Diversity of Terrestrial Gastropods (Molluscsa, Gastropoda ...Nine further spe-cies were typical for...

469

ACTA ZOOLOGICA BULGARICAActa zool. bulg., 70 (4), 2018: 469-477

*Corresponding author: [email protected]

Zoogeography and FaunisticsResearch Article

IntroductionThe Prespa National Park is included in the network of protected territories in Albania and is an integral component of the planned large trans-boundary con-servation area occupying the Lake Prespa drainage basin situated in Albania, Greece and the Republic of Macedonia. These states have declared their will-ingness and commitment to unite their efforts in or-der to protect the unique natural values of the two lakes, Great Prespa Lake and Small Prespa Lake, and the surrounding mountainous ecosystems on Mali i Thatë (Galichica), Suva Gora and Pelister Mountains (Hristovski et al. 2014). Comprehensive informa-tion on the diversity and the conservation status of the terrestrial invertebrate fauna in this region is lacking. It has been predicted that the invertebrate fauna of the Prespa National Park in Albania is di-verse, with some species being important from the conservation point of view, including many endemic and relict taxa (Hristovski et al. 2014).

Previous studies have recorded 13 terrestrial gas-tropod species and subspecies from the the territory of the Prespa National Park and its vicinity: Helix schlae-flii, Lindholmiola corcyrensis, Lindholmiola lens (A. Ferrusac, 1832), Monacha frequens, Montenegrina dofleini dofleini, M. d. prespaensis, M. d. wagneri, M. hiltrudae costulata, M. h. desaretica, M. h. fusca, M. h. sattmanni, Zebrina detrita and Wladislawia tha-teensis (Dhora & Welter-Schultes 1996, Welter-Schultes 1996, Fehér & Drimmer 2004, Erőss et al. 2006, Nordsieck 2008, Subai 2012, Neubert 2014, Fehér & Szekeres 2016).

The aim of the present article is to provide a survey of the diversity of the terrestrial gastropods of the Prespa National Park (Albania) by sampling and identification of new collections and summarising published data, to estimate the conservation value of the malacofauna in the region and to identify the most important habitats for terrestrial molluscs.

Diversity of Terrestrial Gastropods (Molluscsa, Gastropoda) in the Prespa National Park, Albania

Ivaylo K. Dedov1*, Zoltán Fehér2,3, Miklós Szekeres4 & Ulrich E. Schneppat5

1 Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Gagarin Street, 1113 Sofia, Bulgaria; E-mail: [email protected] Natural History Museum, Vienna, 3rd Zoological Department, Burgring 7, A-1010, Vienna, Austria3 Hungarian Natural History Museum, Baross 13, H-1088 Budapest, Hungary; E-mail: [email protected] Institute of Plant Biology, Biological Research Centre, Hungarian Academy of Sciences, Temesvari krt. 62, H-6726 Szeged, Hungary; E-mail: [email protected] Sennereiweg 8, CH-7074 Churwalden-Malix, Switzerland; E-mail: [email protected]

Abstract: In total, 51 terrestrial gastropod taxa are recorded from the Prespa National Park, Albania. Seven species are local endemics for the region of Lake Prespa and adjacent mountains, eight are West-Balkan endemics and seven are Balkan endemics. There are no species considered threatened according to the IUCN cat-egories and only one species, Orcula wagneri, is in the category Near Threatened. Of the habitats studied, limestone rocks and rocky grasslands are the most important for the diversity of the gastropods in the region.

Key words: Albania, Prespa National Park, terrestrial gastropod fauna, distribution

470

Dedov I. K., Z. Fehér, M. Szekeres & U. E. Schneppat

Materials and MethodsThe materials were collected at 41 localities (Table 1, Fig. 1) between 2007 and 2016 in the course of vari-ous field trips. Hand-collecting and the soil-sifting methods were used. The gastropods were preserved in 70% ethanol or kept as dry shells in the collec-tions of the Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, Sofia (IBER-BAS), the Hungarian Natural History Museum, Budapest (HNHM) and the Natural History Museum, Vienna (NHMW). The morphological and anatomical examinations were carried out using ster-eomicroscope.

ResultsThe present study adds 39 species newly re-

corded for the territory of the Prespa National Park (Albania) and, with the present publications, 51 mol-lusks taxa are known for the region (Table 2). One species (Lindholmiola lens) was not confirmed. Several species of high importance for biodiversity conservation were found. Seven gastropods are lo-cal endemics for the region of Lake Prespa and ad-jacent mountains, eight are West-Balkan endemics and seven are Balkan endemics. Only one species, Orcula wagneri, is included in IUCN categories for threatened species as Near Threatened.

Out of the species recorded, 19 were collected nearby Lake Prespa only from altitudes between 850 and 900 m (Oxyloma elegans, Lauria cylindracea, Pyramidula cephalonica, P. pusilla, Truncatellina claustralis, T. cylindrica, T. rothi, Jaminia quad-ridens, Cecilioides acicula, Montenegrina dofleini prespaensis, M. hiltrudae desaretica, Mediterranea inopinata, Tandonia sp., Oligolimax sp., aff. Semilimacella, Limacus flavus, Limax cf. cone-menosi, L. cf. corcyrensis and Cernuella virgata). Twenty-one species occurred in the vicinity between the lake and the rocks and rocky pastures above the forest zone of the mountains at altitudes 850 – above 2200 m (Chondrina arcadica, Chondrula macedon-ica, C. microtragus, Merdigera obscura, Zebrina detrita, Alinda biplicata distincta, Montenegrina hiltrudae costulata, M. h. sattmanni, Strigilodelima conspersa, Vitrea cf. botterii, Mediterranea sp., Morlina glabra, Vitrina sp., Lehmannia cf. brunneri, Lindholmiola corcyrensis, Candidula rhabdotoides, Monacha frequens, Xerolenta obvia, Helix lucorum, H. philibinensis and H. schlaeflii). Nine further spe-cies were typical for the highest limestone areas of the mountains (900 – above 2200 m a.s.l.) and did not occur near the lake (Cochlostoma sp., Orcula

wagneri, Montenegrina dofleini dofleini, M. d. wag-neri, M. hiltrudae fusca, Tandonia macedonica, Limax cf. graecus, L. cf. maximus and Wladislawia thateensis).

Although the forest in the Prespa National Park is growing on limestone, it has not been proved as a habitat especially rich in terrestrial gastropod species by the present study. Only three species were found in such habitats and one of them (Lauria cylindracea) occurred also in Buxus sempervirens shrublands and highly degraded and modified Querco-Carpinetum orientalis forest around Lake Prespa (Table 3).

DiscussionThe biodiversity value of the terrestrial gastropod fauna of the Prespa National Park (Albania) is high, though its relatively small territory (277.5 km2): 51 terrestrial gastropods have been found up to now, including a high number of endemics – 22 species or 43% (Table 2). According to our results, lime-stone rocks and rocky grasslands are the most im-

Fig. 1. Sampling localities in the Prespa National Park, Albania. Numbers correspond to those in Table 1.


471

Tabl

e 1.

Loc

aliti

es (P

resp

a N

atio

nal P

ark)

, Abb

revi

atio

n: C

oll.

No.

– c

atal

og n

umbe

r of t

he c

olle

ctio

n of

Inst

itute

of B

iodi

vers

ity a

nd G

ener

al R

esea

rch,

Bul

garia

n A

cade

my

of S

cien

ces (

IBER

, BA

S); H

NH

M –

col

lect

ion

of th

e H

unga

rian

Nat

iona

l His

tory

Mus

eum

; NH

MW

– c

olle

ctio

n of

the

Nat

ural

His

tory

Mus

eum

, Vie

nna.

№C

oll.N

oLo

calit

ies

Coo

rdin

ates

N/E

Alti

tude

(m

)H

abita

ts

120

412

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa L

ake s

hore

), vi

ll. K

alla

mas

(= v

ill. T

umin

ec),

24 V

201

3, le

g. D

edov

; 17

X 2

014,

leg.

Feh

ér, H

arin

g, Ja

ksch

, Sat

tman

n; 2

9 VI 2

015,

leg.

Erő

ss,

Fehé

r, G

rego

40.8

919°

20.

9395

°85

0-85

6lim

esto

ne ro

cks w

ith b

ushe

s (Pr

uno

web

bii–

Ju-

nipe

retu

m ex

celsa

e)

220

413

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(=G

reat

Pre

spa L

ake s

hore

), 24

V 2

013,

leg.

Ded

ov40

.872

0° 2

0.92

60°

855

limes

tone

rock

s with

bus

hes

320

414

Kor

çë D

istric

t, vi

ll. G

oric

a e M

adhe

(= v

ill. G

orna

Gor

ica)

, nea

r chu

rch,

24

V 2

013,

leg.

Ded

ov40

.893

4° 2

0.90

17°

921

open

terra

in

420

415

2041

6M

ali i

Tha

të (=

Suv

a G

ora

Mts.

), ne

ar th

e M

aced

onia

n bo

rder

, S o

f the

bor

der-c

ross

ing

„Sub

o-tin

o“, 2

5 V

201

3, 2

4 V

I 201

3, le

g. D

edov

40.9

098°

20.

8950

°97

5ro

cky

limes

tone

mea

dow

bet

wee

n fo

rests

of

Que

rcus

frai

netto

and

Que

rcus

cerr

is

5 20

417

2041

8M

ali i

Tha

të (=

Suv

a Gor

a Mts.

), 25

V 2

013,

24

VI 2

013,

leg.

Ded

ov40

.898

0° 2

0.86

47°

1311

Festu

co h

eter

ophy

llae-

Fage

tum

, und

er lo

g

620

419

2042

0M

ali i

Tha

të (=

Suv

a Gor

a Mts.

), “P

ikin

a Vod

a” ar

ea, 2

5 V

201

3, 2

4 V

I 201

3, le

g. D

edov

40.9

055°

20.

8541

°15

65lim

esto

ne m

eado

w w

ith s

catte

red

bush

es a

nd

trees

7 20

421

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

“The

tow

er o

f the

war

” are

a, 25

V 2

013,

leg.

Ded

ov40

.912

2° 2

0.84

98°

1736

alpi

ne ro

cky

limes

tone

mea

dow

8 20

422

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

25 V

201

3, le

g. D

edov

40.9

106°

20.

8477

°17

31Fa

gus f

ores

t

9 20

423

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa

Lake

shor

e), y

ard

of th

e Adm

inist

ratio

n of

Pr

espa

Nat

iona

l Par

k, 2

6 V

201

3, le

g. D

edov

40.8

818°

20.

9207

°86

6 m

eado

w

10

2042

4K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës së

Vog

ël (=

Les

ser P

resp

a Lak

e), v

ill. T

ren,

in fr

ont o

f the

cave

„S

hpel

a e T

reni

t“, 2

6 V

201

3, le

g. D

edov

40.6

722°

20.

9868

°85

6lim

esto

ne ro

cks

1120

425

2042

6K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës së

Vog

ël (=

Les

ser P

resp

a Lak

e), b

etw

een

vill.

Shu

ec an

d vi

ll.

Tren

, ban

k of

the l

ake,

26 V

201

3, 2

4 V

I 201

3, le

g. D

edov

; 13

V 2

016,

leg.

Pap

arist

o40

.684

7° 2

0.99

84°

863-

930

Buxu

s sem

perv

irens

on

limes

tone

rock

-bas

e

12

2042

7K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës së

Vog

ël (=

Less

er P

resp

a La

ke),

„Rak

cko“

are

a, 26

V 2

013,

le

g. D

edov

40.7

245°

20.

9768

°11

09

limes

tone

pas

ture

with

scat

tere

d bu

shes

13

2042

820

429

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa

Lake

shor

e), v

ill. G

llobo

çeni

(= v

ill. G

lobo

-ch

eni),

26

V 2

013,

23-

25 V

I 201

3, le

g. D

edov

40.8

574°

20.

9429

°86

5lim

esto

ne ro

cks

1420

430

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(=G

reat

Pre

spa L

ake s

hore

), ne

ar v

ill. Z

aros

hkë (

= vi

ll. Z

rnos

-ko

), Ra

lnik

area

, 27

V 2

013,

leg.

Ded

ov40

.766

8° 2

0.92

77°

868

Buxu

s sem

perv

irens

shru

blan

d (h

ighl

y de

grad

ed

and

mod

ified

Que

rco-

Carp

inet

um o

rient

alis

for-

est)

on li

mes

tone

rock

s

1520

431

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa

Lake

sho

re),

near

the

chur

ch o

f St.

Tana

s, N

E of

vill

. Liq

enas

(= v

ill. P

uste

c), 2

7 V

201

3, le

g. D

edov

; 1 k

m N

E of

vill

. Liq

enas

(=v

ill.

Puste

c), S

v. A

tana

s and

Ant

oni V

elik

i chu

rch,

at L

iqen

i i P

resp

es, 0

2 V

II 20

03, l

eg. E

rőss

, Feh

ér,

Kon

tschá

n, M

urán

yi; 2

9 V

I 201

3, le

g. E

rőss

, Feh

ér, G

rego

40.7

918°

20.

9134

°85

0-87

0M

orus

spec

. tre

e, lim

esto

ne ro

cks

16

2043

2K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës (=

Gre

at P

resp

a La

ke sh

ore)

, vill

. Gllo

boçe

ni (=

vill

. Glo

bo-

chen

i), b

oat h

arbo

r, 23

VI 2

013,

leg.

I. D

edov

; 30

VI 2

015

leg.

Erő

ss, F

ehér

, Gre

go40

.859

3° 2

0.94

92°

850-

855

limes

tone

rock

s

17

2043

3K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës (=

Gre

at P

resp

a Lak

e sho

re),

vill.

Kal

lam

as (=

vill

. Tum

inec

), 23

VI 2

013,

leg.

Ded

ov

40.8

942°

20.

9373

°85

7m

eado

ws

472


№C

oll.N

oLo

calit

ies

Coo

rdin

ates

N/E

Alti

tude

(m

)H

abita

ts

18

2043

4K

orcë

Dist

rict,

at at

Liq

eni i

Pre

spës

së V

ogël

(= L

esse

r Pre

spa L

ake)

, nea

r vill

. Buz

ëliq

en (=

vill

Za

grad

ets)

, 24

VI 2

013,

leg.

, Ded

ov40

.676

7° 2

1.00

38°

866

limes

tone

rock

s, Bu

xus s

empe

rvire

ns

1920

435

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

near

Cer

je (=

vill

. Cer

je),

24 V

I 201

3, le

g. D

edov

40

.752

3° 2

0.94

34°

1126

rock

y lim

esto

ne m

eado

ws,

Que

rcus

troj

ana

2020

436

2043

7M

ali i

Tha

të (=

Suv

a Gor

a Mts.

), Pl

laja

e Pu

sit p

eak,

25

VI 2

013,

leg.

Ded

ov

40.8

798°

20.

8418

°2

20

4-

2269

limes

tone

rock

s nea

r pat

ch o

f sno

w

2120

438

Mal

i i T

hatë

(=Su

va G

ora

Mts.

), up

to v

ill. G

llobo

çeni

(= v

ill. G

lobo

chen

i), 2

3 IX

201

3, le

g.

Ded

ov40

.852

3° 2

0.94

71°

918

Que

rco

troya

nae-

Carp

inet

hum

orie

ntal

ae,

on

limes

tone

rock

bas

e

2220

439

Mal

i i T

hatë

(=Su

va G

ora

Mts.

), up

to v

ill. G

llobo

çeni

(= v

ill. G

lobo

chen

i), 2

3 IX

201

3, le

g.

Ded

ov40

.845

6° 2

0.94

92°

1035

rock

y lim

esto

ne m

eado

ws

2320

440

2044

1M

ali i

Tha

të (=

Suva

Gor

a Mts.

), “T

he to

wer

of t

he w

ar” a

rea,

24 IX

201

3, le

g. D

edov

40.9

138°

20.

8433

°1

81

4-

1830

alpi

ne m

eado

w o

n lim

esto

ne an

d ro

cky

slope

s

2420

442

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa

Lake

shor

e), r

ock

chur

ch n

ear M

oste

ts, 2

5 IX

201

3, le

g. D

edov

;K

orçë

Dist

rict,

cave

tem

ple

ca. 4

km

S o

f vill

. Gllo

boçe

ni (=

vill

. Glo

boch

eni),

at L

iqen

i i P

re-

spës

, 30

VI 2

015,

leg.

Erő

ss, F

ehér

, Gre

go

40.8

419°

20.

9615

°85

0-87

0lim

esto

ne ro

cks

25

2044

3K

orcë

Dist

rict,

at L

iqen

i i P

resp

ës (=

Gre

at P

resp

a Lak

e sho

re),

Ishu

lli M

alig

rad

(= is

land

“Mal

G

rad”

), 25

IX 2

013,

leg.

Ded

ov40

.791

5° 2

0.93

31°

859

limes

tone

rock

s, bu

shes

2620

444

Kor

cë D

istric

t, at

Liq

eni i

Pre

spës

(= G

reat

Pre

spa L

ake s

hore

), ro

ck ch

urch

“St.

Mar

ena”

, nea

r D

labo

ko, c

a. 4

km E

of v

ill. K

alla

mas

, 25

IX 2

013,

leg.

Ded

ov; 2

9 V

I 201

5, le

g. E

rőss

, Feh

ér,

Gre

go,

40.8

869°

20.

9727

°85

0-85

3lim

esto

ne ro

cks

27H

NH

MK

orcë

Dist

rict,

vill.

Liq

enas

(= v

ill. P

uste

c), b

y th

e sh

ore

of L

iqen

i i P

resp

es (=

Gre

at P

resp

a La

ke),

18 IX

199

2, le

g. F

ehér

; 02

VII

2003

, leg

. Erő

ss, F

ehér

, Kon

tschá

n, M

urán

yi40

.789

7° 2

0.90

87°

844-

860

lacu

strin

e dr

ift m

ater

ial,

degr

aded

rude

ral v

eg-

etat

ion

and

limes

tone

rock

s

28H

NH

MK

orcë

Dist

rict,

E of

vill

. Liq

enas

(= v

ill. P

uste

c), I

shul

l i V

ogël

, sho

re o

f Liq

eni i

Pre

spes

(=

Gre

at P

resp

a Lak

e), 1

7 V

III 2

007,

leg.

Feh

ér, T

amás

40.7

915°

20.

9324

° 86

0lim

esto

ne ro

cks

29N

HM

WK

orçë

Dist

rict,

vill.

Gllo

boçe

ni (=

vill

. Glo

boch

eni),

N s

ide,

17 X

201

4, le

g. F

ehér

, Har

ing,

Ja

ksch

, Sat

tman

n40

.858

° 20.

9426

°86

0lim

esto

ne ro

cks

30N

HM

WK

orçë

Dist

rict,

betw

een

vill.

Gllo

boçe

ni (=

vill

. Glo

boch

eni)

and

vill.

Liq

enas

(= v

ill. P

uste

c), 1

km

N o

f the

Die

las j

unct

ion,

17

X 2

014,

leg.

Feh

ér, H

arin

g, Ja

ksch

, Sat

tman

n40

.812

2° 2

0.90

45°

1040

rock

y gr

assla

nd

31N

HM

WK

orçë

Dist

rict,

2 km

SE

of K

alla

mas

, at L

iqen

i i P

resp

ës (=

Gre

at P

resp

a Lak

e), 2

9 V

I 201

5, le

g.

Erős

s, Fe

hér,

Gre

go40

.880

9° 2

0.95

90°

850

limes

tone

rock

s

32N

HM

WK

orçë

Dist

rict,

sinkh

ole c

ave 1

km

S o

f Gor

icë e

Vog

ël, 2

9 V

I 201

5, le

g. E

rőss

, Feh

ér, G

rego

40.8

699°

20.

9295

°85

0am

ong

boul

ders

at th

e ope

ning

of t

he ca

ve

33H

NH

MK

orçë

Dist

rict,

Mal

i i T

hatë

(= S

uva

Gor

a M

ts.),

ca. 4

.3 k

m E

of t

he v

ill. P

odgo

rije,

ca. 1

.3 k

m

E of

the p

eak

“Bre

gu i

Stan

it”, 2

0 V

200

7, le

g. B

arin

a, Pi

fkó,

Ném

eth

40.8

255°

20.

8528

°18

25lim

esto

ne ro

cks

34H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

ca. 3

.9 k

m E

of t

he v

ill. P

odgo

rije,

betw

een

the

peak

s “Br

egu

i Sta

nit”

and

“Mez

a”, 2

0 V

200

7, le

g. B

arin

a, Pi

fkó,

Ném

eth

40.8

179°

20.

8494

°18

58lim

esto

ne ro

cks

Tabl

e 1.

Con

tinua

tion.


473

Tabl

e 1.

Con

tinua

tion.

№C

oll.N

oLo

calit

ies

Coo

rdin

ates

N/E

Alti

tude

(m

)H

abita

ts

35H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva

Gor

a M

ts.),

ca. 3

.2 k

m E

of t

he v

ill. P

odgo

rije,

ca. 1

.0 k

m

NW

of M

ount

“Mez

a” an

d 1.

5 km

S o

f Mou

nt “S

tani

”, 20

V 2

007,

leg.

Bar

ina,

Pifk

ó, N

émet

h40

.814

3° 2

0.84

02°

1863

limes

tone

bou

lder

s on

gras

sland

36H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva

Gor

a M

ts.),

ca. 2

.9 k

m N

W o

f the

vill

. Liq

enas

(= v

ill.

Puste

c), c

a. 2.

2 km

E/N

E of

the p

eak

“Buz

e K

orut

es”,

22 V

200

7, le

g. B

arin

a, Pi

fkó

40.8

024°

20.

8741

°17

46ka

rstic

shru

blan

d

37H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva

Gor

a M

ts.),

ca. 4

.3 k

m N

W o

f the

vill

. Liq

enas

(= v

ill.

Puste

c), c

a. 70

0 m

S o

f a p

eak,

22

V 2

007,

leg.

Bar

ina,

Pifk

ó40

.818

3° 2

0.87

28°

1913

limes

tone

rock

s on

a scr

ee sl

ope

38H

NH

MK

orcë

dist

rict,

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

ca. 4

.3 k

m N

W o

f vill

. Liq

enas

(= v

ill. P

uste

c),

ca. 4

00 m

E/S

E of

a pe

ak, 2

2 V

200

7, le

g. B

arin

a, Pi

fkó

40.8

220°

20.

8789

° 20

35lim

esto

ne ro

cks o

n a s

cree

slop

e

39H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva

Gor

a M

ts.),

ca. 1

.7 k

m N

of t

he v

ill. Z

vezd

ë, on

Mou

nt

Zvez

da, 2

.1 k

m S

E of

the p

eak,

25

V 2

007,

leg.

Bar

ina,

Pifk

ó, N

émet

h40

.741

9° 2

0.86

35°

1179

limes

tone

rock

s on

gras

sland

40H

NH

MK

orcë

Dist

rict,

Mal

i i T

hatë

(= S

uva G

ora M

ts.),

ca. 2

.7 k

m N

/NW

of t

he v

ill. Z

vezd

ë, on

Mou

nt

Zvez

da, c

a. 50

0 m

SW

of t

he p

eak,

25

V 2

007,

leg.

Bar

ina,

Pifk

ó, N

émet

h40

.756

5° 2

0.84

95°

1718

limes

tone

rock

s

41H

NH

MM

ali i

Tha

të (=

Suva

Gor

a Mts.

), pa

th fr

om G

oric

ë to

the r

idge

, 200

8, le

g. B

arin

a, Pi

fkó,

Ném

eth

40.8

485°

20.

8624

°17

65lim

esto

ne ro

cks

portant habitats for the diversity of the gastropods in the region. Because of total clearing of forests in the border areas of Albania in the near past, in most cases the forests are young and degraded and not rich in species (Table 2). In some small old patches of the deciduous forest in the park, some forest species could be expected, which are present in the Macedonian part of the mountains. These are Acanthinula aculeata (Müller, 1774), Aegopinella minor (Stabile, 1864), Cochlodina laminata (Montagu, 1803), Dinarica serbica (Rossmässler, 1836), Punctum pygmaeum (Draparnaud, 1801) and Triloba thaumasia (Sturany, 1907). In Albania, very close to the border of the park, the endemic species Chilostoma (Josephinella) byshekensis (Knipper, 1941) has been found and perhaps it occurs in the protected territory, too.

According Dhora & Welter-Schultes (1996), Lindholmiola lens occurs in the region. However, the present study does not confirm this information. As far as Goricë i Madhe is the only known local-ity of this species in Albania and this site has been examined by us at several occasions in the course of the present survey, we suggest to remove L. lens from the Albanian checklist of terrestrial gastropods.

Fehér & Szekeres (2016) have identified the population of Montenegrina from St. Marena church as M. dofleini prespaensis but newly collected data show that it is actually M. hiltrudae desaretica; how-ever, some individuals show transitive conchologi-cal features, which might be a sign of hybridization between the two species.

With the present study, the number of the known terrestrial gastropod species from the region is almost tripled. Nevertheless, many questions still remain. In some of the localities, the specimens of Chondrula macedonica are much more slender and without basal plica. The phenomenon of reduction of apertural structures generally is observed at higher altitudes and represents a local variation of the spe-cies but remains without a clear and well-supported explanation. Such forms with reduced plicae are known in the family Clausiliidae as “oreinos-forms” (Nordsieck 2008) but in the family Enidae the phe-nomenon is still weekly studied. From subjective personal observation, we think that many potential predators (e.g. Diptera: Sciomyzidae; Coleoptera: Elateridae, Drilini) of snails with mouth plicae do not go as high up as the snail species does. The pli-cae are generally thought to prevent predators from entering the snail shell through the apertural open-ing. If there is no or only very little pressure by these predators, eventually there is no reason anymore for developing such plicae.

474


Tabl

e.2.

Lis

t of t

he g

astro

pods

in P

resp

a N

atio

nal P

ark

(Alb

ania

). Th

e nu

mbe

rs o

f loc

aliti

es c

orre

spon

d to

thos

e in

the

first

col

umn

of th

e Ta

ble

1. A

bbre

viat

ion:

ID

– R

egis

-tra

tion

num

ber o

f mol

lusk

s in

colle

ctio

n of

Inst

itute

of B

iodi

vers

ity a

nd E

cosy

stem

Res

earc

h, B

ulga

rian

Aca

dem

y of

Sci

ence

s; H

NH

M –

Hun

garia

n N

atur

al H

isto

ry M

useu

m,

Bud

apes

t; N

HM

W –

col

lect

ion

of th

e N

atur

al H

isto

ry M

useu

m, V

ienn

a; M

SS -

mili

eu so

ubte

rrai

n su

perfi

ciel

; BL

– B

alka

n en

dem

ics,

WB

L –

Wes

tern

Bal

kan

ende

mic

s, LE

–

Loca

l end

emic

s (en

dem

ic fo

r the

regi

on a

roun

d th

e la

ke P

resp

a, in

clud

ing

adja

cent

mou

ntai

ns);

IUC

N –

incl

uded

in “

The

IUC

N R

ed L

ist o

f Thr

eate

ned

Spec

ies”

(Eur

ope)

as

LC

– le

ast c

once

rn, N

T –

Nea

r Thr

eate

ned

No

SPE

CIE

SSA

MPL

ES

Mai

n ha

bita

t in

the

park

Alti

tude

ran

ge

in th

e pa

rkE

ndem

ics

Prot

ecte

d

Coc

hlos

tom

atid

ae1

Coc

hlos

tom

a (T

urri

tus)

sp.

HN

HM

: 40

Lim

esto

ne ro

cks (

petro

philo

us)

1718

----

Succ

inei

dae

2O

xylo

ma

(Oxy

lom

a) e

lega

ns (R

isso

, 182

6)H

NH

M: 2

7W

etla

nd85

0--

--L

auri

idae

3La

uria

cyl

indr

acea

(da

Cos

ta, 1

778)

ID: 2

0430

, 204

32, 2

0443

; NH

MW

: 24

Deg

rade

d de

cidu

ous f

ores

t85

5-86

8--

--O

rcul

idae

4O

rcul

a cf

. wag

neri

Stu

rany

, 191

4ID

: 204

37, 2

0440

, 204

41; H

NH

M: 3

3Li

mes

tone

rock

s and

rock

y gr

assl

ands

1814

-226

9W

BL

NT

Pyra

mid

ulid

ae5

Pyra

mid

ula

ceph

alon

ica

(Wes

terlu

nd, 1

898)

ID: 2

0432

Lim

esto

ne ro

cks (

petro

philo

us)

855

WB

L--

6Py

ram

idul

a pu

silla

Gitt

enbe

rger

& B

ank,

19

96ID

: 204

30, 2

0432

, 204

43; N

HM

W: 2

4;

HN

HM

: 28

Lim

esto

ne ro

cks (

petro

philo

us)

855-

868

----

Cho

ndri

nida

e

7C

hond

rina

arc

adic

a (R

einh

ardt

, 188

1)ID

: 204

12, 2

0413

, 204

30 (c

f.), 2

0432

; H

NH

M: 1

5, 3

3, 3

4, 3

9, 4

0; N

HM

W: 2

4R

ocks

(lim

esto

nes,

petro

philo

us)

850-

1858

--LC

Vert

igin

idae

8Tr

unca

telli

na c

laus

tral

is (G

redl

er, 1

856)

HN

HM

: 28

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds86

0--

--9

Trun

cate

llina

cyl

indr

ica

(Fer

ussa

c, 1

807)

ID: 2

0430

, 204

32; N

HM

W: 2

4Li

mes

tone

rock

s and

rock

y gr

assl

ands

855-

868

--LC

10Tr

unca

telli

na ro

thi (

Rei

nhar

dt, 1

916)

ID: 2

0430

, 204

32; N

HM

W: 2

4Li

mes

tone

rock

s and

rock

y gr

assl

ands

855-

868

--LC

Eni

dae

11C

hond

rula

mac

edon

ica

Wag

ner,

1914

ID: 2

0412

(cf.)

, 204

21, 2

0425

, 204

27, 2

0440

, 20

441,

204

42 (c

f.), 2

0443

(cf.)

, 204

44 (c

f.);

HN

HM

: 15,

28,

35,

40;

NH

MW

: 1Li

mes

tone

rock

s and

rock

y gr

assl

ands

850-

1863

BL

LC

12C

hond

rula

mic

rotr

agus

(Ros

smas

sler

, 183

9)

ID: 2

0412

, 204

13, 2

0420

, 204

25,

2042

7,

2043

0, 2

0434

, 204

39, 2

0442

(cf.)

, 204

43,

2044

4 (c

f.); H

NH

M: 1

5, 2

7, 2

8; N

HM

W: 1

, 26

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds85

0-15

65B

LLC

13Ja

min

ia q

uadr

iden

s (M

ülle

r, 17

74)

ID: 2

0413

, 204

32Li

mes

tone

rock

s and

rock

y gr

assl

ands

855

--LC

14M

erdi

gera

obs

cura

(Mül

ler,

1774

)ID

: 204

18, 2

0429

Deg

rade

d de

cidu

ous f

ores

t86

5-13

11--

LC

14Ze

brin

a de

trita

(Mül

ler,

1774

)ID

: 204

12, 2

0419

, 204

20, 2

0425

, 20

427,

20

430,

204

31, 2

0439

, 204

42, 2

0443

, 204

44;

HN

HM

: 15,

27,

28;

NH

MW

: 1Li

mes

tone

rock

s and

rock

y gr

assl

ands

850-

1565

----

Feru

ssac

iidae

16C

ecili

oide

s (C

ecili

oide

s) a

cicu

la

(Mül

ler,

1774

)ID

: 204

30, 2

0443

; HN

HM

: 28

MSS

859-

868

----


475

Tabl

e 2.

Con

tinua

tion.

No

SPE

CIE

SSA

MPL

ES

Mai

n ha

bita

t in

the

park

Alti

tude

ran

ge

in th

e pa

rkE

ndem

ics

Prot

ecte

d

Cla

usili

idae

17Al

inda

bip

licat

a di

stin

cta

(Stu

rany

, 189

4)ID

: 204

12, 2

0413

, 204

21, 2

0424

, 204

30,

2043

1, 2

0432

, 204

34, 2

0442

, 204

43, 2

0444

; H

NH

M: 1

5, 2

8, 4

0; N

HM

W: 1

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds (a

lso

degr

aded

dec

iduo

us fo

rest

s, bu

shes

)85

0-17

36W

BL

--

18M

onte

negr

ina

dofle

ini d

oflei

ni (A

.J. W

agne

r, 19

28)

HN

HM

: 36

Lim

esto

ne ro

cks (

petro

philo

us)

1746

WB

L--

19M

onte

negr

ina

dofle

ini p

resp

aens

is

Nor

dsie

ck, 1

988

ID: 2

0412

; N

HM

W: 1

, 31

Lim

esto

ne ro

cks (

petro

philo

us)

850-

856

LE--

20M

onte

negr

ina

dofle

ini w

agne

ri S

zeke

res,

2006

ID: 2

0421

, 204

36, 2

0440

, 204

41; H

NH

M: 3

3,

37, 3

8, 4

1Li

mes

tone

rock

s (pe

troph

ilous

)17

36-2

204

LE--

21M

onte

negr

ina

hiltr

udae

cos

tula

ta E

rőss

&

Szek

eres

, 200

6ID

: 204

30; H

NH

M: 4

0Li

mes

tone

rock

s (pe

troph

ilous

)86

8-17

18LE

--

22M

onte

negr

ina

hiltr

udae

des

aret

ica

Fehé

r &

Szek

eres

, 201

6ID

: 204

31, 2

0442

, 204

43,

2044

4; H

NH

M:

15, 2

7, 2

8; N

HM

W: 2

4, 2

6Li

mes

tone

rock

s (pe

troph

ilous

)85

0-87

0LE

--

23M

onte

negr

ina

hiltr

udae

fusc

a Fe

her &

Sz

eker

es, 2

006

ID: 2

0440

, 204

41Li

mes

tone

rock

s (pe

troph

ilous

)90

0-18

30LE

--

24M

onte

negr

ina

hiltr

udae

sattm

anni

N

ords

ieck

, 198

8

ID: 2

0424

, 204

25, 2

0426

, 204

27, 2

0428

, 20

429,

204

32, 2

0434

, 204

35, 2

0438

; HN

HM

: 39

; NH

MW

: 11,

16,

29

Lim

esto

ne ro

cks (

petro

philo

us)

850-

1179

LE--

25St

rigi

lode

lima

cons

pers

a (L

. Pfe

iffer

, 184

8)

ID: 2

0430

, 204

31; N

HM

W: 3

0Li

mes

tone

rock

s and

rock

y gr

assl

ands

(als

o de

grad

ed d

ecid

uous

fore

sts,

bush

es)

868-

1040

WB

L--

Pris

tilom

atid

ae26

Vitre

a cf

. bot

teri

i (L.

Pfe

iffer

, 185

3)ID

: 204

21, 2

0430

; HN

HM

: 28;

NH

MW

: 24

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds86

0-17

36--

--27

Vitre

a sp

. ID

: 204

32--

855-

1030

----

Oxy

chili

dae

28M

edite

rran

ea in

opin

ata

(Ulic

ny, 1

887)

HN

HM

: 28

Subt

erra

nean

860

----

29M

edite

rran

ea sp

.ID

: 204

36, 2

0443

Su

bter

rane

an?

859-

2204

----

30M

orlin

a gl

abra

(Ros

smas

sler

, 183

5)ID

: 204

12 (c

f.), 2

0413

(cf.)

, 204

24 (a

ntom

y),

2042

9 (c

f.), 2

0430

(cf.)

, 204

43 (c

f.); H

NH

M:

15 (c

f.), 2

8 (c

f.); N

HM

W: 3

2 (c

f.)

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds (a

lso

degr

aded

dec

iduo

us fo

rest

s, bu

shes

)85

0-10

30--

--

Mila

cida

e

31Ta

ndon

ia m

aced

onic

a (R

ähle

, 197

4)

ID: 2

0421

(ant

omy)

, 204

22 (c

f.)R

ocks

1731

-173

6--

--

32Ta

ndon

ia sp

.ID

: 204

23--

866

----

Vitr

inid

ae33

Olig

olim

ax sp

. ID

: 204

32Li

mes

tone

rock

s and

rock

y gr

assl

ands

855

----

34V

itrin

idae

, af.

Sem

ilim

acel

la sp

. ID

: 204

30--

868

----

476


Tabl

e 2.

Con

tinua

tion.

No

SPE

CIE

SSA

MPL

ES

Mai

n ha

bita

t in

the

park

Alti

tude

ran

ge

in th

e pa

rkE

ndem

ics

Prot

ecte

d

35V

itrin

idae

, af.

Vitr

ina

sp.

ID: 2

0420

, 204

21, 2

0432

, 204

36, 2

0437

, 20

441,

204

43--

855-

2269

----

Lim

acid

ae36

Lehm

anni

a cf

. bru

nner

i (W

agne

r, 19

31)

ID: 2

0417

, 204

28, 2

0429

Roc

ks86

5-13

11B

L--

37Li

mac

us fl

avus

(Lin

naeu

s, 17

58)

ID: 2

0429

Syna

nthr

ope

865

BL

--38

Lim

ax c

f. co

nem

enos

i Böt

tger

, 188

2ID

: 204

28Sy

nant

hrop

e86

5--

--

39Li

max

cf.

corc

yren

sis (

Sim

roth

, 190

5) se

nsu

Räh

le 1

981

ID: 2

0429

Roc

ks86

5W

BL

--

40Li

max

cf.

grae

cus S

imro

th, 1

889

ID: 2

0421

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds17

36B

L--

41Li

max

cf.

max

imus

Lin

naeu

s, 17

58ID

: 204

22D

egra

ded

deci

duou

s for

est

1731

----

Agr

iolim

acid

ae42

Der

ocer

as sp

. (M

ülle

r, 17

74)

ID: 2

0421

--17

36--

--H

elic

odon

tidae

43Li

ndho

lmio

la c

orcy

rens

is (D

esha

yes,

1839

)ID

: 204

15, 2

0416

, 204

19, 2

0421

, 204

25,

2042

7, 2

0430

, 204

34, 2

0435

, 204

38, 2

0440

, 20

441;

HN

HM

: 15;

NH

MW

: 1, 3

0

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds (a

lso

degr

aded

dec

iduo

us fo

rest

s, bu

shes

)86

3-18

30W

BL

LC

Hyg

rom

iidae

44C

andi

dula

rhab

doto

ides

(Wag

ner,

1928

)ID

: 204

12, 2

0421

, 204

25, 2

0427

, 204

30,

2043

4, 2

0440

, 204

42, 2

0443

, 204

44; H

NH

M:

28

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds (a

lso

degr

aded

dec

iduo

us fo

rest

s, bu

shes

)85

3-18

14B

L--

45C

ernu

ella

cf.

virg

ata

(da

Cos

ta, 1

778)

ID: 2

0430

Lim

esto

ne ro

cks a

nd ro

cky

gras

slan

ds (a

lso

degr

aded

dec

iduo

us fo

rest

s, bu

shes

)86

8--

LC

46M

onac

ha c

f. fre

quen

s (M

ouss

on, 1

859)

ID: 2

0421

, 204

33, 2

0438

, 204

41; H

NH

M: 2

7Li

mes

tone

rock

s and

rock

y gr

assl

ands

(als

o de

grad

ed d

ecid

uous

fore

sts,

bush

es, r

uder

al

vege

tatio

n)85

0-18

30--

LC

47Xe

role

nta

obvi

a (M

enke

, 182

8)ID

: 204

12, 2

0415

, 204

16, 2

0419

, 204

33,

2043

6Li

mes

tone

rock

s and

rock

y gr

assl

ands

(als

o de

grad

ed d

ecid

uous

fore

sts,

bush

es)

856-

2204

--LC

Hel

icid

ae

48H

elix

luco

rum

Lin

naeu

s, 17

58

ID: 2

0413

, 204

14, 2

0415

, 204

19,

2042

3,

2042

4, 2

0425

, 204

26, 2

0427

, 204

28, 2

0439

; H

NH

M: 1

5, 2

7Eu

rybi

ont

850-

1565

--LC

49H

elix

phi

libin

ensi

s Ros

smäs

sler

, 183

9 ID

: 204

12, 2

0413

, 204

19, 2

0425

, 204

26,

2043

0, 2

0431

; HN

HM

: 15,

27;

NH

MW

: 1, 2

9Li

mes

tone

rock

s and

rock

y gr

assl

ands

(als

o de

grad

ed d

ecid

uous

fore

sts,

bush

es)

850-

1565

BL

LC

50H

elix

schl

aefli

i Mou

sson

, 185

9ID

: 204

21, 2

0436

, 204

41Li

mes

tone

rock

s and

rock

y gr

assl

ands

866-

2204

WB

LLC

51W

ladi

slaw

ia th

atee

nsis

Sub

ai, 2

012

HN

HM

: 33,

34

Lim

esto

ne ro

cks (

petro

philo

us)

1825

-185

8LE

--N

oSP

EC

IES

LO

CA

LIT

IES

Mai

n ha

bita

t in

the

park

Alti

tude

ran

geE

ndem

ics

Prot

ecte

dL

E-7

BL

-7W

BL

-8

NT-

1L

C-1

4


477

Some species have been found as empty shells only and their determination needs additional studies and anatomical verification. These are Cochlostoma (Turritus) sp., Vitrinidae aff. Semilimacella sp., aff. Vitrina sp. and Oligolimax sp. When juveniles and (or) few specimens only have been found as well when variation of characters is substantial, taxa have been identified at the generic level only (Mediterranea, Vitrea). Therefore, additional spe-cies in the examined area can be expected.

According Sturany & Wagner (1915), the subspecies Orcula wagneri ljubetenensis Sturany, 1914 has a wider diameter than the nominotypical subspecies and has only a single columellar fold. It differs from Orcula schmidtii schmidtii (Küster, 1843) by the fewer number but faster growing whorls and the very narrow umbilicus. These char-acters could vary in different populations and Harl et al. (2014) considered O. schmidtii and O. wagneri as possible synonyms. Our specimens are bigger than O. w. ljubetenensis and resemble O. schmidtii (in some specimens the number of whorls is 9) but, in view of their narrow umbilicus, this might be only a variation of O. w. wagneri.

Such problems exist also with the identification of species of the genus Pyramidula. Despite recent molecular studies (Kirchner et al. 2015, Razkin et al. 2016), it is still not clear where are the morpholog-ical species limits between P. cephalonica and P. pu-silla. Therefore, we identify Pyramidula spp. on con-chological basis only: the population comprising of wider and lower-shelled specimens are identified as P. cephalonica whereas those with higher shell spira and swollen body-whorls are assigned to P. pusilla.

Limax sp. is probably identical with L. corcyren-sis Simroth, 1904 sensu Rähle (1981). It seems, how-ever, our sample from the Lake Prespa area does not fit to the concept of this species as presented by Wiktor (2001) and recognising L. corcyrensis as a synonym of Limax cephalonicus Simroth, 1886. Similar speci-mens to those identified here as “Limax sp.” have been collected from Macedonia and Bulgaria (Dedov, unpublished data). In our opinion, the Limax graecus species-group from the Balkans, southern Italy and Turkey needs an entire taxonomic revision.

Acknowledgement: The present study was completed in the frames of the project task “Biodiversity values of the Prespa Na-tional Park on the basis of selected invertebrate groups” fund-ed by the Association „Friends of the Prespa National Park in Albania“ and the Ministry of Environment, Forests and Water Administration, Directorate of EU Integration and international projects within the project “Trans-boundary Biosphere Reserve Prespa – support to the National Park Prespa in Albania”. The field trips of the NHMW in 2014 and 2015 were funded by the Austrian Science Fund (FWF P 26581-B25).

References

Dhora D. H. & Welter-Schultes F. W. 1996. List of species and atlas of the non-marine molluscs of Albania. Schriften zur Malakozoologie 9: 90–197.

Erőss Z., Fehér Z. & Szekeres M. 2006. New taxa of Montene-grina O. Boettger, 1877 (Gastropoda: Pulmonata: Clausili-idae) from Albania and the neighbouring regions. Annalen des Naturhistorischen Museums in Wien 107 (B): 181–208.

Fehér Z. & Drimmer L. 2004. Mollusks in the Megyeri Collec-tion. Soosiana 31: 21–25.

Fehér Z. & Szekeres M. 2016. Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae). ZooKeys 599: 1–137. doi: 10.3897/zookeys.599.8168

Harl J., Páll-Gergely B., Kirchner S., Sattmann H., Duda M., Kruckenhauser L. & Haring E. 2014. Phylogeography of the land snail genus Orcula (Orculidae, Stylommatophora) with emphasis on the Eastern Alpine taxa: speciation, hybridization and morphological variation. Evolutionary Biology 14: 223.

Hristovski S., Dedov I., Kitanova D., Chobanov D., Hristov G. & Stefanov L. 2014. Biodiversity values of Prespa National Park on the basis of selected invertebrate groups. Final Report, pp. 1–20.

Kirchner S., Harl J., Kruckenhauser L., Duda M., Sattmann H. & Haring E. 2015. Phylogeography and systematics of Pyramidula (Pulmonata: Pyramidulidae) in the eastern Alps: Still a taxonomic challenge. Journal of Molluscan Studies 1–12. doi:10.1093/mollus/eyv047

Neubert E. 2014. Revision of Helix Linnaeus, 1758 in its eastern Mediterranean distribution area, and reassignment of Helix godetiana Kobelt, 1878 to Maltzanella Hesse, 1917 (Gas-tropoda, Pulmonata, Helicidae). Contribution to Natural History 26: 1–200.

Nordsieck H. 2008. Alinda biplicata (Montagu) and Laciniaria plicata (Draparnaud), diversity in comparison, with the description of new subspecies (Gastropoda: Stylomma-tophora: Clausiliidae). Archiv für Molluskenkunde 137 (2): 133–157.

Razkin O., Sonet G., Breugelmans K., José Madeira M., Gómez-Moliner B. J. & Backeljau T. 2016. Species lim-its, interspecific hybridization and phylogeny in the cryptic land snail complex Pyramidula: The power of RADseq data. Molecular Phylogenetics and Evolution 101: 267–278.

Rähle W. 1981 Zur Kenntnis korfiotischer Limax-Arten (Gas-tropoda: Pulmonata). Archiv für Molluskenkunde 111 (1980): 37–41.

Sturany R. & Wagner A. J. 1915. Über schalentragende Land-mollusken aus Albanien und Nachbargebieten. Denk-schriften der kaiserlichen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse 91: 19–138.

Subai P. 2012. Ergänzungen zur Revision der Ariantinae. Schriften zur Malakozoologie 27: 35-46.

Welter-Schultes F. W. 1996. Non-marine molluscs recently col-lected in Albania. Schriften zur Malakozoologie 9: 21–31.

Wiktor A. 2001. The slugs of Greece (Arionidae, Milacidae, Li-macidae, Agriolimacidae – Gastropoda, Stylommatophora). Fauna Graeciae 8: 1–241.

Received: 04.01.2018Accepted: 12.06.2018

Diversity of Terrestrial Gastropods (Molluscsa, Gastropoda ...Nine further spe-cies were typical for...

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Transcript of Diversity of Terrestrial Gastropods (Molluscsa, Gastropoda ...Nine further spe-cies were typical for...