

debr

is5

ga

l.w

ood,

debr

is

102±

991

3±

16

Ap

pe

nd

ixT

ab

leii

(co

ntin

ue

d)

Den

sity

/Hec

tare

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Sub

stra

teCo

mm

ents

Tra

nsec

tff2

10—

S19

1938

1g

al.

alga

e,d

etr

itus

20—

S21

019

249*

15g

al.

alga

e,sh

ell

40—

S15

30

153

15g

al.

alga

e,wo

od80

-S30

50

305

25g

al.

woo

d,al

gae,

cla

yb

alls

110-

S0

00

3g

al.

de

tritu

s,al

gae

wood

ll0-M

380

381

ga

l.wo

rmtu

bes,

wood

rhip

s13

0—N

380

381

ga

l.d

etr

itu

s,

she

ll10

0-N

00

02

ga

l.wo

odch

ips

Ave

rage

95÷

114

5+

910

3±

119

Tra

nsec

tff3

10—

S0

1919

6g

al.

alga

e,sh

ell

20-5

380

3850

ga

l.w

ood,

alga

e40

—S

553

1957

230

ga

l.ba

rk80

-S95

095

8g

al.

rock

,al

gae,

de

tritu

s11

0—5

570

573

ga

l.w

ood,

alga

e13

0-M

760

761

ga

l.wo

rmtu

bes,

woo

d,sh

ell

130-

N19

019

1g

al.

worm

tube

s

Ave

rage

120

+19

45

±9

125

±19

9

Tra

nsec

tff4

10—

S19

019

1g

al.

alga

e,sh

ell

20—

538

3876

6g

al.

alga

e,w

ood,

she

ll40

-S17

238

210

30g

al.

wood

chip

s,b

ott

les

80—

S11

50

115

4g

al.

woo

d,al

gae,

cans

110—

515

30

153

4g

al.

de

tritu

s,w

ood,

grav

el14

5—S

3819

572

ga

l.al

gae,

worm

tube

s13

5-N

00

01

ga

l.wo

rmtu

bes,

wood

chip

s

Ave

rage

76+

6914

÷18

90÷

75

App

endi

xT

able

ii(c

on

tinu

ed

)

De

nsi

ty/H

ect

are

Sta

tion

Fem

ales

Mal

esA

llC

rabs

Sub

stra

teCo

mm

ents

Tra

nsec

t#5

20—

S76

5713

320

ga

l.al

gae,

gra

vel,

woo

d,sh

ell

40-S

496

5755

330

ga

l.w

ood,

rock

,al

gae

80—

S95

1911

440

ga

l.w

ood,

alga

e,ro

ck,

de

bris

110—

S15

30

153

3g

al.

woo

d,d

etr

itu

s16

5-S

00

01

ga

l.wo

rmtu

bes

145-

M0

00

1g

al.

worm

tube

s

Ave

rage

137

±18

622

÷28

159

±20

4

Tra

nsec

t#6

80-S

7619

9550

ga

l.al

gae,

woo

d,ca

ns80

-M19

10

191

20g

al.

woo

d,d

eb

ris,

cans

40-N

760

7610

ga

l.w

ood,

de

bris

20—

N19

019

2g

al.

woo

d,d

etr

itu

s10

-N38

038

1g

al.

de

tritu

s,

woo

d

Ave

rage

80±

674

±8

84

+6

7

App

endi

xT

able

ii(co

ntin

ue

d)

Den

sity

/He

cta

reS

tatio

nFe

mal

esM

ales

All

Cra

bsS

ubst

rate

Com

men

ts

Tra

nsec

t#7

100—

S76

076

40g

al.

wood

chip

s,b

ott

les,

cans

lOO

-M38

038

2g

al.

wood

chip

s10

0-N

00

02

ga

l.w

ood,

de

tritu

s80

-N21

019

229

3g

al.

woo

d,ca

ns40

-N22

90

229

1g

al.

woo

d,d

etr

itu

s,

she

ll20

-N95

095

4g

al.

woo

d,sh

ell

10-N

760

760.

5g

al.

de

tritu

s,

she

ll

Ave

rage

103

±85

3±

710

6-I-

89

GRAN

DAV

ERAG

E92

+11

38

÷15

100

÷11

9

~Fri~

TiTJ

é~1

youn

g—of

—th

e-ye

ar(u

nsex

ed)

crab

,9.

0mm

cara

pace

wid

th.

App

endi

xT

able

12.

Dun

gene

sscr

abd

en

sitie

spe

rhe

ctar

eca

lcu

late

dfr

ombe

amtr

aw

lca

tche

sat

ext

rast

atio

ns

inP

ort

Gar

dner

du

ring

Sep

tem

ber,

1986

.Th

eav

erag

esar

em

eans

±1

stan

dard

de

via

tion

.

Den

sity

/He

cta

reS

tatio

nFe

mal

esM

ales

All

Cra

bsS

ubst

rate

Con

nien

ts

Eas

to

fPS

DDA

Site

1.

Sta

tion

G(1

30m

)S

tatio

nH

(130

m)

Bet

wee

nM

uki

lteo

and

Pic

nic

Poi

nt

Wes

to

fN

avy

Site

Sta

tion

A(1

05m

)19

019

8g

al.

wood

chip

sS

tatio

nB

(llO

m)

00

01

ga

l.wo

rmtu

bes,

wood

Sta

tion

C(

90m

)38

038

1g

al.

de

tritu

s,

wood

chip

sS

tatio

n0

(lO5m

)38

038

1g

al.

de

tritu

s,

wood

chip

sS

tatio

nE

(115

rn)

00

01

ga

l.wo

rmtu

bes,

wood

Sta

tion

F(h

Orn

)38

038

7g

al.

woo

d,d

eb

ris

Ave

rage

22±

190

22±

19

190

193

ga

l.w

ood,

she

ll0

00

4g

al.

woo

d,sh

ell

Sta

tion

1-

40m

190

193

ga

l.w

ood,

de

tritu

sS

tatio

n2

-40

m0

00

5g

al.

woo

d,al

gae

Sta

tion

3—

40m

00

05

ga

l.w

ood,

alga

e,b

ott

les

Sta

tion

4—

lOm

00

010

ga

l.sa

nd,

alga

eS

tatio

n4

-20

m38

038

3g

al.

alga

eS

tatio

n4

-40

m0

00

5g

al.

cla

yb

alls

,al

gae

Sta

tion

4-

80m

00

020

ga

l.cl

ay

ba

lls,

alga

e

8÷

158

+15

Ave

rage

0

135Appendix Table 13. Commercial shrimp densities per hectare

in Port Gardner calculated from beam trawlcatches in February and April, 1986.

STATION 1 FEBRUARY APRIL

Navy Disposal Site (80m)

Station 1 95 0

Station 2 1,069 0

Station 3 935 0

Average 700 ~ 528 o

PSDDA Site 2 (ilOm)

Station 1 76 19

Station 2 76 0

Station 3 95 19

Average 82 11 13 11

PSDDA Site 1 (130m)

Station 1 0 38

Station 2 0 95

Station 3 0 57+Average 0 63 - 29

Transect #1

10-S 0 0

20-S 0 0

40-S 57 0

80-S 57 38

100-M 19 57

80-N 248 0

40-N 95 N.S.3

Average 68 87 16

Transect #2

10-S 0 0

20-S 95 19

40-S 0 0

136

Appendix Table 13. Commercial shrimp densities per hectarein Port Gardner calculated from beam trawl

(Cont’d.) catches in February and April, 1986.

STATION 1 FEBRUARY APRIL

Transect #2

80-S 229 0

110—S 57 191lO-M 38 19

130-N 76 19

100-N 267 19

Average 95 100 12 10

Transect #3

10-s 0 0

20-S. 0 0

40-S . 0 0

80-S . 0 0

110-S 134 57

130—M 76 .19

130-N 57 19

Average 38 53 14 21

Transect #4

10-S 95 0

20-S 0 . 0

40-S 19 0

80-S 191 0

110—S 115 115

• 145—S 76 38

135-N 95 0

Average - 84 63 22 + 43

137

Appendix Table 13. Commercial shrimp densities per hectarein Port Gardner calculated from beam trawl

(Cont’d.) catches in February and April, 1986.

STATION 1 FEBRUARY APRIL

Transect #5

10-S 229 0

40-S 897 38

80—S 153 115

110-S 38 30

165-S 38 19

145-N 0 38

Average 226 340 40 ± 39

Transect #6

80-S 57 38

80-M 172 0

40-N 76 19

20-N 19 0

10-N 0 0.

Average 65 67 11 + 17

Transect #7

100-S 267 0

100-N 57 19

100-N 210 38

80-N 195 0

40-N 0 0

20-N 19 0

10-N 0 0

Average 107 113 8 15

Grand Average 123 + 218 19 + 28

1 Station numbers for the transects indicate depth in meters plus locations where

N North, M = Middle, S South.

peLdw~S~°NSN

UOL3~LALOp~A~pu~sI-

‘9861‘[pd~pu~A~AenJqe~UI(p1~uo~)[M~J~w~qwo~jP~P~flDLPDU~~P9~°dUL

a~r~~sei~isuepdwi~i~sL~D~WWODIe[qPJx~puedd~

App

endi

xTa

ble

14.

De

nsi

ties

per

hect

are

of

Dun

gene

sscr

abs

and

com

mer

cial

shrim

pca

lcu

late

dfr

omo

tte

rtr

awl

catc

hes

from

the

Feb

ruar

yan

dA

pril

cru

ise

sin

Por

tG

ardn

er.

Feb

urar

y19

86A

pril

1986

Sta

tion

’C

ance

rm

apis

ter

Com

mer

cial

shrim

pC

ance

rm

agis

ter

Com

mer

cial

shrim

p

Nav

yD

ispo

sal

Site

(SO

m)

Sta

tion

1S

tatio

n2

Sta

tion

3

Ave

rage

Sta

tion

1S

tatio

n2

Sta

tion

3

Ave

rage

20-S

40-S

100-

N

27 9 27

21±

io2

5 18 9

239

108

216

188

±70

505

410

149 86 1,62

158

135

~43

0 54 441

0 14 32

15~

6

9 0 5

50 36 27

38±

12

36 27 9

24±

14

0 0 27

PSDD

AS

ite2

(hO

rn)

Sta

tion

1S

tatio

n2

Sta

tion

3

Ave

rage

.

PSDD

AS

ite1

(130

m)

90 131

117

113

±21

0 0 0 0 5 0 0

2t3

355

~18

4

Tra

nsec

t1

U)

0 0 0~

0 0 0 0 0

Ave

rage

11±

71

65

±2

41

5±

59

±16

App

endi

xT

hble

14

(co

n’t).

Feb

ruar

y19

86A

pril

1986

Sta

tion

1C

ance

rm

agis

ter

Com

mer

cial

shrim

pC

ance

rm

agis

ter

Com

mer

cial

shrim

p

Tra

nsec

t2

20—

S18

05

540

-SiS

00

011

0—S

099

927

Ave

rage

12±

1033

±57

5±

5ii

±14

Tra

nsec

t4

C

20-S

90

00

40-5

50

90

145-

S9

225

018

Ave

rage

8±

275

±13

03

±5

6~

10

Gra

ndA

vera

ge9

±9

158

~15

85

~8

33±

40

‘Sta

tion

num

bers

for

the

tra

nse

cts

ind

ica

tede

pth

inm

eter

spl

uslo

catio

ns

whe

reN

=nor

th,

M=m

iddl

e,an

dS

=sou

th.

2Mea

n±

1st

anda

rdd

evi

atio

n.

Station1

Navy Site (8Cm)

Station 1

Station 2

Station 3

Average

PSODA Site

Station

Station

Station

Ave rage

PSDDA Site

Station

Station

Station

Average

Transect #1

20-S

40-S

100-M

Average

Transect #2 -

20-S

40-S

110—S

Average

9 0 9

14 4 18

0 0 0

7÷72 1±2 9+9

4 0 4

0 0 0

0 0 0

1÷2 0 1+2

1

0 0 0

2 0 0 0

3 -~ -~ -~

0 0 0

0 0 0

0 4 4

18 4 22

6÷10 3÷2 9~12

•0 0 0

4 0 4

14 0 14

6÷7 0 6÷7

141

Appendix Table 15. Dungeness crab densities per hectare calculated from ottertrawl catches in Port Gardner in June and early July,1986.

Density/HectareFemales Males All crabs

2 (11Cm)

2

3

(13Cm)

142

Appendix Table 15 (Continued)

Station Females Males All crabs

Transect #4

20-S 0 0 0

40-S 0 0 0

145-S 0 0 0

Average 0 0 0

Grand Average 4 ÷ 6 1 ÷ 2 4 ÷ 7

1 Station numbers for the transects indicate depth in meters plus

locations where S = south and M = middle.

2 Mean ÷ 1 standard deviation.

143

19 9

o 0

o 42

6±11 4:~_5

o 228

o 41

o 23

o 117+148

o 131

19 59

o 50

6÷11 80±44

0 N.S.

19 O~

19 0

75 N.S.

0 221

0 N.S.

____ N.S.

19÷29 ~74÷ 128

Appendix Table 16. Commercial shrimp densities per hectare calculatedfrom beam and otter trawis in Port Gardner in Juneand early July, 1986.

Density/HectareBeam trawl Otter trawlStation1

Navy Disposal Site (80m)

Station 1

Station 2

Station 3

Average

PSDDA Site 2 (hOrn)

Station 1

Station 2

Station 3

Average

PSDDA Site I (130m)

Station 1

Station 2

Station 3

Average

Transect #1

10-S

20—S

40-S

80-S

1 O0-M

80-N

40-N

Average

144

Appendix Table 16 (Continued)

Station Beam trawl Otter trawl

Transect #2

10-S 0 - N.S.

20—S 19 0

40—S 19 0

80-S 0 N.S.

110—S - 75 27

110-M 0 N.S.

130—M 0 N.S.

100-N 0 N.S.

Average 14 + 26 9 ~ 16

Transect #3

10-S 0 N.S.

20-S 0 N.S.

40-5 0 N.S.

80-S 0 N.S.

110-5 0 N.S.

130-M 19 N.S.

130-N - 0 N.S.

Average --

Transect #4

10-S 0 N,S.

20-S 0 0

40-S 0 4

80-S 0 N.S.

110-S 0 N.S.

145

Appendix Table 16 (Continued)

Station Beam trawl Otter trawl

Transect #4 — Continued V

145-S ~0 36

135—N N.S.

Average 5 j. 14 13 + 20

Transect #5

20-S 0 N.S.

40-S 787 N.S.

80-S 281 N.S.

110—S 19 N.S.

165—S 19 N.S.

145-M 0 N.S.

Average 184 ÷ 315

Transect #6

80—5 112 N.S.

80-M 19 N.S.

40-N 19 N.S.

20-N 0 V N.S.

10-N 0 NS

Average 30 + 47 V --

Transect #7

100-S 0 V N.S.

100-M 0 N.S.

100—N V 56 N.S.

80-N 19 V N.S.

40-N 0 N.S.

Appendix Table 16 (Continued)

146

1 Station numbers for the transects indicate depth in meters pluslocations where N north, M = middle, and S = south.

2 Mean + standard deviation.

N.S. = not sampled.

Station

Transect #7 — Continued

Beam trawl Otter trawl

20—N 0 N.S.

~ 10—N 0 N.S.

Average 11 4- 21 -—

Grand Average 30 ÷ 112 50 -~- 82

147

Appendix Table 17. Shrimp densities per hectare calculated from both beam andotter trawl catches in Port Gardner during September, 1986.Station numbers for the transects indicate depths in metersand location where N = North, S = South, E = East, and W =

West. The averages are means + 1 standard deviation.N.S. = not sampled. Estimated crab densities are also givenfor the otter trawl.

Beam Trawl Otter Trawl

Station Shrimp/hectare Shrimp/hectare Crab/hectare

Navy Disposal Site (80 m)

Station 1 581 387 5Station 2 169 536 0Station 3 131 405 0

Average 294 ÷ 250 443 + 81 2 ÷ 3

PSDDA~ Site 2 (110 m)

Station 1 19 77 14Station 2 0 72 18Station 3 0 108 27

Average 6 ± 11 86 ÷ 20 20 + 7

PSDDA Site 1 (130 m)

Station 1 19 81 9Station 2 38 104 14Station 3 38 117 5

Average 32 + 11 101 ± 18 12 + 4

Transect #1

10-S 0 N.S. N.S.20-S 375 0 040-S 1760 5 980-S 375 N.S. N.S.

100-N 187 198 080-N 131 N.S. N.S.40-N ~ N.S. N.S.

Average 471 + 648 68 ± 113 3 ± 5

148

Transect #2

1 0-S20-S40-S80-S

110—S1 l0-M1 30-N1 00-N

Transect #3

10—S20-S40-S80-S

110-S1 30-M1 30-N

Transect #4

10-S20—S40-S80-S

110-S145—S135—N

Transect #5

20-S40-S80-S

110-S165—S145-M

Average

0300356730

38193838

00

131206

387556

056

0755656

0

0150936131

075

N.S.68

N.S.N.S.N.S.

23 + 39

N.S.N.S.N.S.N.S.N.S.N.S.N.S.

NS.N.S.N.S.N.S.N.S.N.S.

N.S.1818

N.S.5

N.S.N.S.N.S.

14 ± 8

N.S.N.S.N.S.N.S.N.S.N.S.N.S.

N.S.95

N.S.N.S.

0N.S.

5+5

N.S.N.S.N.S.N.S.N.S.N.S.

Appendix Table 17 (cont.)

Beam Trawl Otter Trawl

Station Shrimp/hectare Shrimp/hectare Crab/hectare

N.S.00

190 + 258Average

Ave rage 72 ± 74

N.S.05

N.S.N.S.

45N.S.

17 + 2555 t~ 33

Average 215 ~i:~ 359

149

Transect #6

6551292

24300

N.S.N.S.N.S.N.S.N.S.

N.S.N.S.N.S.N.S.N.S.

Transect #7

100_S100-M100-N80-N40- N20-N10-N

262412393

10493127

00

749 :~:~ 1106

N.S.N.S.N.S.N.S.N.S.N.S.N.S.

N.S.N.S.N.S.N.S.N.S.N.S.N.S.

Appendix Table 17. (cant.)

Beam Trawl Otter Trawl

Station Shrimp/hectare Shrimp/hectare Crab/hectare

80-S80-M40-N20-N10-N

438 .i: 547Average

Average

Port Gardner Average 269 ± 527 123 ÷ 159 9+8

150

Appendix Table 16. Shrimp densities/heCtare calculated from both beamand otter trawl catches at extra stations in PortGardner during September, 1986. The averages aremeans ÷ 1 standard deviation. N.S. = not sampled.

Shrimp Density/Hectare

Station Beam trawl Otter trawl

West of Navy Site

Station A (105 m) 19 N.S.Station B (110 in) 0 N.S.Station C (90 m) 94 N.S.Station D (105 in) 75 N.S.Station E (115 m) 38 68Station F (110 m) 94 N.S.

Average 53 ÷ 40 68 ÷ 0

East of PSDDA Site 1

Station G (130 m) 38 N.S.Station H (130 in) 19 N.S.

Average 28 + 13

Between Mukilteo and Picnic Point

Station 1 (40 m) 0 N.S.Station 2 (40 m) 0 N.S.Station 3 (40 m) 0 N.S.Station 4 (10 m) 0 N.S.Station 4 (20 m) 0 N.S.Station 4 (40 m) 0 N.S.Station 4 (80 m) 0 N.S.

Average 0

IIniva

XI~IN~~dV

tci

152

APPENDIX A

Analyses of fish data collected by

beam trawla in Commencement Bay, Elliott Bay

and Saratoga Passage during 1986.

153

Commencement Bay

Abundance and biomass. The samples collected during the summer cruise

were the only ones available for analyses. Abundance values ranged from 1 to

43 individuals per location, while biomass values ranged from 45 to 4,191

grams per location (Appendix A, Table 1). At locations where fish were

captured, the two PSDDA sites had by far the lowest value of either measure.

Sample sizes were too small to determine dominant species at each site.

Species richness. Species richness values ranged from 1 to 8 showing a

similar pattern to the abundance and biomass results. The deeper PSDDA sites

had the lowest values. The 20 m station had a much higher value (Appendix A,

Table 1).

Species diversity. Species diversity values ranged from 0.0 to 0.71

(Appendix A, Table 1). The distribution of values among sites was identical

to species richness. The deeper PSDDA sites had the lowest values, while the

20-m station had the highest value.

Elliott Bay

Abundance and biomass. Total abundance ranged from 5 to 22 fish per

location, and total biomass ranged from 69 to 1 ,595 grams per location

(Appendix A, Table 2). The results did not suggest any patterns, seasonally

or by station.

Species

A, Table 2).

biomass, and

Species

(Appendix A,

location was

richness. Species richness values ranged from 2 to 6 (Appendix

The distribution among locations was similar to abundance and

no seasonal or location patterns were evident.

diversity. Species diversity values ranged from 0.2 to 0.9

Table 2), and as with species richness, no pattern by season or

apparent.

Ap

pe

nd

ixA

,T

ab

le2

.A

bu

nd

an

ce,

bio

ma

ss,

sp

ecie

srich

ne

ss

and

sp

ecie

sd

ive

rsity

of

fish

cau

gh

tb

ybe

amtr

aw

lin

Ellio

ttB

ayb

yse

aso

n.

Sp

eci

es

Sp

eci

es

Lo

ca

tio

nA

bu

nd

an

ceB

iom

ass

(gm

)rich

ne

ss

div

ers

ity

Sum

mer

Aut

umn

Sum

mer

Aut

umn

Sum

mer

Aut

umn

Sum

mer

Aut

umn

PSD

DA

1S

ite

82

2.3

60

2.2

41

6.5

56

0.6

30

.90

PSD

DA

1R

efe

ren

ceS

tatio

n7

315

95.0

68

.65

20

.64

0.2

2PS

DD

A2

Site

3.7

2.7

82

3.7

111

3—

6—

—0

.77

Ap

pe

nd

ixA

,T

ab

le1

.A

bu

nd

an

ce,

bio

ma

ss,

sp

ecie

srich

ne

ss

and

sp

ecie

sd

ive

rsity

of

fish

cau

gh

tb

ybe

amtr

aw

lin

Com

men

cem

ent

Bay

du

rin

gsu

mm

er,

19

86

.

Sp

eci

es

Sp

eci

es

Lo

ca

tio

nA

bu

nd

an

ceB

iom

ass

(gm

)rich

ne

ss

div

ers

ity

PSD

DA

1S

ite

121

21

0.0

PSD

DA

2S

ite

145

30

.47

20

mS

tatio

n43

4,1

91

80

.71

155

Saratoga Passage

Abundance and biomass. Saratoga Passage was not sampled during the

autumn, therefore only the summer data were analyzed. Total abundance values

ranged from 3 to 12 fish per location; total biomass values ranged from 51 to

1 ,004 grams per location (Appendix A, Table 3). The PSDDA site was the

deepest and had the highest values of both abundance and biomass. The

abundance and biomass values diminished as station depth decreased.

Species richness. Species richness ranged from 1 to 7 (Appendix A, Table

3). The pattern was the same as that of abundance and biomass: the deeper

PSDDA site had the highest values then values diminished as station depth

decreased.

Species diversity. Species diversity values ranged from 0.0 to 0.5

(Appendix A, Table 3). The highest values were at the deepest PSDDA location,

then values decreased with decreasing depth at all other locations.

Appendix A, Table 3. Abundance, biomass, species richness and speciesdiversity of fish caught by beam trawl in Saratoga Passageduring summer.

Species SpeciesLocation Abundance J3iomass (gm) richness diversity

PSDDA Site 11.7 1,004.2 7 0.4680 m E Station 7.0 109.5 3 0.4140 m S Station 6.0 51 .0 3 0.3820 m S Station 3.0 52.0 1 0.00

156

APPENDIX B

Length frequency histograms of abundant,

non-commercially or recreationally important,

fish caught in Commencement Bay and Elliott Bay.

157

5~

~ Patfish — male

4 ~ Ratfish femaleD Ratfish — juvenile

F _____________________

RE 3QUE~ 2C

1~~1Il I I —r— JI~I II I~II 1111111 I liii

75 95 115 135 155 175 195 215 235 255 275 295 315 335 355 375 395 415 435LENGTH mm

Appendix B, Figure 1, Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during summer at156 m in Commencement Bay.

2 ____________________

~ Ratfish — female

~ Patfish — male

FREQUIEP1CV

0 III e ~—~—i——~-: it :i i I :+—~

75 95 115 135 155 175 195 215 235 255 275 295 315 335 355 375 395 415LENGTH

Appendix B, Figure 2. Length frequency of otter trawl caught ratfish,shown by sex, during summer at PSDDA 1 in Commencement Bay.

158

3

~ Ratfish — male

F ~ P.atfish — femaleR 2 ~ Ratfish — juvenile£QUE11CiV

0 1 1 I i i i I — +— — — — —

75 95 115 135 155 175 195 215 235 255 275 295 315 335 355 375 395LENGTH mm

Appendix B, Figure 3, Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during autumn atPSODA 1 in Commencement Bay,

4~

~ Ratfish — male

~ Ratfish — female3~

F Ratfish — juvenileREQU2ENCV

0 .1 i I I—~ —i—il I I Iii ii I I IllIllIlIf I

75 95 115 135 155 175 195 215 235 255 275 295 315 335 355 375 395LENGTh mm

Appendix B, Figure 4. Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during summer atPSDDA 1 reference site in Elliott Bay.

159

FREQUENCY

315 335 355 375 395 415 435 455

Appendix B, Figure 5. Length frequency of ottershown by sex, during autumn at PSDDAin Elliott Bay.

trawl caught ratfish,2, reference site 1

Figure 6, Length frequency of otter trawl caught ratfish,shown by sex and life history stage, during autumn atPSDDA 2, reference site 2 in Elliott Bay.

Ratfish — male

Patfish — female

4.0

3.5

3.0

2.5

2.0

1.5

1.0

0.5

0.0 I.Ih 11.75 95 115 135 155 175 195 215 235 255 275 295

E TN

3

Ratlish — male

~ Ratfish — female

— juvenileFREaUENCV

2

0

Appendix B,

75 95 115 135 155 175 195 215 235 255 275 295 315 335 355 375 395

LEN6TH mm

160

FREaUENCV

FREaUENCY

Appendix B, Figure 8, Length frequency of otter trawl caught slendersole males during summer at PSDDA 2 in Elliott Bays

Slender sole — male

~ Slender sole — female

D Slender sole — juvenile

16

14

12

10

8

6

4

2

0

Appendix B,

7

5

105 125 145 165 185 205 225 245 265 285 305LEK6TH

Figure 7. Length frequencysole, shown by sex and lifeat PSDDA 1 in Elliott Bay.

of otter trawlhistory stage,

caught slenderduring autumn

Slender sole — male

D Slander sole — female

5

4

3

2

1

0105 125 145 165 185 205 225 245 265 285 305

LENGTh mm

161

2

Appendix B, Figure 9. Length frequency of otter trawl caught slendersole, shown by sex, during autumn at PSDDA 2 in Elliott Bay.

.7

8

F5RE1k4UE3

CV

0

105 125 145 165 185 205 225 245 265 285 305LEN6TH mm

162

APPENDIX C

Abundance and biomass (and range at multiple

sample stations) of otter trawl caught fish

by station and species in Commencement Bay,

Elliott Bay, Saratoga Passage, and

Port Gardner

163

Appendix C, Table 1. Abundance and range at multiple sample stations of ottertrawl-caught fish by station and species in CommencementBay on June 13, 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 ci 40 m 156 m(range) (range)

Spiny dogfish- adult 0.3 0-1 0.3 0-1juvenile

Longnose skateRatfish- adult 4.7 1—9 1.3 0-3 6.0

juvenile 1 .0Pacific herring 5.0Longfin smeltPlainfin midshipman- adult 1 .0

— juvenilePacific cod- adult 24.0

— juvenile 4.0Pacific hake— adult 0.3 0—1

— juvenilePacific tomcod— adult

— juvenileWalleye pollock— adult

— juvenileRed brotulaPallid eelpout 0.3 0—1Shortfin eelpoutBlack eelpout- adult

— juvenileBlackbelly eelpout— adult 1 .0

— juvenileShiner perch- adult

— juvenilePile perch- adult

— juvenileBluebarred pricklebackCopper rockfish- adult

— juvenileSplitnose rockfishQuillback rockfish- adult 0.3 0-1 3.0

‘I U — juvenile

Canary rockfishRockfish UID- juvenileSablefish— adult

— juvenileLingcodRoughback sculpinSpinyhead sculpinSoft sculpinTadpole sculpinNorthern spearnose poacher

164

Appendix C, Table 1. (Continued)

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m 40 in 156 in(range) (range)

Sturgeon poacherBluespotted poacherSnailfish UIDSpeckled sanddab— adult

— juvenileArrowtooth flounderRex sole— adult 6.0

— juvenileFlathead sole— adult

— juvenileRock sole— adult 2.0

— juvenileSlender sole— adult 1 .0 0—2 0.3 0—1

— juvenileDover sole— adult 1.3 1—2 1.0 0—3 5.0 6.0

— juvenileEnglish sole- adult 0.3 0-1 40.0

— juvenile0—0 sole— adult

H H — juvenile

Totals 6.9 5.5 91 13

165

Appendix C, Table 2. Biomass (in grams) and range at multiple sample stationsof otter trawl—caught fish by station and species inCommencement Bay on June 13, 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m 40 m 156 m(range) (range)

Spiny dogfish- adult 16.7 0—50 456.7 0-1,370j uv.

Longnose skateRatfish— adult 1 ,357.7 940—2,923 444.0 0—1 ,012 2,085.0

juvenile 6.0Pacific herring 45.0Longfin smeltPlainfin midshipman- adult 108.0

- juv.Pacific cod- adult 549.0

— juvenile 52.0Pacific hake— adult 13.5 0—40.5

— juvenilePacific tomcod— adult

H — juvenile

Walieye pollock— adult— juvenile

Red brotulaPallid eelpout 1.7 0-5.0Shortfin eelpoutBlack eelpout— adult

— juvenileBlackbelly eelpout— adult 9.0

II — juv.

Shiner perch— adult— juvenile

Pile perch- adult— juvenile

Bluebarred pricklebackCopper rockfish— adult

— juvenileSplitnose rockfishQuillback rockfish— adult 643.3 0—970 501 .0

- juv.Canary rockfishRockfish UID- juvenileSablefish— adult

‘I — juvenile

LingcodRoughback sculpinSpinyhead sculpinSoft sculpinTadpole sculpinNorthern spearnose poacher

166

Appendix C, Table 2. (Continued)

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 in 40 in 156 m(range) (range)

Sturgeon poacherBluespotted poacherSnailfish UIDSpeckled sanddab— adult

I, —

Arrowtooth flounderRex sole— adult 161 .0

— juvenileFlathead sole— adult

I’ — juvenile

Rock sole— adult 114.0— juvenile

Slender sole- adult 74.7 0-174 40.2 0-92.5— juvenile

Dover sole— adult 587.7 360—1,013 1,203.0 0—2,700 399.0 2,718.0H — juvenile

English sole— adult 122.7 0—368 5,824.0— juvenile

C—0 sole- adult— juvenile

Totals 2052 2909.9 7762 4809

167

Appendix C, Table 3. Abundance and range at multiple sample stations ofotter trawl—caught fish by station and species inCommencement Bay on September 8, 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m 40 m 156 m(range) (range)

Spiny dogfish- adultjuvenile

Longnose skateRatfish— adult 8.0 5—15 5.0 5—8 1 2 26

juvenile 0.5 0—1 7Pacific herringLongfin smelt 0.5 0-1Plainfin midshipman- adult 6

II — juvenile

Pacific cod— adult 0.7 0—1 0.3 0—1— juvenile

Pacific hake— adult 0.3 0—1— juvenile 0.5 0—1

Pacific tomcod— adult 7— juvenile 54

Walleye pollock— adult 0.7 0—2 0.7 0-1— juvenile

Red brotulaPallid eelpoutShortfin eelpoutBlack eelpout— adult 0.3 0—1

— juvenileBlackbelly eelpout— adult 7

B B — juvenile

Shiner perch— adult 2— juvenile

Pile perch- adultI’ — juvenile

Bluebarred pricklebackCopper rockfish— adult 2

I’ — juvenile

Splitnose rockfishQuiliback rockfish— adult 0.7 0—1 8 2

— juvenileCanary rockfishRockfish UID— juvenileSablefish- adult

— juvenileLingcodRoughback sculpin 16Spinyhead sculpin 0.7 0-1Soft sculpin 0.5 0—1Tadpole sculpinNorthern spearnose poacher

168

Appendix C, Table 3. (Continued)

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 in 40 in 156 in

(range) (range)

Sturgeon poacherBluespotted poacherSnailfish UIDSpeckled sanddab— adult 2

— juvenileArrowtooth flounder 2Rex sole- adult 0.3 0-1 8 3

— juvenileFlathead sole— adult

— juvenileRock sole— adult 16 10

— juvenileSlender sole— adult 3.7 0—6 1.3 1—2 12 3

— juvenileDover sole— adult 8.3 5-10 3.3 1—8 15 12

— juvenile 3English sole— adult 0.7 0—1 19 182 3

— juvenileC—0 sole— adult 1

— juvenile

Totals 23.6 12.6 51 307 59

169

Appendix C, Table 4. Biomass (in grams) and range at multiple samplestations of otter trawl—caught fish by station andspecies in Commencement Bay on September 8, 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m 40 m 156 m(range) (range)

Spiny dogfish- adult‘I

Longnose skateRatfish- adult 1 ,830.0 720—3,610 1,533.3 850-2,550 635 290 3140.0

It juvenile 7.7 0—23 550.0

Pacific herringLongfin smelt 0.7 0-2Plainfin midshipman- adult 219.2

- juv.Pacific cod- adult 1 ,573.3 0—4,200 180 0-540

— juvenilePacific hake- adult 140.0 0-420

II — juvenile 0.5 0—1.5

Pacific tomcod- adult 355— juvenile 210

Walleye pollock— adult 53.3 0—160 134 0—212— juvenile

Red brotulaPallid eelpoutShortfin eelpoutBlack eelpout— adult 1 .2 0—3.5 4.5

— juvenileBlackbelly eelpout— adult 116.5

- juv.Shiner perch— adult 57 22

— juvenilePile perch- adult 300

— juvenileBluebarred pricklebackCopper rockfish— adult 820

— juvenileSplitnose rockfishQuillbackrockfish— adult 301.7 0—565 1,190 2,480 1,180

- juv.Canary rockfishRockfish UID- juvenile 12.5Sablefish— adult

I’ — juvenile

LingcodRoughback sculpin 177.5Spinyhead sculpin 5.0 0-8.0Soft sculpin 1.8 0—5.5Tadpole sculpinNorthern spearnose poacher 33.4

170

Appendix C, Table 4. (Continued)

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 in 40 in 156 in(range) (range)

Sturgeon poacherBluespotted poacherSnailfish lID 5Speckled sanddab— adult 21

— juv.Arrowtooth flounder 330Rex sole— adult 36.3 0—109 308 285

— juvenileFlathead sole- adult

— juvenileRock sole- adult 490 519

— juvenileSlender sole— adult 170 65—360 129.3 98—190 443.7 110

— juvenileDover sole— adult 28,533 1,970—4,460 1,238.3 390—2,810 1,210 3975

— juvenile 50English sole— adult 190 0—290 4,890 29,865 855

— juvenileC—C sole— adult 240

— juvenile

Totals 6,821.3 3,558.4 8,343 36,929.3 10,117

171

Appendix C, Table 5. Abundance and range of multiple sample stations forotter trawl—caught fish by station and species inElliott Bay on July 3, 1986.

Spiny dogfish— adultjuvenile

Longnoae skateRatfish— adult

juvenilePacific herringLongfin smeltPlainfin midshipman- adult

— .Juv.Pacific cod- adult

— juvenilePacific hake— adult

— juvenilePacific toscod— adult

— juvenileWalleye pollock— adult

— juvenileRed brotulaPallid eelpoutShortf in eelpoutBlack eelpout— adult

— juvenileBlackbelly aelpout— adult

- .Juv.Shiner perch— adult

— juvenilePile perch— adult

— juvenileBluebarred pricklsbackCopper rockfiah— adult

— juvenileSplitnoae rockfiahQuillhack rockfiah— adult

— Juv.Canary rockfiahRockfiah RID— juvenileSablefish— adult

— juvenileLingcodRoughback aculpinSpinyhead aculpinSoft aculpinTadpole aculpinNorthern spearnoae poacherSturgeon poacherBlueapotted poacherSnailfiah RIDSpeckled sanddab— adult

- Juv.Arrowtooth flounderRex mole— adult

— juvenileFlathead aole— adult

— juvenileRock mole— adult

— juvenileSlender sole— adult

— juvenileDover sole— adult

— juvenileEnglish sole— adult

— juvenile0—0 sole- adult

— juvenile

2.3 4.7 0—5 0—11 16.0 2.00.3 0.7 0—1 0-2

0.3 0-10.3 0—1 1.0

1.0 4.01.0

2.0

PSDDA 2Reference site II

ART

36.04.0

Species PSDDA I

RangeSR AUT SR ART

1.0 15.3 0—3 2—42

PSDDA IReference mite

SR ART

1.0 10.0

6.0

0.3 0.7

2.0 17.01.0

0.3 5.3

5-8

0-1 0—1

0—5 2—440—3

0-1 0-12

2.0 2.0

1.0

2.013.7 93.3 3—22 67—137 13.0

1.3 0—4

2.7 9.7 0—5 8—13 3.0 1.0

6.0 2-12 1.0 3.0

0.3 1.3 0-1 0-4

PSDDA 2 PSDDA 2Reference site I

RangeSR ART SR ART SR ART

0.3 0-15.0 23.7 2-7 9-34 8.0 20.00.3 3.3 0-1 0—7 1.0

1.7 1—2

0.3 0.3 0—1 0-I2.0 0-6

1.3 0—2 3.0

0.7 1.3 0—2 0—4 7.0

2.0 0-5 1.0

1.00.7 0-2

0.7 0-1

1.0

7.0

0.3 0-1

6.0 9.3 4—9 2—15 3.0 8.0 9.0

4.3 6.7 3—6 4—11 9.0 2.0 3.0

0.3 16.3 0—I 0-29 1.0 28.0 33.0

0.3 0-10.3 0-I

1.3 2.31 .3

5.7 24.7

0.3

15.7 46.72.3 7.32.3 2.31.00.3

0-4 0-70—3

0—12 15—42

0—I

14—18 33—540—5 5—90-3 0-70-20-1

1.0

15.0

12.01.01.0

1.0

8.0

Totals 51.8 248.1 66.0 30.0 16.9 69.9 30.0 67.0 95.0

Biomass (in grams) and range at multiple sample stationsfor otter trawl—caught fish by station and species inElliott Bay on July 3~ 1986.

Species PSDDA I

Range311 SOT SO

P5BPS IReference Site

AUT SU AUT SR AUT

PSDDA 2 PSDDA 2Reference site I

RangeSO SOT SD API

Spiny dogfish— adultjuveni le

Longnoae skateRatfish— adult

juvenilePacific herringLongfin smeltPlainfin midshipman— adult

—joy-Pacific cod— adult

— juvenilePacific hake— adult

— juvenilePacific tomcod— adult

— juvenileWalleye pollock— adult

- juvenileRed brotulaPallid eelpoutShortfin eelpoutBlack eelpout— adult

— juvenileBlackbelly eelpout— adult

— Joy.Shiner perch— adult

— juvenilePile perch— adult

— juvenileBluebarred pricklebackCopper rockf jab— adult

— juvenileSplitnose rockfish

Quillback roetfish— adult- Joy.

Canary rockfishRockfiah DID— juvenileSablefish— adult

— juvenileLingcodRoughback sculpinSpinyhead aculpinSoft aculpinTadpole sculpinNorthern spearnose poacherSturgeon poacherBluespotted poacherSnailfish DIDSpeckled sanddab— adult

— Juv.Arrowtooth flounderRex sole- adult

— juvenileFlathead sole— adult

— juvenileRock sole— adult

— juvenileSlender sole— adult

— juvenileDover sole— adult

— juvenileRriglish sole— adult

— juvenileC—0 sole— adult

— juvenile

615.0 1,675.0 0-1,845.0

318.2 423.3 0—822.012.2 16.0 0—36.5

510.0—715 .0

0-1,090.0

490.0-10,210.0 443.0 510.0 108.50—37.5

3.00-45.0

0-2.50-4.0 2.5

292.0 550.01.5

110.0

172

Appendix C, Table 6.

270-4,230.0 448.5 1000.0

25.30—835.0 1195.5 550.0 1684.2 7066.7 1682.0—1707.50-48.0 15.8 100.0 0-47.5

137.7

130.412.522.7

621 .7

504.0

3,788.3

28.3

0-413.0

0-204.5

0-68.0

0-76.0590.0-11070.0 3704.0 4440.0

0—200.0 20.0

I 9583

126.74.0

3.6

410.0-4005.0

0-325.5 0-380.00—11.9

0-5.5

3.5 6.1 0—10.5 0—18.2

19.1 0—53.4

224.8 1,641.0 100.3—462.0 690.0-2,733.0 407.0 67.0

29.8 0-89.5

91.8 52.3 0-260.0 45.0—60.0 13.5 3.5

1,625.0 840.0-2,045.0 330.0 110.0 360,0 0-1080.0

666.7 4,916.7 0—2,000.0 0—14,750.0

866.7 0-1750.0

0.8I .3

9.0 0—276.7 0-20.0

103.0 205.0 0—309 0-615.0 88.5 3.7 0-11.08.0 0-15.0 110.0

410.8 2,850.3 0—774.5 1,860.0-4,315.0 1,481.0

48.3 0—145.0

854.2 1,770.0 461.5—1292.0 870.0—2320.0 441.0 445.0 893.9 613.3 200.0—443.0 170.0-1,020.017.2 56.5 0—39.5 32.0—72.0 6.5

280.7 361.7 143.0—435.0 0-1,085.0 138.5 4,277.0 2,533.3 491.0—2,132.0 980.0-5,300.05.3 0-9.0

57-7 0-173.0 100.3 3,810.0 0-301.0 0-6,280.0

18.5

25.0

206.0 745.0

4,291.0 620.05,280.0

252.0

Totals 3,969.9 20,630 4,993 2,801 7,003.2 17,496.8 8,790 12,255

173

Appendix C, Table 7. Abundance and range at multiple sample stations of ottertrawl-caught fish by station and species in SaratogaPassage on July 1, 1986.

Location

Species PSDDA PSDDA PSDDA 20m E 40m B 40m W 80m B(range) reference

Spiny dogfish— adult 0.5 0—1 5juvenile

Longnose skateRatfish— adult 1 1—1

juvenilePacific herringLongfin smeltPlainfin midshipman- adult

— juvenilePacific cod— adult

— juvenilePacific hake- adult 2.5 0-4 2

— juvenilePacific tomcod— adult

— juvenileWalleye pollock— adult

— juvenileRed brotulaPallid eelpoutShortfin eelpout 0.7 0-2Black eelpout- adult

— juvenileBlackbelly eelpout— adult 2

— juvenileShiner perch- adult

— juvenilePile perch— adult

— juvenileBluebarred pricklebackCopper rockfish- adult

— juvenileSplitnose rockfishQuillback rockfish- adult 0.5 0-1

— juvenileCanary rockfishRockfish UID- juvenileSablefish- adult

— juvenileLingcod 0,3 0—1Roughback sculpinSpinyhead sculpinSoft sculpinTadpole sculpin 5 0-9Northern spearnose poacher

174

Appendix C, Table 7. (Continued)

Location

Species PSDDA PSDDA PSDDA 2Cm E 4Cm E 4Cm W 80 in E(range) reference

Sturgeon poacherBluespotted poacherSnailfish UIDSpeckled sanddab— adult

— juvenileArrowtooth flounderRex sole— adult

— juvenileFlathead sole— adult

— juvenileRock sole— adult 2 5

II — juvenile

Slender sole- adult 2.7 0-7 3 4— juvenile

Dover sole- adult— juvenile

English sole- adult 4 7 2 7— juvenile

C—C sole— adultII It — juvenile

Totals 10.60 5 6 15 2 19

175

Appendix C, Table 8. Biomass (in grams) and range at multiple sample stationsof otter trawl caught fish by station and species inSaratoga Passage on July 1 , 1986.

Location

PSDDASpecies PSDDA PSDDA Reference 20m E 40m B 40m W 80m B

(range)

Spiny dogfish- adult 351.7 0-1055 978juvenile

Longnose skateRatfish— adult 441.7 255—695 142

juvenilePacific herringLongfin smeltPlainfin midshipman- adult

— juvenilePacific cod— adult

— juvenilePacific hake- adult 741 .5 0—1 ,381 .5 678

— juvenilePacific tomcod— adult 1 .5

— juvenileWalleye pollock— adult

— juvenileRed brotulaPallid eelpoutShortfin eelpout 15.2 0—45.5Black eelpout— adult 56

— juvenileBlackbelly eelpout— adult

— juvenileShiner perch- adult

— juvenilePile perch— adult

— juvenileBluebarred pricklebackCopper rockfish- adult

— juvenileSplitnose rockfishQuillback rockfish- adult 34.7 0-104

— juvenileCanary rockfishRockfish UID- juvenileSablefish— adult

— juvenileLingcod 1,666.7 0-5,000Roughback sculpin 9.5Spinyhead sculpinSoft sculpinTadpole sculpin 119.0 0-357

176

Appendix C, Table 8. (Continued)

Location

PSDDASpecies PSDDA PSDDA Reference 2Orn E 40m E 40m W 8Oni E

(range)

Northern spearnose poacherSturgeon poacherBluespotted poacherSnailfish lIDSpeckled sanddab- ad~ilt

— juvenileArrowtooth flounderRex sole— adult loT

— juvenileFlathead sole— adult

I’ — juvenile

Rock sole- adult 65 448— juvenile 19

Slender sole— adult 97 0—245 120 142— juvenile

Dover sole— adult 142— juvenile

English sole— adult 424 863 137 417— juvenile

C—0 sole— adult— juvenile

Totals 3,467.5 798 498.5 1,3315 137 1,984

Ap

pe

nd

ixC

,T

ab

le9

.N

umbe

r(a

bu

nd

an

ce)

of

fish

,b

iom

ass

(in

gra

ms)

and

ran

ge

at

Nav

yan

dPS

DD

Asite

so

fo

tte

rtr

aw

lca

ug

ht

fish

by

sta

tio

nan

dsp

ecie

sin

Po

rtG

ard

ne

ron

Fe

bru

ary

12an

d1

3,

1986

(W86

)~

7.4

1690

440

312

01

30

219

8.1

134

.5

sort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erW

86W

86W

86W

OO

W8

6W

OO

Fidn

spec

ies

NAV

YO

TPS

OD

A2

PSO

DA

ITr

anI

205

Tran

14O

STr

anI

1005

Sci

en

tific

Nam

eC

omm

onN

ame

Abu

rul

Ran

geS

iom

ase

Ran

geA

bund

Ran

geB

iom

ass

Ran

geA

bund

Ran

geB

iom

ase

Ran

geA

bund

Bio

maa

aA

bund

Bio

maa

aA

bund

Bio

mas

s

Lam

petra

tdde

nlal

aP

acifi

cla

mpr

ey0.

30-

13

0-9

Squ

aius

acan

o,ia

ssp

iny

dogf

ish

0.5

0-1

115

0-2

30

0.3

0-1

498.

30

-14

95

Raj

arh

ine

Iong

noce

skat

ei4

ydro

lagu

sco

lliel

ratfi

sh-

adul

t9.

72

-22

1348

.33

80

-24

20

332

2-4

469

67.5

59

20

-80

15

25.3

12-3

651

62.8

28

93

-87

40

3769

60-ji

Nef

lhle

0.7

0-2

11.2

0-3

3.6

95

-13

197.

514

5-25

09

0-2

796

.70

-29

0A

iosv

sapk

iissi

ma

Am

eiic

ansh

ad0.

30-

141

.50

-12

4.6

Clu

pea

palla

sii

Pac

ific

herr

ing

Por

ichi

hys

nola

lus

plai

ntin

mId

ship

man

0.3

0-1

350-

105.

10.

50

110

00

20

00.

30.

110

03

0G

aotis

mac

roce

phai

usPa

cific

cod

0.3

01

1333

.50

-40

00

.5M

icro

gadu

spro

xim

t,sPa

cific

iom

cod

-ad

ult

5.7

0-1

221

5.3

0-1

66

0.3

0-1

33.3

0-1

00

-juve

nile

3.7

2-5

48.1

20

.4-8

2.9

420

Mer

iucc

ius

pror

iucl

usPa

cific

hake

-ad

ult

482

4-5

830

48.3

14

25

-59

60

199

-29

6105

47

20

-74

90

18.3

5-3

965

91.7

11

50

-99

35

1016

00-J

uven

ile9.

70

-24

2006

.70

-28

5B

rosm

ophy

cis

mar

gina

tare

dbr

otul

aLy

coda

pus

man

dibu

laris

palli

dee

lpou

tLy

code

sdi

aple

nis

blac

kee

ipou

tLy

code

spa

cifie

sbi

ackb

eliy

eelp

out

8.7

1-16

96.8

16.9

-234

11

12.4

Cym

alog

asle

ragg

tega

tash

iner

perc

h-

adul

t67

.75

-18

716

93.5

12

0.5

-48

60

-juve

nile

10

-315

0-4

5D

amal

ichl

hys

Caer

epi

lepe

rch

-ad

ult

-ju

veni

leR

onqv

ilus

jord

ani

north

ern

ronq

ull

Lurn

penu

ssag

ilia

snak

epr

lckl

ebac

k0.

30-

15

0-1

5S

ebas

tesm

alig

erqu

lilba

ckro

ckfis

h2.

71-

449

1.7

150-

1035

21-

311

105

20

-17

00

10-

246

1.7

0-9

00

1228

02

860

Ano

plop

oma

timbr

iasa

hief

ish

Arte

dius

op.

scui

pin

Dao

ycol

luss

etlg

ersp

inyh

eads

culp

in0.

70

-235

0-1

05

0.3

01

8.3

02

5C

tdto

notu

opug

elen

sis

roug

hbac

ksc

tiple

Giib

erlid

iasi

galu

tes

soft

scul

pin

Icei

lnus

bore

alis

north

ern

scul

pin

Lept

ocof

fiis

arm

aluu

Paci

ficst

agho

rnsc

ulpi

nN

autic

hlhy

soc

uio(

asci

aius

sailt

insc

ulpi

nR

adul

inus

aspr

ellu

ssl

imsc

ulpi

nB

aihy

agon

usni

grip

inni

sbl

ackt

nst

arsn

out

poac

her

Xen

erel

mvs

lalif

rons

blac

ktp

poac

her

3.3

3-4

30.5

27

.9-3

51

0-2

160

-32

1.3

0-3

11.3

0-2

01

11.8

Xen

erel

mus

Ola

can(

hus

biue

spot

ted

poac

her

Cith

aric

hlhy

sup

.sa

ndda

b-

adul

t2

709

295

-juve

nile

i13

.26

95C

ilhar

icht

hys

sord

idus

Paci

ficsa

ndda

bC

ilhar

ichl

hys

stig

mae

ussp

eckl

edsa

ndda

b-

adul

t-—

nile

Ath

eres

thes

slom

jas

arro

wto

oth

floun

der

0.3

01

100

03

00

119

.5G

lypt

ocep

halu

szac

hiru

sre

oso

le-

adul

t2

0-3

240

0-3

75

0.5

0-1

120

24

0.3

0.1

8.3

02

5-

Juve

nile

Hip

pogl

osso

ides

eias

sodo

nfia

thea

dsol

e25

.717

-42

3328

.32

13

5-5

61

00.

50

-I80

0-1

60

120

05

500

Lepl

dops

ella

bThn

eata

rock

sole

-ad

ult

0.3

0-1

163.

40

-49

00.

30.

150

0-1

50

266

013

2080

-juve

nie

27.

85

23.1

Lyop

selta

exiis

slen

der

sole

-ad

ult

30.3

21

-45

2085

12

50

-30

40

14.5

8-21

692.

54

10

-97

526

24

-28

1301

.71

25

5-1

39

0-ju

veni

le9

0-3

17.7

0-41

Mic

rost

omus

paci

ficus

Dov

erso

le7.

33

-12

1686

91

0-2

57

55

3-7

1215

11

70

-12

60

3.7

0-11

818.

30

-24

55

Par

ophy

rsve

llus

Engl

ioh

sole

-ad

ult

111

58

-17

420

325

81

70

-34

75

53.

53

475

04

90

10

10

73

14

1860

79

5.3

80

51

320

1797

0-ju

veni

le0.

70-

18.

30

-13

.51

715

715

00P

ialic

hthy

sst

eila

lus

stor

ryflo

unde

rP

leur

snic

hihy

sco

enos

usC

Osd

eI

60P

selil

chth

ysm

eian

oslic

lus

sand

sole

118

0.1

118

.42

564

26NA

TOTA

L3

44

.83

69

76

891

73

61

94

.31

83

49

101

27

925

56

13

094

10

43

9

Ap

pe

nd

ixC

,T

ab

le9

(co

ntin

ue

d).

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

WaS

W8

6W

86

Wa

sW

86

W86

Flab

Spe

cies

Tran

220

STr

an2

40$

Tran

211

05Tr

an4

20$

Tran

440

$Tr

an4

145$

Sci

en

tific

Nam

eC

omm

onN

ame

Abu

ndS

iom

asa

Abu

ndB

iom

ass

Abu

ndB

iom

ass

Abu

ndB

iom

asa

Abu

ndB

iom

ass

Abu

ndB

iom

ass

Lam

petra

tride

ntal

aP

acifi

cla

mpr

eySq

ualu

sac

antri

assp

iny

dogf

ish

Flaj

arh

ina

Iong

nouo

skat

eH

ydro

lagu

sco

lliel

ratta

h-

adul

t2

580

838

00-ju

veni

le‘

Alos

asa

pidi

ssim

aAm

eric

ansh

adC

lupe

apa

llasi

iP

acifi

che

rrin

gP

oric

hthy

sno

tatu

spl

ainf

inm

idsh

ipm

anG

adt.o

mac

roce

phal

usP

acifi

cco

dM

icro

gadu

spr

oxim

usP

acifi

cto

mco

d-

adul

t13

105

-juve

nile

1410

0M

erlu

cciu

spr

oduc

tus

Pac

ific

hake

-ad

ult

578

0I

410

-ju

veni

leB

rosm

ophy

cis

mar

ylna

tare

dbr

otul

aLy

coda

pus

man

dibu

laris

palli

dee

lpou

tLy

code

sdi

apte

nis

blac

kee

lpou

tLy

code

spac

ilica

blac

kbel

lyae

lpou

t1

102

50C

ymat

ogas

tera

ggre

gata

slen

orpe

rch

-ad

ult

230

120

-juve

nile

4125

534

200

Dam

atch

thys

vacc

api

lepe

rch

-ad

ult

710

512

922

10-ju

venI

le1

1026

310

Ron

qullu

sJo

rdan

!no

rther

nro

nqui

fLu

irpen

ussa

gitta

snak

epr

lckl

ebac

k36

1065

Seb

aste

amal

iger

quill

back

rock

fish

850

213

203

400

Ano

plop

oma

limbr

iasa

blef

ish

Arte

dius

op.

scul

pin

230

17

565

Das

ycot

tus

setig

ersp

lnyh

ead

scul

pin

Chi

tono

tus

puge

tens

isro

ughb

ack

scul

pin

Gilb

ertid

iaoi

galu

tes

soft

scul

pin

Icei

lnus

bore

alis

north

ern

scul

pin

Lept

ocot

tus

arm

atus

Pac

ific

stag

horn

scul

pln

722

52

380

Nau

ticht

hys

ocul

ofaa

ciat

ussa

ilfin

scul

pin

Rad

ulin

usas

prel

lus

slim

scul

pin

Bat

hyag

onus

nlgr

ipln

nis

blac

kfin

star

anou

tpo

ache

rX

ener

etm

usla

titio

nsbl

ackl

ippo

ache

r2

30X

ener

etm

uatri

acan

thus

blue

spot

ted

poac

her

Cith

atic

hthy

aop

.sa

ndda

b-

adul

t1

2712

300

418

0-

juve

nile

580

Cith

aric

hthy

sso

rdid

usPa

cific

sand

dab

115

Cith

aric

hthy

sst

igm

aeus

spec

kled

sand

dab

-ad

ult

2681

0-ju

veni

le1

25A

ther

esth

esst

omia

sar

row

loot

hflo

unde

r1

145

Gly

ptoc

epha

lus

zach

lrus

rea

sole

-ad

ult

155

113

0-

juve

nile

Hog

loss

rAde

sela

ssoc

ion

fath

ead

sole

431

01

110

Lepi

dops

etta

bilin

eata

rock

sole

-ad

ult

123

017

1480

2216

50-ju

veni

le2

81

51

59

145

Lyop

setta

exits

slen

der

sole

-ad

ult

640

190

361

014

830

-ju

veni

leM

icro

stom

uspa

cific

usD

over

sole

180

II31

9028

4900

Par

ophy

rsve

tlus

Eng

lish

sole

-ad

ult

6719

208

1710

140

1490

-juve

nile

5459

01

7.2

Pla

ticht

hys

stel

latu

sst

arry

floun

der

Pleu

ronl

chth

yoco

enos

usC

Ood

e1

50P

setti

chth

ysm

elan

oslic

tus

sand

sole

TOTA

L4

26a

242

54

35

3fi

77

80

201

24

a.2

44

78

59

562

10

45

5

Fish

Spe

cies

Sci

en

tific

Nam

eC

omm

onN

ame

Lam

pelra

tride

ntal

aP

acifi

cla

mpr

eySq

uaki

sac

antrl

assp

iny

dogf

ish

Raj

ath

ina

long

nose

skat

eH

ydro

lagu

sco

lliei

raffi

sh-

adul

tju

veni

leAl

osa

sapi

diss

ima

Amer

ican

shad

ClLr

pea

palla

sil

Pac

ific

herr

ing

Por

icht

hys

nola

tus

plal

nfin

mid

ship

man

Gad

usm

acro

ceph

alus

Pad

ficco

dM

icro

gadu

spr

oxim

usP

acifi

cto

mco

d-

adul

t-ju

veni

leM

erlu

cciu

spr

oduc

tus

Paci

ficha

ke-

adul

tju

veni

leB

rosm

ophy

cis

mar

gina

tare

dbr

otul

aLy

coda

pus

man

dibr

iaris

palli

dea

lpou

tLy

code

sdl

aple

rus

blac

kee

lpou

tLy

code

spac

illca

blac

kbel

lyee

lpsu

tC

yrrra

toga

ster

aggr

egat

esh

iner

perc

h-

adul

t-ju

veni

leD

amal

ithth

ysva

cca

pile

perc

h-

adul

t-

juve

nile

Ron

qullu

sjo

rdan

ino

rther

nro

nqui

lLw

npen

ussa

gitta

snak

epr

ickl

ebac

kSe

bast

esm

alig

erqu

lilba

ckro

ckfis

hA

nopl

opom

afim

bria

sabl

efis

hA

rtedi

usrp

.sc

ulpi

nD

asyc

otlu

sse

llger

spin

yhea

dsc

ulpi

nC

hlto

notu

spug

eten

sis

roug

hbac

ksc

ulpi

nG

ilber

lidia

slga

lute

sso

ftsc

ulpl

nIc

elin

usbo

real

isno

rther

nsc

ulpi

nLa

pfo

cotlu

sar

mat

usP

acifi

cst

agho

rnsc

ulpi

nN

aulic

hthy

soc

ulof

asci

atus

sailf

insc

ulpi

nR

adul

inus

aspr

ellu

ssl

imsc

ulpi

nB

athy

agon

usni

grip

innl

sbl

ackf

lnst

arsn

out

poac

her

Xen

erel

mus

laifm

nsbl

ackt

ippo

ache

rX

ener

ebnu

sbl

acan

thus

bhie

spot

ted

poac

her

Cilh

aric

hlhy

sup

.sa

ndda

b-

adul

t-J

uven

ileC

ithar

ichl

hys

surd

idus

Paci

ficsa

ndda

bC

ithar

lcht

hys

sbgm

aeus

spec

lcle

dsa

ndda

b-ad

ult

juve

nile

Ath

eres

thes

stsm

las

arro

wto

sth

floun

der

Gly

ptoc

epha

lus

zach

lrus

rex

sole

-ad

ult

-ju

veni

leH

ippo

glos

scrid

esel

asso

don

fath

ead

sole

Lepi

dops

etta

bilin

eala

rock

sole

-ad

ult

juve

nile

Lyop

setta

exilis

slen

der

sole

-ad

ult

-ju

veni

leM

icro

stom

uspa

cific

usD

over

sole

Par

ophy

rsve

llus

Engl

ish

sole

-ad

ult

-ju

veni

leP

latic

hthy

sst

ella

ftrs

star

ryflo

unde

rP

leur

onic

hlhy

sco

enos

usC

Oer

ieP

selu

chth

ysm

elan

oslic

tus

sand

sole

0.3

0-1

30

-90.

30-

117

00

-51

00.

30-

114

3.3

0-4

30

5.7

1-14

411.

71

10

-79

00.

30-

10.

30-

15.

70

-13

781.

70

-16

95

2014

514

220

Ap

pe

nd

ixC

,T

ab

le1

0.

Nu

mb

er

(ab

un

da

nce

)o

ffish

,b

iom

ass

(in

gra

ms)

an

dra

ng

ea

tN

avy

an

dP

SD

DA

site

sfo

ro

tte

rtr

aw

lca

ug

ht

fish

,b

ysta

tio

nan

dsp

ecie

sin

Po

rtG

ard

ne

ron

Ap

ril

18an

d2

1,

1986

(SP

86

).

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

SP88

SF

86SP

86S

F86

SP86

SF

86

NAV

YO

TPS

OD

A2

PSO

DA

1Tr

anI

20S

Tran

I40

ST

ran

IlO

OM

Abu

ndR

ange

Sia

mes

eR

ange

Abu

ndR

ange

Bio

mas

sR

ange

Abu

ndR

ange

Bio

mas

sR

ange

Abu

ndB

iom

ass

Abu

ndB

iom

asa

Abu

ndB

iom

ass

3.7

3-4

300

17

0-4

35

3.7

2-5

806.

73

80

-13

30

10-

243

8.3

0-1

10

10-

212

0-3

5

0.7

0-2

31.7

0-7

0

0.7

0-1

21.7

0-4

01

140

6.7

4-8

390

36

0-4

20

10

-219

50

-40

51

85

146-

2125

8.3

90

-45

50.

30-

12.

70

-81

11

11-

126

015

0-33

51

605

0.3

0-1

166.

70

-50

0

4.3

3-7

392

7-5

00.

30-

15.

70

-17

250

11

0.3

0-1

31.7

0-9

52

210

0.7

0-1

30-

72

450

114

5

175

-30

915

23

5-1

49

04.

73

-624

3.3

150-

295

8.7

7-1

041

1.7

25

5-5

00

1583

01.

70

-36.

30

-10

30

-934

6.7

0-1

04

06

5-1

014

158

85

-19

65

0.7

0-1

156.

70

-25

012

125

265

530

.32

0-3

660

604

47

0-7

09

02.

70

-756

6.7

0-1

37

50.

70

-211

00

-33

02

455

6110

870

112

5

TOTA

L9

4.7

98

78

.71

9.7

34

76

.11

3.4

13

49

.118

90

012

01

42

21

Ap

pe

nd

ixC

,T

ab

le10

(co

ntin

ue

d).

Fish

Spe

cies

Sci

en

tific

Nam

eC

omm

onN

ame

Lam

petra

idde

ntat

aP

acifi

cla

mpr

eySq

uatu

sac

anu’

rias

spin

ydo

gfis

hR

aja

thin

slo

ngno

sesk

ate

Hyd

rola

gus

colli

eira

tfish

-ad

ult

juve

nile

Alos

asa

pidi

ssim

aAm

eric

ansh

adC

iupe

apa

llasi

iP

acifi

che

rrin

gPo

rlctrt

hys

nola

lus

plai

ntin

mid

ship

man

Gad

usm

aas3

ceph

atis

Paci

ficco

dM

icro

godu

spr

oxim

usP

acifi

cto

nico

d-

adul

t-ju

verd

leM

erlu

cciu

spr

oduc

tus

Pac

ific

hake

-ad

ult

-ju

veni

leB

rosm

ophy

cis

mar

gina

tare

dbr

otul

aLy

coda

pus

man

dibu

laris

palli

dee

lpou

tLy

code

sdl

apte

rus

blac

kee

lpou

tLy

code

spac

iilca

blac

kbel

lyee

lpos

tC

ymat

ogas

tara

ggre

gata

shin

erpe

rch

-ad

ult

-ju

veni

leD

an7a

litht

hys

vacc

api

lepe

rch

-ad

ult

juve

nile

Ron

quilu

sjo

rdan

ino

rthem

ronq

uil

Lum

penu

ssa

gitta

snak

epr

ickl

ebac

kS

ebas

tesm

alig

erqu

iliba

ckro

cktis

hA

nopl

opom

aIir

nbria

sabt

efis

hA

rledi

us~

.sc

ulpl

nD

asyc

ottu

sse

tiger

spln

yhea

dsc

ulpi

nC

hito

nolu

spug

eten

sls

roug

hbac

ksc

ulpi

nG

ilber

tidia

siga

lute

sso

ftsc

ulpi

nIc

elir,

usbo

real

isno

rther

nsc

ulpi

nLe

ptsc

ottu

sar

mat

usP

acifi

cst

agho

rnsc

ulpl

nN

autic

hthy

soc

ulot

asci

alus

sailt

insc

ulpi

nR

adul

inus

aspr

ellu

ssl

imsc

slpi

nB

athy

agon

usni

grip

inni

sbl

ackf

lnot

arsn

out

poac

her

Xen

eret

mus

latit

rons

blsc

kbp

poac

her

Xen

eret

mus

biac

anih

usbl

uesp

otte

dpo

ache

rC

ithar

icht

hys

op.

sand

dab

-ad

ult

-ju

veni

leC

ithar

lcht

hys

sord

idus

Paci

ficsa

ndda

bC

ithar

icht

hys

stig

mse

ussp

eckl

edsa

ndda

b-ad

ult

juve

nile

Ath

eres

thes

slom

ias

arro

wto

oth

floun

der

Gly

ptoc

epha

lus

zach

irus

ran

sole

-ad

ult

-juve

nile

Hip

pogl

osso

ides

elas

sodo

nfa

thea

dso

leLe

pido

pset

tabi

linea

taro

ckod

e-

adul

t-ju

venl

eLy

opse

ttaex

ilissl

ende

rso

le-

adul

t-ju

veni

leM

icro

stom

uspa

cjfic

usD

over

sole

Par

ophy

rsca

llus

Eng

lish

sole

-ad

ult

-ju

veni

leP

latic

hthy

sst

ella

tus

star

ryflo

unde

rP

leur

onic

hlhy

sco

enos

usC

Onn

ieP

sets

chlh

ysm

elan

ostic

lus

sand

sole

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

SP86

SP86

SP86

SP86

SP86

SP86

Tran

220

STr

act

24f

lSTr

an2

liES

Trac

t4

205

Trac

t4

40S

Trac

t4

1455

Abu

ndB

iom

ass

Abu

ndB

lom

ass

Abu

ndB

iom

ass

Abu

ndB

iom

ass

Abu

ndB

lom

ass

Abu

ndB

tom

ass

15

127

19

215

300

560

0

12

250

221

0

255

211

203

335

115

0

540

520

0

115

412

02

5

1183

02

52

15

1114

208

80

816

10

417

60

I2

110

l—~ co CO

TOTA

L8

32

712

88

69

15

85

525

482

71

814

33

82

Ap

pe

nd

ixC

,T

ab

le1

1.

Nu

mb

er

(ab

un

da

nce

)o

ffish

,b

iom

ass

(in

gra

ms)

an

dra

ng

ea

tN

avy

an

dP

SD

DA

site

sfo

ro

tte

rtr

aw

lca

ught

fish

bysta

tio

nan

dsp

eci

es

inP

ort

Gar

dner

onJu

ne30

and

Ju

ly2,

1986

(StJ

86).

Fish

Spe

cies

Sci

en

tific

Nam

e

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

SU

BS

SU

86S

U86

SU

BS

NAV

YO

TP

SD

DA

2P

SD

DA

1T

rw1

20$

Com

mon

Nam

eA

bund

Ran

geB

iom

ess

Ran

geA

bund

Ran

geB

iom

ass

Ran

geA

bund

Ran

geB

iom

ass

Ran

geA

bund

Bio

mas

s

Lam

pel

ralri

dent

ata

Pac

ific

lam

prey

Squ

aksa

can#

rias

spin

ydo

gfis

h3.

71-

710

92.5

58

0-1

70

2R

aja

chin

alo

ngno

sesk

ate

Hyd

rola

gus

coilf

elra

ttuh

-ad

ult

53

-923

39.7

37

6-5

33

81.

30-

441

8.3

0-1

25

51

0-2

316.

50

-71

2-ju

veni

le1

0-2

5.3

0-1

410

.31-

2714

8.3

21

-47

7Al

coa

sapi

dlss

ima

Amer

ican

shad

Clu

pea

palla

sil

Pac

ific

herr

ing

Poric

Mth

ysno

lalu

spl

aint

inm

idsh

ipm

anG

adw

maa

vcep

haiu

sPa

cific

cod

Mic

roga

dus

prox

imus

Paci

ficto

mco

d-

adul

t-ju

veni

leM

erlu

cciu

spr

oduc

lus

Pac

ific

hake

-ad

ult

7.3

5-9

890.

54

64

-12

00

0.7

0-1

166.

30

-31

8-

juve

nile

Bro

smop

hyci

sm

argi

nata

red

brot

ula

Lyco

dapu

sm

andi

bula

rispa

llid

eelp

out

Lyco

des

diap

teru

sbl

ack

eelp

out

0.3

0-I

9.3

0-2

8Ly

code

opac

ilica

btac

kbel

lyee

lpou

t5.

73

-10

223

11

9-4

08

C,p

nalo

gast

erag

greg

ata

shin

erpe

rch

-ad

ult

-ju

veni

leD

amal

icht

hys

vacc

api

lepe

rch

Ron

quilu

sJo

rdan

north

ern

ronq

ull

Lum

penu

ssa

gitta

snak

epr

ickl

ebac

k$e

bast

esm

alig

erqu

iltba

ckro

ckfia

h1.

70

-247

9.7

12

0-9

35

0.7

0-1

212.

30

-38

3A

nspi

opom

alir

nb,la

sahi

efis

hA

rtedi

us~

.sc

ulpi

nD

asyc

otlu

sse

liger

spin

yhea

dsc

ulpi

nC

hito

notu

spug

eten

sis

roug

hbac

ksc

tdpi

nG

ilber

tidia

siga

luto

sso

ftsc

tdpi

nIc

olin

usbo

real

isno

rther

nsc

ulpi

nLe

ptoc

otlu

sar

mat

usP

acifi

cst

agho

rnsc

ulpi

nN

autlc

hthy

soc

tiofa

scla

o,s

sailt

insc

iripi

nR

aciu

linus

aspr

eilu

ssli

msc

ulpi

nB

alhy

agon

usni

grip

inni

sbl

ackt

nst

arsn

out

poac

her

10-

220

.70

-34

Xen

eret

mus

lalif

rons

blac

ktp

poac

her

0.3

0-I

2.5

0-7

Xen

ereb

nus

lflac

anth

usbk

resp

ofle

dpo

ache

rC

ithar

lchl

hys

sp.

sand

dab

-ad

ult

-ju

veni

leC

ithar

icht

hys

sord

idus

Paci

ficsa

ndda

bC

ilhar

ichl

hys

slig

mae

ussp

eckl

edsa

ndda

b-a

dult

-juve

nile

Ath

eres

thes

slom

ias

arro

wto

oth

tinun

der

Gly

ploc

eplta

ius

zach

irus

rev

sole

-adu

lt-

juve

nile

Hjo

pogi

sssc

ides

eias

sodo

nfa

thea

dso

le2.

30

-739

80-

119

4Lo

pido

pset

labi

liiie

ata

rock

sole

-ad

ult

-ju

veni

leLy

opse

tlaex

ilissl

ende

rso

le-

adul

t5

3-7

314.

717

5-58

12

1-3

97.3

57

-12

92.

70-

610

80-

212

-juve

nile

0.3

0-1

20-

6M

icro

stsm

uspa

ciltc

usO

over

uole

-adu

lt0.

70-

117

9.5

0-4

02

6.7

3-11

1927

.21

41

5-2

35

54

2-5

1142

,74

72

-17

72

-juve

nile

0.3

0-1

20

-60,

70-

13.

30

-5P

arop

hyrs

vetlu

sE

nglis

hso

le-

adul

t28

.72

3-2

553

78.2

38

37

-70

34

12.3

2-3

224

87.2

30

6-6

72

93

2-4

658.

82

62

-10

88

-ju

veni

leP

latic

hlhy

sst

ella

tus

star

ryflo

unde

rPl

etKo

nich

thys

coen

osus

COru

leP

selti

chlh

ysm

elan

oslic

tus

sand

sole

Por

tG

ardn

erP

ort

Gar

dner

SU

BS

SU

BS

Trai

tI

40$

Tran

IlO

OM

Abu

ndB

iom

ass

Abu

ndB

iom

ass

288

31

707

819

41.5

1428

1

531

9.5

I54

.589

3719

495

1,5

I10

.5

241

61

104.

5

236

228

.5

133

623

9

211

51

51

6.2

158

2

249.

515

1315

.528

6475

.5

GD

I-’

TOTA

L6

1.4

11

30

43

3.9

52

55

.91

3.4

24

64

32

48

5.5

122

66

84

.563

11

20

3

Ap

pe

nd

ixC

,T

ab

le11

(co

ntin

ue

d).

Fish

Spe

cies

Sci

en

tific

Nam

eC

omm

onN

ame

Lam

petra

tride

ntat

aP

acifi

cla

mpr

eySq

ualu

sac

antil

assp

iny

dogf

ish

Raj

arh

ina

long

nose

skat

eH

ydro

lagu

sco

llie!

raffi

sh-

adul

t-

juve

nile

Alos

asa

pidi

ssim

aAm

eric

ansh

adC

lupe

apa

llasi

iP

acifi

che

rrin

gP

oric

hthy

sno

tatu

spl

ainh

inm

idsh

ipm

anG

adus

mac

roce

phal

usPa

ciltc

cod

Mic

mga

dus

prox

imus

Paci

ficto

mco

d-

adul

t-ju

veni

leM

erlu

cciu

spr

oduc

tus

Paci

ficha

ke-

adul

t-

juve

nile

Bro

smop

hyci

sm

argi

nata

red

brol

ula

Lyco

dapu

sm

arrd

ibul

aris

palli

dee

lpou

tLy

code

sdi

apta

ivs

blac

kee

lpou

tLy

code

spac

illca

blac

kbel

lyee

lpou

tC

just

atog

aste

ragg

rega

tash

iner

perc

h-

adul

t-

juve

nile

Dam

alic

hlhy

sva

cca

pile

perc

hR

orrq

uilu

sjo

rdan

!no

rther

nro

nqul

lLu

mpe

nus

sagi

ttasn

ake

prlc

kleb

ack

Seb

aste

smal

iger

qulll

back

rock

tish

Ano

plop

oma

limbr

iasa

blef

ish

Arle

dius

~.

scul

pin

Das

ycot

tus

sellg

ersp

lnyh

ead

scul

pin

Chi

tono

lusp

ugot

ensi

srs

ughb

ack

scul

pin

Gilb

erO

dia

siga

lute

sso

ftsc

ulpi

nIc

elin

usbo

real

isno

rther

nsc

ulpl

nLe

ptoc

otlu

sar

mal

usP

acifi

cst

agho

rnsc

ulpi

nN

autic

hlhy

soc

ulof

asci

atus

sailt

insc

ulpi

nR

adul

inus

aspr

ellu

ssl

imsc

ulpi

nB

athy

agon

usni

grip

inni

sbl

ackf

inst

arsn

out

poac

her

Xen

erel

mus

IaO

7ron

sbl

ackt

ippo

ache

rX

ener

etnr

usld

acan

lhus

blue

spol

ted

poac

her

Cilh

aflc

hthy

sop

.sa

ndda

b-

adul

t-

juve

nile

Cith

aric

hihy

sso

rdid

usPa

cific

sand

dab

Cith

adch

thys

sllg

rnae

ussp

ecki

edsa

ndda

b-a

dult

-ju

veni

leA

ther

esth

esst

omia

sar

row

loot

hflo

unde

rG

lypt

ocep

halu

sza

chiru

sre

xso

le-a

dult

-ju

veni

leH4

xpog

loss

oide

sel

amod

onta

thea

dso

leLe

pido

pset

tabi

linea

taro

ckso

le-

adul

t-

juve

nile

Lyop

setta

exilis

slen

der

sole

-ad

ult

-ju

veni

leM

icro

stom

uspa

cific

usD

over

sole

-ad

ult

-ju

veni

leP

arop

hyrs

vellu

sEn

glis

hso

lead

ult

-ju

veni

leP

latic

hthy

sst

ella

lzJS

star

ryflo

unde

rP

leur

onic

hlhy

sco

enos

usCO

sole

Pse

ttich

lhys

mel

anos

O’c

tus

sand

sole

Por

tG

erdn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

SU

86SO

86SU

86S

o86

5086

SO86

SO86

SO86

Trai

t2

20S

Tra

it2

405

Trai

t2

lbS

Tran

420

STr

ait

440

STr

ait

414

5SPS

DD

AR

ef1

PS

DD

AR

ef2

Abu

ndB

lom

ass

Abu

ndB

iom

ess

Abu

ndS

iam

ese

Abu

ndB

tom

ase

Abu

ndB

iom

ass

Abu

ndB

tom

aee

Abu

ndB

iom

aee

Abu

ndB

iom

ese

476

2518

2115

116

002

522

139

703

1055

142

01

111

13

128

.51

148.

51

1500

1683

3

114

31

181

371

1

366

17

119

761

.5

142

11

270

136

0.5

143

123

1t2

0

I22

231

310

.5

317

4.5

16.

51

23.5

560

6.5

113

224

.5I

186

292

.5

150

316

31

67.5

415

636

1955

449

4.5

14

224

9.5

2218

70.5

447

56

439.

55

1540

510

50

125

9.5

l-~ CO

TOTA

L14

65

7.5

561

13

46

303

85

011

481

16

7.5

125

08

322

56

65

.515

55

88

Fish

Spe

cies

Sci

en

tific

Nam

eC

omm

onN

ame

Pac

ific

lam

prey

spin

ydo

gfis

hlo

ngno

sesk

ate

raffi

sh-

adul

t-

jLPjen

hleAm

eric

ansh

adP

acifi

che

rrin

gpl

ainf

inm

idsh

ipm

anP

aclfc

cod

Paci

ficto

mco

d-

adul

t-ju

veni

leP

acifi

cha

ke-

adul

t-

juve

nIle

red

brot

ula

palli

dee

ipou

tbl

ack

eeip

out

blac

kbei

lyee

ipou

tsh

iner

perc

h-

adul

t-ju

veni

lepi

lepe

rch

-ad

ult

-ju

veni

leR

onqu

ilus

Jord

an!

north

ern

ronq

uil

Lso~

penu

ssa

gitta

snak

epr

ickl

ebac

kSe

bast

osm

aiig

erqu

lilba

ckro

ckfis

hA

nypl

opom

atin

sbrla

sabl

efis

hA

rtodl

usop

.sc

uipi

nD

asyc

oiiu

sse

tiger

spin

yhea

dsc

ulpi

nC

hito

noiu

spug

elen

sis

roug

hbac

ksc

ulpi

nG

ilber

iidia

siga

luie

sso

ftsc

ulpi

nIc

eiln

usbo

real

isno

rther

nsc

uipi

nLo

pioc

otiu

san

natu

sP

acifi

cst

agho

rnsc

ulpi

nN

autlc

hthy

soc

ulof

ascl

atus

sailt

insc

ulpi

nR

adul

inus

aspr

eilu

ssl

imac

ulpl

nB

athy

agon

usni

grip

inni

sbl

ackf

inst

atsn

out

poac

her

Xer

iere

tmus

latif

rons

blac

klip

poac

her

Xen

vrei

mus

frlac

anlls

jsbl

uesp

otte

dpo

ache

rC

iihar

icht

hys

sp.

sand

dab

-ad

ult

-ju

veni

leC

ithar

icht

hys

sord

idus

Pac

ific

sand

dab

-ad

ult

-juve

nile

Clih

sflc

hthy

sst

igm

aeus

spec

kled

sand

dab

-ad

ult

-juve

nile

Athe

rest

hea

stom

las

arro

Wto

oth

floun

der

Gly

ptoc

epha

lus

zach

irus

rev

sole

-ad

ult

-ju

veni

leH

jopo

gios

scld

esel

asso

don

tath

ead

sole

Lepi

dops

eita

bilin

eata

rock

sole

-ad

ult

-ju

veni

eLy

opse

ttaex

ilissl

ende

rso

le-

adul

t-

juve

nile

Mic

rost

omus

paci

ftcus

Dov

erso

leP

arop

hyrs

vetlu

sE

nglis

hso

le-

adul

t-

juve

nIle

Pla

ticht

hys

stel

latu

sst

arry

floun

der

Ple

uron

icht

hys

coen

osus

COso

leP

setil

chth

ysm

eian

ostic

tus

sand

sole

6.7

0-1

541

1.7

0-7

25

0.7

30

-923

3.3

0-7

00

630

635

04

6~5

031

24li)

.)

Ap

pe

nd

ixC

,T

ab

le1

2.

Num

ber

(ab

un

da

nce

)o

ffish

,b

iom

ass

(in

gra

ms)

and

ran

ge

at

Nav

yan

dPS

DD

Asite

sfo

ro

tte

rtr

aw

lca

ught

fish

bysta

tio

nan

dsp

eci

es

inP

ort

Gar

dner

onS

ep

tem

be

r11

and

15,

1986

(AU

86).

Lam

petra

fride

nial

sS

quah

sac

anth

ias

Raj

arh

ina

Hyd

rola

gus

colli

e!

Alos

asa

pkiis

sim

aC

lupe

apa

iasl

iP

oric

hthy

sno

taiu

sG

aolu

xmac

roce

phsl

usM

ic-ro

gadu

spr

oxim

us

Mer

lucc

ius

prod

uctu

s

Bro

sn,o

phyc

ism

argi

nata

Lyco

dapu

sm

ar,d

ibul

aris

Lyco

des

diap

teni

sLy

code

spa

ciflc

aC

ymal

ogas

iera

ggre

gata

Dam

alic

hihy

sva

cca

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

AU86

AU86

AU86

AU86

AU86

AU86

NAV

YO

TP

SD

DA

2PS

DD

AI

Tran

I20

STr

anI

40S

Tran

1lO

fiMA

bund

Ran

geB

iom

asa

Ran

geA

bund

Ran

geB

iom

aea

Ran

geA

bund

Ran

geB

iom

ass

Ran

geA

bund

Blo

maa

aA

bund

Bio

mas

sA

bund

Bio

maa

a

0.7

0-1

298.

30

-46

5

8.3

3-1

710

32.5

41

2-1

51

01

1-1

611.

710

5-88

01

0-2

498.

30

-13

90

18.3

3-3

648

9.2

137-

1090

0.3

0-1

4.8

0-1

42

80

0.3

0-1

990

0-2

97

00.

30-

10.

50-

112

602

40-

139

1.7

0-8

60

247

56.

35-

78.

57

-Il

0.3

0-1

30-

91

0-3

7.5

0-2

21.

30-

325

0-71

255

0.7

0-1

241.

70

-38

50.

30-

110

8.3

0-3

25

0.3

0-I

23.3

0-7

0

0-2

30.

70-

111

.30

-20

1.7

0-3

26.3

0-4

6

0.7

0-1

230

-40

335

1.3

0-2

385

0-6

10

0.7

0-2

7.8

113

.5

18

253

0.7

0-2

83.3

0-2

50

175

210

0

10

-284

.30

-11

05

760

239

2219

-26

1148

.79

50

-14

00

7.7

7-8

471.

74

25

-53

57.

72

-16

353.

26

9-7

50

490

IS

1.7

0-4

461.

70

-97

01.

70-

456

6.7

0-1

61

02

405

56.3

30

-72

9000

51

70

-10

97

03.

30-

667

1.7

0-1

34

52.

72-

676

1.7

54

0-9

60

551

017

370

348

0

415

TOTA

L1

18

.61

37

84

18.1

32

74

.224

23

56

89

60

581

86

820

22

77

.5

Ap

pe

nd

ixC

,T

ab

le12

(co

ntin

ue

d).

Fish

Spe

cies

Sci

en

tific

Nam

eC

omm

onN

ame

Lam

pet

ratri

dent

ala

Pac

ific

lam

prey

Squa

lus

acan

t’ias

spin

ydo

gfis

hR

ajar

hina

longn

ose

skat

eH

ydro

lagu

sco

uld

raIl

sh-

adul

t-

juve

nile

Alco

asa

pklis

oim

aAm

eric

ansh

adC

lupe

apa

llasi

iP

acifi

che

rrin

gP

cric

hthy

sno

tatu

spl

ainh

inm

idsh

ipm

anG

adus

maa

vcep

haki

oPa

cific

cod

Mic

roga

dus

prox

imus

Paci

ficto

mco

dad

ult

juve

nile

Mer

lucc

ius

prad

uctu

sPa

cific

hake

-ad

ult

-ju

veni

leB

rosm

ophy

cis

mar

gina

tare

dbr

otul

aLy

coda

pus

man

dibu

laris

palli

dee

lpou

tLy

code

sdi

apte

rus

blac

kee

lpou

tLy

code

spac

iflca

blac

kbel

lyee

lpou

tC

ymat

gsle

ra~g

rega

tash

iner

perc

h-

adul

t-ju

venI

le19

175

Dam

alic

hthy

sva

cca

pile

perc

h-

adul

t-

juve

nile

Ron

quilu

sjo

rdan

ino

rther

nro

nqui

lLu

mpe

nus

sagi

ttasn

ake

pric

kleb

ack

Seba

stes

mal

lger

quill

back

rock

fish

Ano

plop

oma

hmbr

lasa

blef

ish

Arte

dlus

up.

scul

pin

Das

ycot

tus

seSg

ersp

inyh

ead

scul

pin

Chl

tono

lusp

ugel

ensi

oro

ughb

ack

scul

pin

Gilb

erlid

iaoi

galu

tes

soft

scul

pin

Icel

inus

bore

alis

north

ern

scul

pin

Lept

ocot

luo

arm

atus

Pac

ific

stag

horn

scul

pin

Nau

tlchl

hyo

ocul

olas

ciat

ussa

ilfin

ocul

pin

Rad

ulin

usao

prel

lus

slim

scul

pin

Bat

hyag

onus

nlgr

ipin

nis

blac

ktn

star

snou

tpo

ache

rX

ener

etm

usla

tifro

nsbl

acki

lppo

ache

rX

ener

elm

usbi

acan

thus

blue

spot

ted

poac

her

Cilh

aric

hthy

asp

.oa

ndda

b-

adul

t-

juve

nile

Cith

aric

hlhy

sso

rdid

usPa

cific

oand

dab

-ad

ult

juve

nile

Cith

adch

thys

shEgm

aeus

spec

kled

oand

dab

-ad

ult

248

-juve

nil

117

Alh

eres

lhes

stom

las

arro

wto

oth

floun

der

Gly

ptoc

epha

lus

zach

irus

reu

sole

-ad

ult

-juve

nile

Hpi

pogi

oosc

ldes

elas

sodo

nfa

thea

dso

leLe

pido

poel

labi

li,te

ata

rock

sole

-ad

ult

-ju

veni

leLy

opse

ltaex

iliosl

ende

rso

lead

ult

-ju

veni

leM

icro

slom

uspa

cilic

usD

over

sole

Par

ophy

rsve

tlus

Eng

lish

sole

-ad

ult

-ju

veni

leP

lalic

hlhy

ssl

ella

luo

star

ryflo

unde

rPl

euro

rrlch

lhys

coen

osus

COso

leP

setli

chth

ysm

elax

ostic

tus

sand

sole

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erP

ort

Gar

dner

Por

tG

ardn

erAU

86AU

86AU

86AU

86AU

86AU

86AU

86AU

86AU

86Tr

an2

20S

Tran

240

STr

ait

2lb

STr

an4

20S

Tran

440

STr

an4

1455

PS

DD

AR

ef1

PSO

DA

Ref

2S

teS

Abu

ndB

lom

ass

Abu

ndB

iom

ees

Abu

ndB

iom

esa

Abu

ndB

iom

ess

Abu

ndB

iom

ass

Abu

ndB

iom

ess

Abu

ndB

iom

ess

Abu

ndB

iom

ess

Abu

ndB

iom

ess

282

51

510

140

0

131

53

510

818

0I

32

14.

5

369

5

28.

537

165

13

13.

52

1035

426

69

12.5

125

1614

11

252

271

332

21

221

15.

52

23.5

21

t23

485

176

131

176

26.

52

7

24.

5

13

230

138

123

.52

80

Ina

322

0

325

53

125

525

012

230

211

53

364

185

412

01

753

125

840

06

240

77.

57

575

139

.52

575

144

52

220

332

552

2685

691

513

1300

2817

554

1105

717

303

660

18.

5

TOTA

L29

692

683

18

739

49

34

.521

15

63

892

69

74

418

341

95

8.5

343

97

15

171

58

7.5

185

APPENDIX D

Abundance and biomass (and range of multiple

sample stations) of beam trawl-caught fish by

station and species in Commencement Bay,

Elliott Bay and Saratoga Passage.

186

Appendix D, Table 1. Abundance and range at multiple sample stations of beamtrawl-caught fish by station and species in CommencementBay during July 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m(range) (range)

Ratfish- adult 0.67 0-1 2— juvenile

Plainfin midshipman- adult 9- juv,

Pacific cod— adult— juvenile

Pacific hake— adult— juvenile

Pallid eelpoutBlackfin eelpoutBlackbelly eelpoutSnake pricklebackBluebarred pricklebackQuiliback rockfish— adult

— juvenileRoughback sculpin 0.3 0-1Tadpole sculpinSlim sculpinBigeye poacher 0.3 0-1Blackfin poacherArrowtooth flounder 0.3 0—1Flathead sole- adult

— juvenileRock sole— adult 7

— juvenile 5Slender sole— adult 0.3 0—1

— juvenileDover sole— adult

— juvenileEnglish sole— adult 17

— juvenile

Total 1 43

187

Appendix D, Table 2. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Commencement Bay during July 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 2 PSDDA 2 20 m(range) (range)

Ratfish- adult 143.7 0-381 770.0— juvenile

Plainfin midshipman- adult 406.5- Juv.

Pacific cod— adult 106.0— juvenile

Pacific hake- adultI’ — juvenile

Pallid eelpoutBlackfin eelpoutBlackbelly eelpoutSnake pricklebackBluebarred pricklebackQuillback rockfish— adult 175.0

— juvenileRoughback sculpin 3.3Tadpole sculpinSlim sculpinBigeye poacher 6.7 0—20Blackfin poacherArrowtooth flounder 68.5 0-205Flathead sole— adult

— juvenileRock sole- adult 304.5

— juvenile 52.5Slender sole— adult 35.0 0—105

— juvenileDover sole- adult

— juvenile 7.5English sole— adult 2,369.5

— juvenile

Total 212.0 45.0 4,191.5

188

Appendix D, Table 3. Abundance and range at multiple sample stationsof beam trawl-caught fish by station and species inElliott Bay during June 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 1-Ref PSDDA 2 PSDDA 2(range) (range)

Ratfish— adult 2.7 2—4— juvenile

Plainfin midshipman- adult 0.3 0-1- juv.

Pacific cod— adult— juvenile

Pacific hake- adult 1— juvenile

Pallid eelpoutBlackfin eelpout 0.3 0-1Blackbelly eelpout 0.7 0-2 1Snake prickleback 1 .7 0-4Bluebarred prickleback 1 .0 0-3Quillback rockfish- adult

— juvenileRoughback sculpinTadpole sculpin 0.3 0-1Slim sculpinBigeye poacherBlackfin poacherArrowtooth flounderFlathead sole— adult 1

— juvenileRock sole— adult

— juvenileSlender sole— adult 2.0 0—5 3

I’ — juvenile 2.3 0—5

Dover sole— adult 1 0.3 0—1— juvenile

English sole— adult— juvenile

Total 8 7 3.7

189

Appendix D, Table 4. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Elliott Bay during June 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 1-Ref PSDDA 2 PSDDA 2(range) (range)

Ratfish- adult 632.3 386—1,102— juvenile

Plainfin midshipman— adult 4.3 0-13.0— juv,

Pacific cod— adult— juvenile

Pacific hake— adult 258.0— juvenile

Pallid eelpoutBlackfin eelpout 5.7 0—17Blackbelly eelpout 2.5 0—7.5 41.0Snake prickleback 5.4 0-13.1Bluebarred prickleback 4.4 0-13.3Quillback rockfish— adult

— juvenileRoughback sculpinTadpole sculpin 1 .0 0-3Slim sculpinBigeye poacherBlackfin poacherArrowtooth flounderFlathead sole- adult 147.0

I’ — juvenile

Rock sole— adultU U — juvenile

Slender sole— adult 58.3 0—153.0 191.0— juvenile 9.9 0—23.3

Dover sole- adult 950.0 184.7 0—554- juvenile

English sole— adult— juvenile

Total 602.2 1595.0 823.7

190

Appendix D, Table 5. Abundance and range at multiple sample stationsof beam trawl—caught fish by station and speciesat Elliott Bay during September 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 1-Ref PSDDA 2 PSDDA 2(range) (range)

Ratfish- adult— juvenile 2 1 0—2

Plainfin midshipman- adult- Juv.

Pacific cod— adult— juvenile

Pacific hake- adult— juvenile

Pallid eelpout 0.5 0—1Blackfin eelpout 0.7 0-2Blackbelly eelpout 5.0 1—12Snake pricklebackBluebarred prickleback 5.3 0-5Quiliback rockfish- adult

— juvenileRoughback sculpinTadpole sculpinSlim sculpinBigeye poacherBlackfin poacherArrowtooth flounderFlathead sole— adult 1.3 0—3

— juvenileRock sole— adult

— juvenileSlender sole— adult 5.3 2—9 0.3 0—1

— juvenile 7.0 0—14 1Dover sole— adult 0.3 0-1 0.3 0—1

— juvenileEnglish sole— adult

— juvenile

Total 22.3 3 2.7

191

Appendix B, Table 6. Biomass (in grams) and range at multiple samplestations of beam trawl—caught fish by station andspecies in Elliott Bay during September 1986.

Location

Species PSDDA 1 PSDDA 1 PSDDA 1-Ref PSDDA 2 PSDDA 2(range) (range)

Ratfish- adultIt - juvenile 61 14.1 0—24.9

Plainfin midshipman- adultII — juv.

Pacific cod— adult— juvenile

Pacific hake- adult— juvenile

Pallid eelpout 1.7 0—5.2Blackfin eelpout 2.4 0—7.3Blackbelly eelpout 75.6 1.5—210Snake pricklebackBluebarred prickleback 15.9 14.5—17.9Quillback rockfish- adult

— juvenileRoughback sculpinTadpole sculpinSlim sculpinBigeye poacherBlackfin poacherArrowtooth flounderFlathead sole— adult 131 .3 0—330

— juvenileRock sole- adult

— juvenileSlender sole— adult 97.5 26.5—175 24.8 0—74.5

— juvenile 25.5 0—45.6 7.6Dover sole— adult 70.7 0—212 68.3 0—205

— juvenileEnglish sole— adult

— juvenile

Total 416.5 68.6 111.3

192

Appendix D, Table 7. Abundance and range of multiple sample stationsof beam trawl—caught fish by station and speciesin Saratoga Passage during June 1986.

Location

PSDDASpecies PSDDA range 80 m E 40 m E 20 rn E

Ratfish- adult 1 0—3— juvenile

Plainfin midshipman- adult- juv.

Pacific cod— adult— juvenile

Pacific hake- adult 0.3 0-1— juvenile

Pallid eelpoutBlackfin eelpout 0.3 0—1Blackbelly eelpout 0.3 0—1Snake pricklebackBluebarred prickleback 4Quillback rockfish- adult 0.3 0—1

— juvenileRoughback sculpinTadpole sculpinSlim sculpin 4Bigeye poacherBlackfin poacher 1 0—2Arrowtooth flounderFlathead sole- adult

— juvenileRock sole— adult

— juvenileSlender sole— adult 8.0 4—13 2

— juvenileDover sole— adult

— juvenile 0.3 0—1English sole— adult 3

— juvenile

Total 11.7 7 6 3

193

Appendix D, Table 8. Biomass (in grams) and range at multiple samplestations of beam trawl-caught fish by station andspecies in Saratoga Passage during June 1986.

Location

PSDDASpecies PSDDA (range) 80 m E 40 in E 20 in E

Ratfish— adult 421.7 0—1 ,265— juvenile

Plainfin midshipman- adult- juv.

Pacific cod— adult— juvenile

Pacific hake- adult 228.7 0—686— juvenile

Pallid eelpoutBlackfin eelpout 4.5 0—13Blackbelly eelpout 27.0Sna]ce pricklebackBluebarred prickleback 19.5Quillback rockfish— adult 65.3 0—196

— juvenileRoughback sculpinTadpole sculpinSlim sculpin 10.5Bigeye poacherBlackfin poacher 6.7 0—15Arrowtooth flounderFlathead sole- adult

— juvenileRock sole- adult

— juvenileSlender sole— adult 276.3 117—483 63.0 21.0

It It — juvenile 21.5

Dover sole— adult— juvenile 1.2 0—3.5

English sole- adult 52.0II — juvenile

Total 1,004.2 109.5 51.0 52.0

194

APPENDIX E

Number of flatfish per hectare

caught by otter trawl in Commencement

Bay, Elliott Bay, Saratoga Passage and

Port Gardner, shown by season, station

and species.

Ap

pe

nd

ixE

,T

ab

le1

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Com

men

cem

ent

Bay

du

rin

gsu

mm

eran

da

utu

mn

,1

98

6.

PS

DD

A1

PS

DD

A2

20m

40

m1

56

mS

UB

6A

U8

6S

U86

AU

86S

U86

AU

86S

U86

AU

86S

U86

AU

B6

Fla

tfis

hS

peci

esF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

a

spec

kled

sand

dab

-adu

lt9

arro

wto

oth

floun

der

1.35

279

13.5

rex

sole

-adu

lt36

flath

ead

sole

-adu

lt72

rock

sole

-ad

ult

slen

der

sole

-ad

ult

4.5

16.6

51.

355.

8554

13.5

Dov

erso

le-

adul

t5.

853

7.3

54.

514

.85

22.5

67.5

2754

-ju

veni

le13

.5U

i

Eng

lish

sole

-ad

ult

3.15

1.35

85.5

180

819

13.5

CO

sole

-ad

ult

4.5

TO

T.

FLA

TF

ISH

/Ha

10

.35

57

.15

7.2

22

.05

017

12

38

.51

04

427

94

.5

Ap

pe

nd

ixE

,T

ab

le2

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Ellio

ttB

ayd

urin

gsu

mm

eran

da

utu

mn

,1

98

6.

PS

DD

A1

PS

DD

A1

Ref

PS

DD

A2

PS

DD

A2

Ref

1P

SD

DA

2R

ef2

SU

86A

US

6S

U86

AU

86S

U86

AU

86S

U86

AU

86A

U86

Fla

tfis

hS

peci

esF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

rex

sole

-adu

lt5.

8510

.35

4.5

1.35

-juve

nile

5.85

flath

ead

sole

-adu

lt25

.65

111.

1567

.54.

5ro

ckso

le-

adul

t1.

35sl

ende

rso

le-

adul

t70

.65

21

0.1

554

3627

41

.85

13.5

3640

.5-ju

veni

le10

.35

32.8

54.

5D

over

sole

-ad

ult

10.3

510

.35

4.5

19.3

53

0.1

540

.59

13.5

-juve

nile

4.5

Eng

lish

sole

-ad

ult

1.35

1.35

73.3

54.

512

614

8.5

TO

T.

FLA

TF

ISH

/Ha

12

8.7

38

2.0

51

35

40

.54

7.7

14

6.7

58

.517

12

02

.5

Ap

pe

nd

ixE

,T

ab

le3

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Sa

rato

ga

Pa

ssa

ge

du

rin

gsu

mm

er,

19

86

.

PSD

DA

PSD

DA

Ref

2Om

E4O

mE

40m

W8O

mE

SU

86

SU

86

SU

86S

U8

6S

U86

SU

86

Fla

tfis

hS

pe

cie

sF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

a

rex

sole

-adu

lt4.

5fla

thea

dso

le-a

dult

rock

sole

-ad

ult

922

.5sl

ende

rso

le-

adul

t12

.15

13.5

4.5

18D

over

sole

-ad

ult

4.5

Engl

ish

sole

-ad

ult

1831

.59

31.5

TOT.

FLA

TFIS

H/H

a1

2.1

51

3.5

275

8.5

95

8.5

Ap

pe

nd

ixE

,T

ab

le4

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Po

rtG

ard

ne

rd

urin

gw

inte

r,1

98

6.

Tra

nsec

t1

Tran

sect

1T

rans

ect

1Tr

anse

ct2

Tra

nsec

t2

Tran

sect

2Tr

anse

ct4

Tran

sect

4T

rans

ect

4N

avy

PS

DD

A2

PS

DD

A1

20

S4

09

lOO

M2

09

40

S1

10

82

0S

40

81

45

SC

omm

onN

ame

Fis

h/H

aF

ish/

Ha

Fis

h/H

aF

ish/

Ha

Fis

h/H

aF

ish/

Ha

Fis

h/H

aF

ish/

Ha

Fis

h/H

aF

ish/

Ha

Fis

h/H

aF

ish/

Ha

sarid

dab

-ad

ult

00

09

40.5

04.

554

018

00

-ju

veni

le0

00

4.5

370

00

022

.50

0Pa

ciiic

sand

dab

00

00

00

00

00

4.5

0sp

eckl

edsa

ndda

b-

adul

t0

00

00

00

00

011

70

-juve

nile

00

00

00

00

00

4.5

0ar

row

toot

hflo

unde

r0

01.

350

4.5

00

4.5

00

00

rex

solo

-ad

uft

92.

251.

350

09

04.

54.

50

00

-juve

nile

00

00

04.

50

00

00

0fla

thoa

dso

le11

5.65

2.25

04.

522

.50

00

018

4.5

0ro

ckso

lo-

adul

t1.

350

1.35

958

.50

4.5

76.5

00

990

-juve

nile

00

09

22.5

09

4.5

04.

540

.50

slen

der

sole

-ad

ult

136.

565

.25

117

00

4.5

027

4.5

13.5

063

-juve

nile

40.5

00

00

4.5

00

00

00

Dove

rso

le32

.85

22.5

16.6

50

09

04.

549

.50

012

6En

glis

hso

le-

adul

t49

9.5

15.7

531

.54.

576

.511

70

301.

536

063

00

-juve

nile

3.15

00

4.5

706.

50

024

30

4.5

00

star

ryflo

unde

r0

00

00

00

00

00

0DO

sole

00

00

4.5

00

00

4.5

00

sand

sole

00

00

00

00

00

00

TO

T.

FLA

TF

ISH

/He

83

8.5

10

81

69

24

59

73

14

8.5

18

72

09

4.5

85

.59

00

18

9

Ap

pe

nd

ixE

,T

ab

le5

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Po

rtG

ard

ne

rd

urin

gsp

rin

g,

1986

.

Tra

nse

ctI

Tra

nse

ct1

Tra

nse

ct1

Tra

nse

ct2

Tra

nse

ct2

Tra

nse

ct2

Tra

nse

ct4

Tra

nse

ct4

Tra

nse

ct4

NA

VY

PS

DD

A2

PS

DD

A1

20

S4

05

lOO

M2

0S

40

51

10

52

05

40

S1

45

5C

omm

onN

ame

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

uand

dab

-ad

ult

00

00

90

13.5

00

4.5

180

-juvo

nilo

00

04.

50

00

00

90

0Pa

cific

sand

dab

00

00

00

00

00

00

spec

kled

sand

dab

-ad

ult

00

00

00

00

00

00

-juve

nile

00

00

00

00

00

00

arro

wto

oth

floun

der

00

00

00

00

00

00

rex

solo

-ad

ult

25

.65

01.

350

09

00

00

00

-juve

nile

1.35

03

.15

00

00

00

00

0ila

thoa

dso

le2

5.6

50

00

09

00

00

00

rock

solo

-ad

ult

00

00

4.5

04.

522

.50

049

.50

-juve

nile

00

00

00

00

09

90

slen

der

solo

-adu

lt76

.521

.15

39.1

50

067

.50

09

00

4.5

-juve

nile

7.65

00

00

00

00

00

0D

ovor

uole

13.5

273.

150

549

00

90

00

Engl

ish

sole

-ad

ult

13

6.3

51

2.1

53

.15

09

27

4.5

013

.54.

50

49

.50

-juve

nile

00

00

00

00

00

360

star

ryflo

unde

r0

00

00

00

00

00

0co

sole

00

00

00

00

00

00

sand

sole

00

00

4.5

04.

50

00

00

TO

T.

FLA

TF

ISH

/Hs

28

6.6

56

0.3

49

.95

4.5

813

69

22

.53

62

2.5

22

.51

62

4.5

Ap

pe

nd

ixE

,T

ab

le6

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sp

er

he

cta

reca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Po

rtG

ard

ne

rd

urin

gsu

mm

er,

19

86

.

Tra

nsec

t1

Tra

nsec

t1

Tra

nsec

t1

Tra

nsec

t2

Tra

nsec

t2

Tra

nsec

t2

Tra

nsec

t4

Tra

nsec

t4

Tra

nsec

t4

PS

DD

AP

SD

DA

Com

mon

Nam

eN

AVY

PS

DD

A2

PS

OD

A1

20S

405

lOO

M20

54

05

11

05

20S

40S

145S

Ref

1R

ef2

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/He

Fis

hlH

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

sand

dab

-ac

ctt

00

00

4.5

04.

50

09

00

00

-juve

nile

00

00

00

00

013

.50

00

0P

acltc

sand

dab

00

00

00

00

00

00

00

spec

kled

sand

dab

-adu

lt0

00

00

00

00

00

00

0-ju

venf

e0

00

00

00

00

00

00

0ar

row

toot

hflo

unde

r0

00

00

00

00

00

00

0re

xso

le-a

dult

00

00

00

013

.50

00

00

0-ju

veni

le0

00

0I)

00

00

00

00

0fa

thea

dco

le10

.35

00

00

00

4.5

00

00

00

rock

sole

-ad

ult

00

00

270

4.5

22.5

00

00

00

-juve

nile

00

00

00

4.5

00

00

00

0sl

ende

rso

le-

adul

t22

.59

12.1

50

09

09

00

04.

59

0-ju

venf

e0

01.

350

4.5

4.5

00

00

00

00

Dov

erso

le-

adul

t3.

1530

.15

180

04.

50

4.5

120

4.5

1827

18-ju

veni

le0

1.35

3.15

00

00

00

00

4.5

00

Eng

lish

sole

-ad

ult

129.

1555

.35

13.5

4.5

67.5

126

999

1827

00

22.5

22.5

-juve

nile

00

00

00

00

00

00

00

star

ryfo

unde

r0

00

00

04.

50

00

00

00

CO

eds

00

00

00

00

00

00

00

sand

sole

00

00

00

00

00

00

00

TOT.

FLA

TFiS

H/H

a1

65

.15

99_O

S4

51

54_

a1

03

.51

44

271

53

304

95

4.5

275

8.5

40

.5

Ap

pe

nd

ixE

,T

ab

le7

.N

umbe

ro

fe

ach

fla

tfis

hsp

ecie

sca

ug

ht

by

ott

er

tra

wl

at

ea

chsta

tio

nin

Po

rtG

ard

ne

rd

urin

ga

utu

mn

,1

98

6.

Tra

nsec

tI

Tra

nsec

tI

Tra

nsec

t1

Tra

nsec

t2

Tra

nsec

t2

Tra

nsec

t2

Tra

nsec

t4

Tra

nsec

t4

Tra

nsec

t4

PS

DD

AP

SD

DA

P50

0AC

omm

onN

ame

NAV

YP

SD

DA

2P

SD

DA

12

00

40S

lOO

M20

540

Slb

S2

0$

405

14

55

Ref

1R

ef2

Sta

EF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

Fis

h/H

aF

ish

/Ha

sand

dab-

adul

t0

00

00

00

00

00

00

00

-juve

nile

00

00

00

00

00

00

00

0P

acili

csa

ndda

b-

adul

t0

00

00

00

00

00

00

00

-juve

nIle

00

00

4.5

00

90

00

00

00

spec

kled

sand

dab

-ad

ult

00

00

90

94.

50

4.5

90

00

0-ju

veni

le0

00

00

04.

50

00

00

00

0ar

row

toot

hflo

unde

r0

00

00

00

00

00

00

00

rex

sole

adul

t0

3.15

00

4.5

90

04.

50

13.5

00

00

-juve

nile

00

00

00

00

00

00

00

0lla

thea

dsol

e4.

50

00

00

00

00

00

00

0ro

ckso

le-

adul

t0

00

022

.50

013

.50

13.5

22.5

00

00

-juve

nile

00

00

90

4.5

00

90

00

00

slen

der

sole

-ad

ult

9934

.65

34.6

50

180

913

.518

018

4.5

13.5

3627

-juve

nile

00

00

4.5

00

00

00

00

31.5

0D

over

sole

07.

657.

650

09

00

31.5

04.

50

94.

59

Engl

Ish

sole

-ad

ult

253.

3514

.85

12.1

522

.576

.513

.513

.523

427

58.5

126

018

31.5

13.5

-juve

nile

00

00

00

00

04.

50

00

00

star

ryS

ound

er0

00

4.5

00

00

00

00

00

0C

Oed

e0

00

00

00

00

00

00

00

sand

sole

00

00

00

00

00

00

00

0

TOT.

FLA

TFiS

H/H

a3

56

.85

60

.35

4.4

527

14

85

31

.54

0.5

27

4.5

8190

19

3.5

45

40

.51

03

54

8.5