Current Distribution, Status, and Threats of Canada Lynx ... · Current Distribution, Status, and...

John Squires, Rocky Mountain Research Station, Missoula, MT

Current Distribution, Status, and Threats of Canada Lynx in Montana and Wyoming

3

Basis of Assessment - Montana:

a) Lynx Research Program at RMRS initiated in 1998 b) Captured and collared 175 individuals c) Recorded 169,782 GPS and 3043 VHF locations that document lynx movements and resource- use

d) Investigated the following topics regarding lynx in Montana:

4

Basis of Assessment:

• Resource selection Squires, J. R, N. J. DeCesare, J. A. Kolbe, and L. F. Ruggiero. 2008.

Hierarchical den selection of Canada lynx in western Montana. Journal of Wildlife Management 72:1497-1506.

Squires, J. R., N. J. DeCesare, J. A. Kolbe, and L. F. Ruggiero. 2010. Seasonal Resource Selection of Canada Lynx in Managed Forests of the Northern Rocky Mountains. Journal of Wildlife Management 74:1648-1660.

• Prey Selection Squires, J. R. and L. F. Ruggiero. 2007. Winter prey selection of Canada lynx

in northwestern Montana. Journal of Wildlife Management 71:310-315.

• Competition Kolbe, J. A., J. R. Squires, D. H. Pletscher, and R. F. Ruggiero. 2007. The

effect of snowmobile trails on coyote movements within lynx home ranges. Journal of Wildlife Management 71:1409-1418.

5


• Activity Patterns Kolbe, J. A. and J. R. Squires. 2007. Circadian activity patterns of Canada

lynx in western Montana. Journal of Wildlife Management 71:1607-1611.

Olson, L. E., J. R. Squires, N. J. DeCesare, J. A. Kolbe. 2011. Den use and activity patterns in female Canada lynx (Lynx canadensis) in the Northern Rocky Mountains. Northwest Science 85(3):455-462.

• Detection/Monitoring Squires, J. R., K. S. McKelvey, L. F. Ruggiero. 2004. A snow-tracking protocol

used to delineate local lynx, Lynx canadensis, distributions. Canadian Field-Naturalist 118:583-589.

McKelvey, K. S., J. Von Kienast, K. B. Aubry, G. M. Koehler, B. T. Maletzke, J. R. Squires, E. L. Lindquist, S. Loch, M. K. Schwartz. 2006. DNA analysis of hair and scat collected along snow tracks to document the presence of Canada lynx (Lynx canadensis). Wildlife Society Bulletin 34:451-455.

Squires, J. R., L. E. Olson, D. L. Turner, N. J. DeCesare, and J. A. Kolbe. 2012. Estimating detection probability for Canada lynx using snow-track surveys in the Northern Rocky Mountains. Wildlife Biology 18:215-224.

6


• Connectivity

Squires, J. R., Nicholas J. DeCesare, Lucretia E. Olson, Jay A. Kolbe, Mark Hebblewhite, and Sean A. Parks. 2013. Combining resource selection and movement behavior to predict corridors of Canada lynx at their southern range periphery. Biological Conservation 157:187–195.

7

Basis of Assessment - Wyoming:

a) Wyoming Game and Fish Department (WGF), in cooperation with the Shoshone National Forest (SNF), initiated lynx surveys during winter 1995/96 - no lynx detected on SNF - Impetus for WGF to fund additional surveys, trapping, and telemetry in the Wyoming Range b) During 1997-98, WGF searched approximately 2055 km of maintained snowmobile routes and 2400 km of non-maintained trails for lynx tracks in 12 areas (Laurion and Oakleaf 1998) – 6 lynx tracks detected

8


c) During winter 1998-99, three general areas were searched (Laurion and Oakleaf 1999) - 6 tracks located in Wyoming Range.

d) RMRS, in cooperation with WGD, conducted lynx surveys in 2000, 2001, and 2002 - 2000 – Wyoming Range: 5 lynx tracks - kitten and female - 2001 – Wyoming Range: 5 lynx tracks - 2002 – no detections

9


e) Yellowstone Park Lynx survey from 2001 – 2004 (1,143 km ski-based snow tracks, 749 km snowmobile-based survey, 693 km aircraft survey, and 35 hair snare transects - 105-175 stations; Murphy et al. 2006. Distribution of Canada Lynx in Yellowstone National Park. Northwest

Science 80:199-206)

f) DNA confirmed detections of 3 lynx – 1 female, 1 female with male kitten, and 1 male. All detections on east side of Yellowstone Park – east shore Yellowstone Lake.

10


g) Endeavor Wildlife Research Foundation conducted track/DNA surveys between 2004-2005 in southern GYA (Bridger-Teton Nat. Forest including Gros Ventre and Teton Wilderness areas) – searched 4,320 miles and detected (DNA confirmed) 18 lynx tracks. Confirmed presence in Wyoming Range.

h) In 2008-2009, Endeavor Wildlife Research searched 2,854 miles for winter tracks throughout the GYA – documented 6 detections on Togwotee Pass, 2 possible detections in Yellowstone National Park, and 1 possible detection on the Beartooth Plateau.

11

Basis of Assessment – Wyoming (telemetry):

e) From 2006-2007, WGF collared 2 lynx (one male, 1 female) in Wyoming Range – monitored throughout the year with conventional ground and aerial telemetry (1996 – 2001) – (N=219 locations – male, N = 212 - female)

f) In 2000, female died. WGF asked RMRS to replace the collar on male with ARGOS (N = 258). Documented summer exploratory movements of male (1999 –2001) based on conventional and Argos telemetry (Squires and Oakleaf. 2005. Movements of a male Canada lynx crossing the Greater Yellowstone Area, including highways. Northwest Science 79:196-201).

12

Status - Montana

13

Status - Montana

Litter size of lynx in western Montana, 1999 – 2007

Seeley Lake

Purcell Mountains

Combined

Year Litters Kittens Kittens / litter

Litters Kittens Kittens / litter

Litters Kittens Kittens / litter

1999 2 4 2.00 2 4 2.00 2000 4 6 1.50 4 6 1.50 2001 3 8 2.67 3 8 2.67 2002 3 4 1.33 3 4 1.33 2003 5 14 2.80 5 14 2.80 2004 5 15 3.00 5 16 3.20 10 31 3.10 2005 5 12 2.40 6 19 3.17 11 31 2.82 2006 3 5 1.67 3 8 2.67 6 13 2.17 2007 3 7 2.33 8 22 2.75 11 29 2.64

MLE Mean 1 33 75 2.24 22 65 2.95 55 140 2.53 MLE Var 1 0.002 0.084 0.008 95% CI 2.21-2.27 2.67-3.23 2.51-2.55

Litter size of lynx in western Montana, 1999 – 2007

Seeley Lake

Purcell Mountains

Combined

Year

Litters

Kittens

Kittens / litter

Litters

Kittens

Kittens / litter

Litters

Kittens

Kittens / litter

1999

2

4

2.00

2

4

2.00

2000

4

6

1.50

4

6

1.50

2001

3

8

2.67

3

8

2.67

2002

3

4

1.33

3

4

1.33

2003

5

14

2.80

5

14

2.80

2004

5

15

3.00

5

16

3.20

10

31

3.10

2005

5

12

2.40

6

19

3.17

11

31

2.82

2006

3

5

1.67

3

8

2.67

6

13

2.17

2007

3

7

2.33

8

22

2.75

11

29

2.64

MLE Mean 1

33

75

2.24

22

65

2.95

55

140

2.53

MLE Var 1

0.002

0.084

0.008

95% CI

2.21-2.27

2.67-3.23

2.51-2.55

14

Status - Montana Proportion of successful adult Canada lynx

Seeley Lake (N=52 breeding-age females)

Purcell Mountains (N=28)

Combined (N = 80)

Year Females Females w kittens

Prop Females Females w kittens

Prop Females Females w kittens

Prop

1999 4 2 0.50 4 2 0.50 2000 6 4 0.67 6 4 0.67 2001 9 3 0.33 9 3 0.33 2002 6 2 0.33 6 2 0.33 2003 5 3 0.60 5 3 0.60 2004 6 4 0.68 5 5 1.00 11 9 0.82 2005 6 5 0.83 7 7 1.00 13 12 0.92 2006 3 3 1.00 6 4 0.67 9 7 0.78 2007 3 3 1.00 10 7 0.70 13 10 0.77

MLE 1 Mean 0.61 0.83 0.67

MLE Var 1 0.01 0.01 0.02

95% CI 0.42-0.81 0.43-0.98 0.45-0.82

Proportion of successful adult Canada lynx

Seeley Lake

(N=52 breeding-age females)

Purcell Mountains

(N=28)

Combined

(N = 80)

Year

Females

Females w kittens

Prop

Females

Females w kittens

Prop

Females

Females w kittens

Prop

1999

4

2

0.50

4

2

0.50

2000

6

4

0.67

6

4

0.67

2001

9

3

0.33

9

3

0.33

2002

6

2

0.33

6

2

0.33

2003

5

3

0.60

5

3

0.60

2004

6

4

0.68

5

5

1.00

11

9

0.82

2005

6

5

0.83

7

7

1.00

13

12

0.92

2006

3

3

1.00

6

4

0.67

9

7

0.78

2007

3

3

1.00

10

7

0.70

13

10

0.77

MLE 1

Mean

0.61

0.83

0.67

MLE Var 1

0.01

0.01

0.02

95% CI

0.42-0.81

0.43-0.98

0.45-0.82

15

We estimated survival based on 125 lynx monitored monthly from 1999-2007 using a staggered entry design; we documented 2376 lynx-use months during this period.

16

Annual Survival Rate of female lynx on the Seeley Lake(1999-2007) and Purcell (2003-2007) Study Areas including a combined estimate.

Female

Subadult Survival 1 Female

Adult Survival 1

Seeley Mean 0.515 0.747

Variance 0.014 0.003

95% CI 0.283 - 0.746 0.648 - 0.846

Purcells Mean 0.683 0.846


95% CI 0.428 - 0.937 0.721 - 0.970

Combined Mean 0.520 0.753


95% CI 0.322 - 0.718 0.659 - 0.847

Annual Survival Rate of female lynx on the Seeley Lake(1999-2007) and Purcell (2003-2007) Study Areas including a combined estimate.

Female

Subadult Survival 1

Female

Adult Survival 1

Seeley

Mean

0.515

0.747

Variance

0.014

0.003

95% CI

0.283 - 0.746

0.648 - 0.846

Purcells

Mean

0.683

0.846

Variance

0.017

0.004

95% CI

0.428 - 0.937

0.721 - 0.970

Combined

Mean

0.520

0.753

Variance

0.010

0.002

95% CI

0.322 - 0.718

0.659 - 0.847

17

Status - Montana

0200400600800

1000120014001600

0.70

0.80

0.90

1.00

1.10

1.20

1.30

1.40

1.50

1.60

Lambda

# of

repl

icat

es

SeeleyPurcellsCombined

Frequency distribution of λ values from 5000 replicates in which lynx vital rates were chosen from a uniform distribution bounded by their 95% confidence intervals

Chart2

0.70.70.7

0.750.750.75

0.80.80.8

0.850.850.85

0.90.90.9

0.950.950.95

111

1.051.051.05

1.11.11.1

1.151.151.15

1.21.21.2

1.251.251.25

1.31.31.3

1.351.351.35

1.41.41.4

1.451.451.45

1.51.51.5

1.551.551.55

1.61.61.6

1.651.651.65

Seeley

Purcells

Combined

Lambda

# of replicates

0

0

0

11

0

0

213

0

7

608

0

178

1080

16

611

1285

86

1079

995

240

1383

549

456

1002

199

680

560

52

817

154

8

798

25

0

708

1

0

499

0

0

356

0

0

204

0

0

92

0

0

39

0

0

7

0

0

2

0

0

0

0

determ_lambdas

Note: This worksheet shows the λ values from our determinisitc models.

λD= the deterministic lambda from the mean matrix for each population, or the matrix in which vital rates are equal to their MLE means

mean λD_rep= the mean lambda from the 5000 replicate matrices in which vital rates were chosen from uniform distributions bound by the 95% CL's for each vital rate

λ from Replicates comes from the the 5000 replicate matrices; the data are stored in each study area's Deterministic folder in a file called "(studyarea)_randomlambdas"

Study Area:SeeleyPurcellsCombinedFrequency Dist. of λ from 5000 replicatesCumulative Dist. of λ from 5000 replicates

λD:0.9231.2230.975

mean λD_rep:0.9251.1680.973λSeeleyPurcellsCombinedλSeeleyPurcellsCombined

std_dev of mean:0.0750.1170.0700.700000.70000

95% UCL:0.9271.1710.0020.7511000.750.2200

95% LCL:0.9231.1650.0000.80213070.804.4800.14

min_λ:0.7200.8530.7800.8560801780.8516.6403.7

max_λ:1.1921.5941.2070.901080166110.9038.240.3215.92

median_λ:0.9241.1630.9720.9512858610790.9563.942.0437.5

λ values from 5000 Replicates:1.21911.01161.0099524013831.0083.846.8465.16

11.07221.14581.08531.0554945610021.0594.8215.9685.2

20.948141.09270.985161.101996805601.1098.829.5696.4

30.905821.23831.01091.15528171541.1599.8445.999.48

40.910631.24121.02461.208798251.2010061.8699.98

50.899351.13891.05571.25070811.2510076.02100

61.11041.16860.982771.30049901.3010086100

70.987181.01840.910251.35035601.3510093.12100

80.972351.12750.862671.40020401.4010097.2100

90.88011.26290.874111.4509201.4510099.04100

100.926211.2271.08111.5003901.5010099.82100

110.882541.20321.03721.550701.5510099.96100

120.927591.19741.02931.600201.60100100100

130.887121.14370.970851.650001.65100100100

140.942971.37970.98392

150.849441.12110.87596

160.909111.17831.12990.70000

170.882181.09430.845970.751100

180.925981.31380.970330.8022407

191.02261.08471.01770.858320185

201.01991.27650.987390.90191216796

210.93561.11161.0720.9531971021875

220.933651.27181.03061.0041923423258

230.863850.994861.00731.0547417984260

240.787711.06730.91121.10494014784820

250.982741.13820.995951.15499222954974

260.908821.33251.07291.20500030934999

270.841681.06990.991031.25500038015000

281.0261.09311.02331.30500043005000

290.862261.31690.995821.35500046565000

300.885011.02450.954191.40500048605000

310.903151.15230.974961.45500049525000

320.989661.1140.971521.50500049915000

330.877481.34450.947221.55500049985000

340.904521.13190.959681.60500050005000

351.00691.09591.0041.65500050005000

360.804261.04741.1463

370.867181.30720.94618

380.810271.14570.83689

390.803571.02921.0378

400.872421.30.93367

410.963851.06130.82504

420.830171.34610.9045

430.971771.41190.9782

440.877161.37641.0758

450.965171.27131.0635

460.883231.09860.96167

470.913971.12141.0812

480.904381.14930.79761

490.839711.12080.98884

500.831651.0540.95876

510.844681.02381.0095

520.934531.08590.99387

530.952591.07440.87838

541.00611.18910.92305

550.950520.917140.91573

560.831331.09790.85254

570.795131.03811.0164

580.894721.02260.97779

590.938521.35230.98565

600.981631.06160.93798

610.925151.30940.9799

620.935151.08860.99776

630.860921.08671.0075

640.930511.0740.82092

650.922721.29070.91295

660.884761.23450.92155

670.939391.14190.97168

680.920251.26250.93703

691.00191.12290.9161

700.841161.37020.94362

710.956841.1511.0146

720.811221.23840.85968

730.921951.30611.055

740.84551.21140.91431

750.825811.18680.98178

761.02931.29410.93738

770.943741.15340.95626

780.956411.10990.94583

791.09661.18560.97578

801.00681.05541.078

810.945341.13370.83511

820.848941.06261.117

830.98031.19281.0667

840.939831.21890.88514

850.924791.13790.83901

860.853741.02630.98255

870.815381.26540.87745

880.850891.43690.91552

890.869221.14780.99491

900.831551.20691.0772

910.935741.27521.0268

920.865471.25451.0193

930.929371.08850.91067

940.986741.17020.87507

950.982321.14031.0074

960.907461.21820.97069

970.852831.11821.0307

980.772651.41780.96541

990.94031.19841.0281

1000.977751.0760.99449

1010.789141.20841.0468

1020.977091.11270.97941

1030.949141.11751.0431

1041.02891.08321.0935

1051.14471.33180.90652

1060.965961.00281.0087

1070.919241.2881.1146

1081.00091.5140.97512

1091.03391.29581.0498

1100.974341.08930.89624

1110.93861.00310.90952

1120.874981.01111.0053

1130.933921.11760.91446

1140.905091.28370.88739

1150.785410.985211.0133

1160.800111.10331.0845

1170.989711.4010.91996

1181.0031.12330.92016

1190.818851.11990.95115

1200.946341.36870.91711

1210.908131.221.0407

1220.86831.28890.9972

1231.05391.15030.9386

1240.866571.3040.86497

1250.977621.26151.0324

1260.923761.21290.92383

1270.790171.26181.0696

1280.855611.19540.83798

1291.09081.22750.96169

1300.869581.36540.9311

1310.911961.11980.93033

1320.905471.25280.90556

1330.839441.38430.9171

1340.865011.15711.0485

1350.884661.2610.97832

1360.849951.30110.99421

1371.00331.36221.0202

1380.895251.25560.94605

1390.969811.29030.97906

1400.927011.18031.055

1410.919651.11131.1016

1421.00631.1120.97294

1430.918211.20171.1256

1440.963881.30441.0468

1450.905151.30430.84398

1460.990871.13911.015

1470.787371.1151.0006

1480.812541.27671.0354

1490.967141.11021.0683

1500.945451.38620.93908

1510.874441.1980.98496

1520.852341.17370.95106

1530.920061.17390.90819

1540.944921.20820.91947

1550.937751.23470.94108

1560.887751.09871.0448

1570.831691.26181.0755

1580.954820.932490.92493

1590.918121.17360.86793

1600.993571.05370.93094

1610.858151.110.96746

1620.975361.33541.0113

1630.962151.33750.9161

1640.99061.31891.0566

1651.01051.40321.0287

1661.00891.10991.0511

1670.882011.08710.90267

1680.921431.13870.89239

1690.924421.19170.92883

1700.801710.980360.983

1710.937291.25850.86821

1720.973951.29940.93049

1730.992181.33351.0063

1740.865351.36480.93471

1750.884521.12410.99329

1760.856381.10840.94299

1770.92190.959260.98652

1780.896581.29890.92373

1790.9831.41171.1038

1800.958231.22060.98936

1810.961981.14421.0913

1821.00871.07780.81181

1831.02271.11440.87117

1840.819941.04581.0254

1850.998431.23530.97976

1860.933881.03341.1629

1871.07981.14790.96356

1880.775831.32380.94831

1890.911051.24351.0218

1900.947581.15561.0281

1910.811821.15620.94609

1920.957791.33061.0961

1930.796321.22981.1324

1940.868081.24770.8981

1951.0131.21750.84481

1960.903651.10621.0038

1970.974691.09591.0128

1980.792781.2160.99659

1990.887191.14380.97214

2000.779111.09981.0515

2011.00441.26840.89833

2020.846531.1380.94539

2031.03321.26031.1056

2040.821441.2331.0299

2050.941221.11051.0716

2060.878061.27220.88033

2070.891291.19740.97583

2080.804511.22780.98685

2090.94941.07930.91425

2100.956571.09790.97221

2110.988771.07251.0172

2120.971691.1271.0137

2130.83271.17250.96876

2140.991911.15341.0116

2150.995410.981950.89298

2161.00851.24460.91787

2170.943791.0260.96212

2180.877941.0580.99448

2190.916491.09531.1326

2200.875091.00331.1523

2210.898821.2431.0602

2220.905791.17420.95258

2230.91681.2750.95241

2240.958871.11131.0728

2250.944110.984580.95289

2260.87031.24260.90668

2270.905841.07761.0439

2280.938341.02720.95157

2290.926681.24531.0161

2301.00441.11941.0463

2310.916761.19361.0072

2321.05751.06170.86868

2330.851.36310.92659

2341.09021.34281.0296

2350.822321.02141.1178

2360.954171.2980.99558

2370.774011.37411.0038

2380.893471.37910.94602

2390.834321.13610.9285

2400.828991.21230.91297

2410.835211.2690.90743

2421.09551.03160.88018

2430.974481.26440.99

2440.848471.05170.93035

2450.956261.03090.99334

2461.03961.03271.0441

2470.990670.98131.0205

2480.857731.38670.99236

2491.0241.13891.0305

2500.95051.22731.0116

2510.918031.0850.95817

2520.869381.03320.98098

2530.880471.32651.0329

2540.83481.26050.90459

2550.992811.17391.071

2560.905491.10240.98019

2570.902211.28690.87163

2580.97311.05181.0137

2591.01591.19931.0403

2600.959231.06810.92144

2610.959331.1520.98653

2620.8681.05930.91701

2630.898231.12791.0162

2640.945221.11150.92401

2650.998071.1420.99881

2660.908611.08450.84298

2671.02871.01590.96118

2681.01931.20150.86678

2690.936311.29410.87872

2700.98931.24120.8669

2711.08631.34390.97072

2720.874421.05330.90332

2730.9341.19590.88756

2740.941581.1290.86591

2751.02931.10391.0375

2761.02830.987540.91478

2770.892871.05790.99883

2781.02191.15980.92437

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Current Distribution, Status, and Threats of Canada Lynx ... · Current Distribution, Status, and...

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