Comparisons of growth and early yields of potato … · Comparisons of growth and early ... harvest...

16
J. agric. Sci., Camb. (1984), 103, 443-458 443 With 1 text-figure Printed in Great Britain Comparisons of growth and early yields of potato varieties of contrasting maturity classification at three sites BY R. L. GRIFFITH, E. J. ALLEN*, SUSAN A. O'BRIEN* AND P. J. O'BRIEN* Department of Agriculture (Crop Husbandry), University College of Wales, Aberystwyth, Dyfed {Received 17 April 1984) SUMMARY Six experiments which compared a range of contrasting varieties are reported. Two experiments were at a very early, coastal site in N. Pembrokeshire (Trefin), two at a site in South Pembrokeshire (Trefloyne) and two at a coastal site in N. Norfolk (Bran- caster). No variety was in every experiment but several were in the majority of experiments. In some experiments sprout growth was manipulated by control of storage temperatures. A variety classified as early or second early produced the highest yields in all experiments but the yields of the best maincrop varieties were high and substantially better than many early varieties. The highest-yielding variety at the first harvest maintained this advantage throughout June and July. At two sites (Trefin and Brancaster) the highest-yielding variety was consistent from year to year but overall the ranking order of varieties for yields was inconsistent, e.g. Arran Comet was the highest-yielding variety at Trefin in both years and one of the lowest yielding at Brancaster. The results suggest that it is unwise to attempt to rank varieties for early yields by an overall score. The maturity classification of varieties is not necessarily an accurate guide to the utility of a variety. A consideration of the physiological age of the seed and growing conditions provided some explanation of these effects and their implication for breeding are discussed. INTRODUCTION out-yielded maincrops only for a short period at the beginning of harvesting. Moreover, several authors In the United Kingdom potato varieties are have reported that the growth pattern of varieties, classified into three maturity groups, early, second especially many classified as earlies (O'Brien et al. early and maincrop which broadly indicate the 1983; O'Brien & Allen, 1981) can be modified by appropriate time of harvesting. The classification management of the seed tuber. The low yields of is not meant to be rigid but early varieties are early varieties later in the season may be primarily generally associated with lower yields later in the determined by the use of old seed of certain varie- season than second earlies and maincrops and to ties, for O'Brien et al. (1983) have shown that in obtain the highest yields over a long period several some early varieties old seed leads to smaller leaf changes in variety are assumed to be necessary. areas which senesce earlier than younger seed, Few direct comparisons of contrasting varieties thereby reducing yields later in the season, have been reported and some available data suggest Such results suggest that high yields early in the that differences between varieties are smaller than season may be achieved without prejudice to later hitherto assumed. Gunasena & Harris (1971) found yields and this possibility has considerable impli- that Craigs Royal (second early) produced similar cations for the breeding and testing of new varie- high final yields (80 t/ha) to Pentland Dell (main- ties as well as the most effective use of existing crop) and Allen (1977) reported that early varieties varieties. This paper presents comparisons of con- trasting potato varieties grown at three different * Present address: Department of Applied Biology, sites and harvested during the first 2-3 months of University of Cambridge. the season (May-July). In some experiments sprout at http:/www.cambridge.org/core/terms. http://dx.doi.org/10.1017/S0021859600047420 Downloaded from http:/www.cambridge.org/core. Faculty of Classics, University of Cambridge, on 24 Nov 2016 at 15:48:51, subject to the Cambridge Core terms of use, available

Transcript of Comparisons of growth and early yields of potato … · Comparisons of growth and early ... harvest...

J agric Sci Camb (1984) 103 443-458 4 4 3With 1 text-figurePrinted in Great Britain

Comparisons of growth and earlyyields of potato varieties of contrasting maturity

classification at three sites

BY R L GRIFFITH E J ALLEN SUSAN A OBRIEN AND P J OBRIENDepartment of Agriculture (Crop Husbandry) University College of Wales

Aberystwyth Dyfed

Received 17 April 1984)

SUMMARYSix experiments which compared a range of contrasting varieties are reported Two

experiments were at a very early coastal site in N Pembrokeshire (Trefin) two at a sitein South Pembrokeshire (Trefloyne) and two at a coastal site in N Norfolk (Bran-caster) No variety was in every experiment but several were in the majority ofexperiments In some experiments sprout growth was manipulated by control ofstorage temperatures A variety classified as early or second early produced the highestyields in all experiments but the yields of the best maincrop varieties were high andsubstantially better than many early varieties The highest-yielding variety at the firstharvest maintained this advantage throughout June and July At two sites (Trefin andBrancaster) the highest-yielding variety was consistent from year to year but overallthe ranking order of varieties for yields was inconsistent eg Arran Comet was thehighest-yielding variety at Trefin in both years and one of the lowest yielding atBrancaster

The results suggest that it is unwise to attempt to rank varieties for early yields byan overall score The maturity classification of varieties is not necessarily an accurateguide to the utility of a variety A consideration of the physiological age of the seed andgrowing conditions provided some explanation of these effects and their implication forbreeding are discussed

INTRODUCTION out-yielded maincrops only for a short period at thebeginning of harvesting Moreover several authors

In the United Kingdom potato varieties are have reported that the growth pattern of varietiesclassified into three maturity groups early second especially many classified as earlies (OBrien et alearly and maincrop which broadly indicate the 1983 OBrien amp Allen 1981) can be modified byappropriate time of harvesting The classification management of the seed tuber The low yields ofis not meant to be rigid but early varieties are early varieties later in the season may be primarilygenerally associated with lower yields later in the determined by the use of old seed of certain varie-season than second earlies and maincrops and to ties for OBrien et al (1983) have shown that inobtain the highest yields over a long period several some early varieties old seed leads to smaller leafchanges in variety are assumed to be necessary areas which senesce earlier than younger seedFew direct comparisons of contrasting varieties thereby reducing yields later in the seasonhave been reported and some available data suggest Such results suggest that high yields early in thethat differences between varieties are smaller than season may be achieved without prejudice to laterhitherto assumed Gunasena amp Harris (1971) found yields and this possibility has considerable impli-that Craigs Royal (second early) produced similar cations for the breeding and testing of new varie-high final yields (80 tha) to Pentland Dell (main- ties as well as the most effective use of existingcrop) and Allen (1977) reported that early varieties varieties This paper presents comparisons of con-

trasting potato varieties grown at three different Present address Department of Applied Biology sites and harvested during the first 2-3 months of

University of Cambridge the season (May-July) In some experiments sprout

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444 R L GRIFFITH AND OTHERS

Table 1 Details of varieties used in Expls 1-6

Variety

Home GuardArran CometMaria BardPentland CrownStormont EnterpriseDesireePrimuraPentland JavelinF46USpuntaRenovaJuliverDragaAmincaWiljaEstimaUlster SceptreRed Craigs RoyalMaria PiperFoxtonRecordVanessa

Origin

BritishBritiahBritishBritishBritishDutchDutchBritiahBritiahDutchDutchDutchDutchDutchDutchDutchBritishBritishBritishBritishDutchDutch

Maturityclassification

EarlyEarlyEarlyMaincropMaincropMaincropEarlyEarlyEarlySecond earlySecond earlyMaincropMaincropEarlySecond earlySecond earlyEarlySecond earlyMaincropMaincropMaincropEarly

growth of varieties was manipulated by control ofstorage temperature so as to achieve the requiredphysiological age for early harvests As the datacovered five seasons the influence of site andseason in relation to the testing of varieties couldbe assessed

THE EXPERIMENTS

Two experiments were carried out on the farmof Mr I Davies Park Court Trefin SW Dyfed onlight gravelly soil of the Arfon Series (Rudeforth ampBradley 1972) Both experiments used 66 cm rowsA fertilizer dressing of 166 kg N 107 kg P and138 kg Kha was applied by hand over the ridgesprior to planting Sampling grading and recordingtechniques were essentially as reported by Allen(1977) Details of the varieties grown are given inTable 1

Experiment 1 1976-7Treatments were five varieties Home Guard

Arran Comet Maris Bard Pentland Crown andStormont Enterprise replicated four times in arandomized-block design Plots were four rowswide and 6-1 m long Seed tubers of 55-65 65-7575-85 and 85-95 g were used for the four replicatesand spaced 30 cm apart in the row Numbers ofplants and stems were recorded on two rows on8 and 22 April and yields estimated from ten plantsper plot on 31 May 10 June and 1 July

The seed of Arran Comet Pentland Crown and

Stormont Enterprise was once-grown from mid-Wales and weighed out at University College ofWales in late January when Arran Comet hadconsiderable sprout growth The seed was placedapical end uppermost in trays and stored at 12 degCin a glasshouse with supplementary heating untilplanting The seed of the other varieties was selectedfrom the farmers own Scottish seed in his glass-house on 11 February After selection the seed wasplaced apical end uppermost in trays and left inthe farm glasshouse until planting Sprout measure-ments were made on ten tubers per seed size of eachvariety on 12 March and the experiment plantedby hand on 20 March

Experiment 2 1977-8There were six treatments comprising Home

Guard Arran Comet Maris Bard a once-grownstock (OG) and a certified stock (CC) PentlandCrown and Desiree replicated four times in arandomized-block design The once-grown stock ofMaris Bard was the progeny of the certified seedused in Expt 1 and had been multiplied in a fieldof early potatoes It had been defoliated and har-vested early (early July and mid-July respectively)and consequently had an early end to dormancywhich resulted in very long sprouts (advancedphysiological age) by planting The seed of PentlandCrown and Desiree was once-grown in mid-Waleswhile for the remaining three varieties the seed wasfrom the purchased certified stock of the farmerThe seed from Trefin was weighed out in the farmglasshouse on 22 December and only Maris Bard(certified stock) had no sprout growth Tubers wereplaced in trays as in Expt 1 and remained in thefarm store until planting Seed of Desiree and Pent-land Crown were weighed out at University Collegeof Wales on 13 January when there was some sproutgrowth Tubers were placed in trays as in Expt 1and stored at 12 degC until planting Sprout measure-ments were taken on 9 February on ten tubers ofeach size on all varieties and the experimentplanted on 3 March Plots were four rows wide and5-3 m long Seed tubers of 70-80 80-90 90-100and 110-115 g were used for the four replicates andspaced 27 cm apart in the row Numbers of plantsand stems were counted on 30 March 7 April and1 May using 10-6 m of row Growth analyses usingtwo adjacent plants per plot were carried out on4 and 18 May and yields estimated from ten plantsper plot on 30 May 7 and 16 June

Two experiments were carried out at TrefloyneField Station on Old Red Sandstone soils of thePembroke Series (Soil Survey Record No 24 1974)Both experiments used 71 cm rows and receivedfertilizer dressings of 154 kg N 66 kg P and 160 kgKha applied over the opened ridges prior toplanting

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Growth of contrasting potato varieties 445

Experiment 3 1978-9Treatments were nine varieties Primura Home

Guard Arran Comet Pentland Javelin F4611(earlies) Spunta Renova (second earlies) Juliverand Draga (maincrops) replicated three times in arandomized-block design Plots were three rowswide and 11-25 m long Seed tubers of 40-50 50-60and 60-70 g were used for the three replicates andspaced 25 cm apart in the row Number of plantsand above-ground stems were recorded on 18 Mayusing one complete row Growth analysis (twoplants per plot) and estimates of yields (ten plantsper plot) were carried out on 21 June 3 16 and31 July

All seed was once-grown in mid-Wales Seedwas selected before the onset of sprout growth andplaced in trays as in Expts 1 and 2 The seed wasstored in a frost-free glasshouse until planting(average daily temperature 84 degC) Sprout measure-ments were taken on 8 March using ten seed tubersof each size and the experiment planted by handon 19 April

Experiment 4 1979-80In this experiment 11 varieties were compared

Aminca and Maris Bard (earlies) and Wilja (secondearly) were added to those in Expt 3 and HomeGuard deleted There were three replicates plotdimensions and seed size were as in Expt 3Numbers of plants and above-ground stems werenot recorded in this experiment Growth analysisand estimates of yield were carried out as in Expt 3on 18 27 June 8 and 28 July

Seed production treatment selection and storagewere as in Expt 3 The average daily temperatureof the glasshouse was 9-9 degC Sprout measurementswere taken on 3 March using ten tubers of eachseed size and the experiment planted by hand on21 March

Two experiments were carried out at DeepdaleFarms Limited Brancaster Norfolk on light sandysoils of the Wick Series Rows were 76 cm apartand tubers spaced 20 cm apart Both sites werestone-separated prior to planting

Experiment 5 1979-80Nine varieties Home Guard Arran Comet Maris

Bard (earlies) Red Craigs Royal Wilja Estima(second earlies) Maris Piper Pentland Squire andDesiree (maincrop) were compared in three random-ized blocks Seed of 40-55 g was used for all repli-cates Plots were four rows wide and 5 m longPlant and stem emergence were recorded every3-4 days using one whole row per plot Yields wereestimated from 14 plants per plot on 24 June 8 and22 July

All seed was once-grown in mid or west Wales

Table 2 Number of day-degrees gt4C experiencedby seed from onset of sprouting (3 mmsprout) toplant-ing in three varieties

Experiment

Variety

Home GuardArran CometMaris Bard

(once grown)Maris Bard(certified)

1 5 6

640 390 890 mdash 270 mdash630 340 880 760 300 mdashmdash 740 mdash mdash mdash mdash

330 80 mdash 430 230 530

and selected in late December when all seed hadbroken dormancy The first three varieties werestored at 6 degC the next three at 8 degC and the threemaincrop varieties at 12 degC Sprout measurementswere made on 30 tubers per variety on 29 Februaryand the seed moved to Norfolk The experimentwas planted by hand on 21 March A liquid fertilizerdressing of 150 kg N 86 kg P 167 kg K and 30 kgMgha was applied at planting Four applicationsof 25 mm of water were made by rain-gun on 1422 May and 4 18 June

Experiment 6 1980-1Seven varieties Maris Bard Ulster Sceptre

Vanessa (earlies) Wilja Estima (second earlies)Record and Foxton (maincrop) were compared inthree randomized blocks Seed of 70-80 80-90 and90-100 g was used for the three replicates plotswere the same size as in Expt 5 Emergence was notrecorded and yields were estimated from 14 plantsper plot on 8 22 June and 6 July

Seed of all varieties was stored at 6 degC until mid-February when the two maincrop varieties weretransferred to 12 degC Sprout measurements weremade on ten tubers per seed size of each variety on3 March and the experiment planted by hand on5-6 March

A liquid fertilizer dressing of 145 kg N 61 kg Pand 119 kg Kha was applied at planting Threeapplications of 25 mm of water were made by rain-gun on 7 22 June and 7 July In all experimentsthe control of weeds was satisfactory and theapplication of fungicides and aphicides ensuredthat both blight (Phytophthora infestans) and virusdiseases were controlled

In all experiments the yields per hectare wereobtained by harvesting the area occupied by thestated number of plants irrespective of any missingplants From growth analysis samples on someoccasions yields are presented per plant and careshould be exercised if these values are converted toyield per hectare as the sample area was small andinappropriate for such conversion

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446 R L GRIFFITH AND OTHERS

(a) Aberporth

40

30

20

10

0

50

U 40 H

A

pound 30MU

bullvI 20O

I o

I

1977

-10

50

40 -

30-

20-

10-

0 -

-10

March April

(b) Trefloyne

1979

April

1980

May

(c) Hemsby

V

March April

40-

30-

20

i o H

o

50

40-

30-

20

10

OH

1978

-10J r

50-

40-

30-

20

- 1 0 -

March April

1980

March April

1981

March April

Fig 1 For description see opposite

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60 -i

40-

20

140 n

120 -

100 -

BO-

S 60

40-

20-

lOO-i

80-

60-

40-

20-

0

Growth of contrasting potato varieties(d) Aberporth

1978

447

1977

(e) Trefloyne1979 1980

1980Hemsby

1981

March April May June March April May June

Fig 1 Weekly number of day-degrees gt 4 degC (a-c) and total monthly rainfall (d-f) in 2 years at Aber-porth (north Pembrokeshire) Trefloyne (south Pembrokeshire) and Hemsby (Norfolk) T Time ofplanting v time of 50 plant emergence

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

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Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

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Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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444 R L GRIFFITH AND OTHERS

Table 1 Details of varieties used in Expls 1-6

Variety

Home GuardArran CometMaria BardPentland CrownStormont EnterpriseDesireePrimuraPentland JavelinF46USpuntaRenovaJuliverDragaAmincaWiljaEstimaUlster SceptreRed Craigs RoyalMaria PiperFoxtonRecordVanessa

Origin

BritishBritiahBritishBritishBritishDutchDutchBritiahBritiahDutchDutchDutchDutchDutchDutchDutchBritishBritishBritishBritishDutchDutch

Maturityclassification

EarlyEarlyEarlyMaincropMaincropMaincropEarlyEarlyEarlySecond earlySecond earlyMaincropMaincropEarlySecond earlySecond earlyEarlySecond earlyMaincropMaincropMaincropEarly

growth of varieties was manipulated by control ofstorage temperature so as to achieve the requiredphysiological age for early harvests As the datacovered five seasons the influence of site andseason in relation to the testing of varieties couldbe assessed

THE EXPERIMENTS

Two experiments were carried out on the farmof Mr I Davies Park Court Trefin SW Dyfed onlight gravelly soil of the Arfon Series (Rudeforth ampBradley 1972) Both experiments used 66 cm rowsA fertilizer dressing of 166 kg N 107 kg P and138 kg Kha was applied by hand over the ridgesprior to planting Sampling grading and recordingtechniques were essentially as reported by Allen(1977) Details of the varieties grown are given inTable 1

Experiment 1 1976-7Treatments were five varieties Home Guard

Arran Comet Maris Bard Pentland Crown andStormont Enterprise replicated four times in arandomized-block design Plots were four rowswide and 6-1 m long Seed tubers of 55-65 65-7575-85 and 85-95 g were used for the four replicatesand spaced 30 cm apart in the row Numbers ofplants and stems were recorded on two rows on8 and 22 April and yields estimated from ten plantsper plot on 31 May 10 June and 1 July

The seed of Arran Comet Pentland Crown and

Stormont Enterprise was once-grown from mid-Wales and weighed out at University College ofWales in late January when Arran Comet hadconsiderable sprout growth The seed was placedapical end uppermost in trays and stored at 12 degCin a glasshouse with supplementary heating untilplanting The seed of the other varieties was selectedfrom the farmers own Scottish seed in his glass-house on 11 February After selection the seed wasplaced apical end uppermost in trays and left inthe farm glasshouse until planting Sprout measure-ments were made on ten tubers per seed size of eachvariety on 12 March and the experiment plantedby hand on 20 March

Experiment 2 1977-8There were six treatments comprising Home

Guard Arran Comet Maris Bard a once-grownstock (OG) and a certified stock (CC) PentlandCrown and Desiree replicated four times in arandomized-block design The once-grown stock ofMaris Bard was the progeny of the certified seedused in Expt 1 and had been multiplied in a fieldof early potatoes It had been defoliated and har-vested early (early July and mid-July respectively)and consequently had an early end to dormancywhich resulted in very long sprouts (advancedphysiological age) by planting The seed of PentlandCrown and Desiree was once-grown in mid-Waleswhile for the remaining three varieties the seed wasfrom the purchased certified stock of the farmerThe seed from Trefin was weighed out in the farmglasshouse on 22 December and only Maris Bard(certified stock) had no sprout growth Tubers wereplaced in trays as in Expt 1 and remained in thefarm store until planting Seed of Desiree and Pent-land Crown were weighed out at University Collegeof Wales on 13 January when there was some sproutgrowth Tubers were placed in trays as in Expt 1and stored at 12 degC until planting Sprout measure-ments were taken on 9 February on ten tubers ofeach size on all varieties and the experimentplanted on 3 March Plots were four rows wide and5-3 m long Seed tubers of 70-80 80-90 90-100and 110-115 g were used for the four replicates andspaced 27 cm apart in the row Numbers of plantsand stems were counted on 30 March 7 April and1 May using 10-6 m of row Growth analyses usingtwo adjacent plants per plot were carried out on4 and 18 May and yields estimated from ten plantsper plot on 30 May 7 and 16 June

Two experiments were carried out at TrefloyneField Station on Old Red Sandstone soils of thePembroke Series (Soil Survey Record No 24 1974)Both experiments used 71 cm rows and receivedfertilizer dressings of 154 kg N 66 kg P and 160 kgKha applied over the opened ridges prior toplanting

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Growth of contrasting potato varieties 445

Experiment 3 1978-9Treatments were nine varieties Primura Home

Guard Arran Comet Pentland Javelin F4611(earlies) Spunta Renova (second earlies) Juliverand Draga (maincrops) replicated three times in arandomized-block design Plots were three rowswide and 11-25 m long Seed tubers of 40-50 50-60and 60-70 g were used for the three replicates andspaced 25 cm apart in the row Number of plantsand above-ground stems were recorded on 18 Mayusing one complete row Growth analysis (twoplants per plot) and estimates of yields (ten plantsper plot) were carried out on 21 June 3 16 and31 July

All seed was once-grown in mid-Wales Seedwas selected before the onset of sprout growth andplaced in trays as in Expts 1 and 2 The seed wasstored in a frost-free glasshouse until planting(average daily temperature 84 degC) Sprout measure-ments were taken on 8 March using ten seed tubersof each size and the experiment planted by handon 19 April

Experiment 4 1979-80In this experiment 11 varieties were compared

Aminca and Maris Bard (earlies) and Wilja (secondearly) were added to those in Expt 3 and HomeGuard deleted There were three replicates plotdimensions and seed size were as in Expt 3Numbers of plants and above-ground stems werenot recorded in this experiment Growth analysisand estimates of yield were carried out as in Expt 3on 18 27 June 8 and 28 July

Seed production treatment selection and storagewere as in Expt 3 The average daily temperatureof the glasshouse was 9-9 degC Sprout measurementswere taken on 3 March using ten tubers of eachseed size and the experiment planted by hand on21 March

Two experiments were carried out at DeepdaleFarms Limited Brancaster Norfolk on light sandysoils of the Wick Series Rows were 76 cm apartand tubers spaced 20 cm apart Both sites werestone-separated prior to planting

Experiment 5 1979-80Nine varieties Home Guard Arran Comet Maris

Bard (earlies) Red Craigs Royal Wilja Estima(second earlies) Maris Piper Pentland Squire andDesiree (maincrop) were compared in three random-ized blocks Seed of 40-55 g was used for all repli-cates Plots were four rows wide and 5 m longPlant and stem emergence were recorded every3-4 days using one whole row per plot Yields wereestimated from 14 plants per plot on 24 June 8 and22 July

All seed was once-grown in mid or west Wales

Table 2 Number of day-degrees gt4C experiencedby seed from onset of sprouting (3 mmsprout) toplant-ing in three varieties

Experiment

Variety

Home GuardArran CometMaris Bard

(once grown)Maris Bard(certified)

1 5 6

640 390 890 mdash 270 mdash630 340 880 760 300 mdashmdash 740 mdash mdash mdash mdash

330 80 mdash 430 230 530

and selected in late December when all seed hadbroken dormancy The first three varieties werestored at 6 degC the next three at 8 degC and the threemaincrop varieties at 12 degC Sprout measurementswere made on 30 tubers per variety on 29 Februaryand the seed moved to Norfolk The experimentwas planted by hand on 21 March A liquid fertilizerdressing of 150 kg N 86 kg P 167 kg K and 30 kgMgha was applied at planting Four applicationsof 25 mm of water were made by rain-gun on 1422 May and 4 18 June

Experiment 6 1980-1Seven varieties Maris Bard Ulster Sceptre

Vanessa (earlies) Wilja Estima (second earlies)Record and Foxton (maincrop) were compared inthree randomized blocks Seed of 70-80 80-90 and90-100 g was used for the three replicates plotswere the same size as in Expt 5 Emergence was notrecorded and yields were estimated from 14 plantsper plot on 8 22 June and 6 July

Seed of all varieties was stored at 6 degC until mid-February when the two maincrop varieties weretransferred to 12 degC Sprout measurements weremade on ten tubers per seed size of each variety on3 March and the experiment planted by hand on5-6 March

A liquid fertilizer dressing of 145 kg N 61 kg Pand 119 kg Kha was applied at planting Threeapplications of 25 mm of water were made by rain-gun on 7 22 June and 7 July In all experimentsthe control of weeds was satisfactory and theapplication of fungicides and aphicides ensuredthat both blight (Phytophthora infestans) and virusdiseases were controlled

In all experiments the yields per hectare wereobtained by harvesting the area occupied by thestated number of plants irrespective of any missingplants From growth analysis samples on someoccasions yields are presented per plant and careshould be exercised if these values are converted toyield per hectare as the sample area was small andinappropriate for such conversion

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446 R L GRIFFITH AND OTHERS

(a) Aberporth

40

30

20

10

0

50

U 40 H

A

pound 30MU

bullvI 20O

I o

I

1977

-10

50

40 -

30-

20-

10-

0 -

-10

March April

(b) Trefloyne

1979

April

1980

May

(c) Hemsby

V

March April

40-

30-

20

i o H

o

50

40-

30-

20

10

OH

1978

-10J r

50-

40-

30-

20

- 1 0 -

March April

1980

March April

1981

March April

Fig 1 For description see opposite

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

60 -i

40-

20

140 n

120 -

100 -

BO-

S 60

40-

20-

lOO-i

80-

60-

40-

20-

0

Growth of contrasting potato varieties(d) Aberporth

1978

447

1977

(e) Trefloyne1979 1980

1980Hemsby

1981

March April May June March April May June

Fig 1 Weekly number of day-degrees gt 4 degC (a-c) and total monthly rainfall (d-f) in 2 years at Aber-porth (north Pembrokeshire) Trefloyne (south Pembrokeshire) and Hemsby (Norfolk) T Time ofplanting v time of 50 plant emergence

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

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Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

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Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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Growth of contrasting potato varieties 445

Experiment 3 1978-9Treatments were nine varieties Primura Home

Guard Arran Comet Pentland Javelin F4611(earlies) Spunta Renova (second earlies) Juliverand Draga (maincrops) replicated three times in arandomized-block design Plots were three rowswide and 11-25 m long Seed tubers of 40-50 50-60and 60-70 g were used for the three replicates andspaced 25 cm apart in the row Number of plantsand above-ground stems were recorded on 18 Mayusing one complete row Growth analysis (twoplants per plot) and estimates of yields (ten plantsper plot) were carried out on 21 June 3 16 and31 July

All seed was once-grown in mid-Wales Seedwas selected before the onset of sprout growth andplaced in trays as in Expts 1 and 2 The seed wasstored in a frost-free glasshouse until planting(average daily temperature 84 degC) Sprout measure-ments were taken on 8 March using ten seed tubersof each size and the experiment planted by handon 19 April

Experiment 4 1979-80In this experiment 11 varieties were compared

Aminca and Maris Bard (earlies) and Wilja (secondearly) were added to those in Expt 3 and HomeGuard deleted There were three replicates plotdimensions and seed size were as in Expt 3Numbers of plants and above-ground stems werenot recorded in this experiment Growth analysisand estimates of yield were carried out as in Expt 3on 18 27 June 8 and 28 July

Seed production treatment selection and storagewere as in Expt 3 The average daily temperatureof the glasshouse was 9-9 degC Sprout measurementswere taken on 3 March using ten tubers of eachseed size and the experiment planted by hand on21 March

Two experiments were carried out at DeepdaleFarms Limited Brancaster Norfolk on light sandysoils of the Wick Series Rows were 76 cm apartand tubers spaced 20 cm apart Both sites werestone-separated prior to planting

Experiment 5 1979-80Nine varieties Home Guard Arran Comet Maris

Bard (earlies) Red Craigs Royal Wilja Estima(second earlies) Maris Piper Pentland Squire andDesiree (maincrop) were compared in three random-ized blocks Seed of 40-55 g was used for all repli-cates Plots were four rows wide and 5 m longPlant and stem emergence were recorded every3-4 days using one whole row per plot Yields wereestimated from 14 plants per plot on 24 June 8 and22 July

All seed was once-grown in mid or west Wales

Table 2 Number of day-degrees gt4C experiencedby seed from onset of sprouting (3 mmsprout) toplant-ing in three varieties

Experiment

Variety

Home GuardArran CometMaris Bard

(once grown)Maris Bard(certified)

1 5 6

640 390 890 mdash 270 mdash630 340 880 760 300 mdashmdash 740 mdash mdash mdash mdash

330 80 mdash 430 230 530

and selected in late December when all seed hadbroken dormancy The first three varieties werestored at 6 degC the next three at 8 degC and the threemaincrop varieties at 12 degC Sprout measurementswere made on 30 tubers per variety on 29 Februaryand the seed moved to Norfolk The experimentwas planted by hand on 21 March A liquid fertilizerdressing of 150 kg N 86 kg P 167 kg K and 30 kgMgha was applied at planting Four applicationsof 25 mm of water were made by rain-gun on 1422 May and 4 18 June

Experiment 6 1980-1Seven varieties Maris Bard Ulster Sceptre

Vanessa (earlies) Wilja Estima (second earlies)Record and Foxton (maincrop) were compared inthree randomized blocks Seed of 70-80 80-90 and90-100 g was used for the three replicates plotswere the same size as in Expt 5 Emergence was notrecorded and yields were estimated from 14 plantsper plot on 8 22 June and 6 July

Seed of all varieties was stored at 6 degC until mid-February when the two maincrop varieties weretransferred to 12 degC Sprout measurements weremade on ten tubers per seed size of each variety on3 March and the experiment planted by hand on5-6 March

A liquid fertilizer dressing of 145 kg N 61 kg Pand 119 kg Kha was applied at planting Threeapplications of 25 mm of water were made by rain-gun on 7 22 June and 7 July In all experimentsthe control of weeds was satisfactory and theapplication of fungicides and aphicides ensuredthat both blight (Phytophthora infestans) and virusdiseases were controlled

In all experiments the yields per hectare wereobtained by harvesting the area occupied by thestated number of plants irrespective of any missingplants From growth analysis samples on someoccasions yields are presented per plant and careshould be exercised if these values are converted toyield per hectare as the sample area was small andinappropriate for such conversion

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446 R L GRIFFITH AND OTHERS

(a) Aberporth

40

30

20

10

0

50

U 40 H

A

pound 30MU

bullvI 20O

I o

I

1977

-10

50

40 -

30-

20-

10-

0 -

-10

March April

(b) Trefloyne

1979

April

1980

May

(c) Hemsby

V

March April

40-

30-

20

i o H

o

50

40-

30-

20

10

OH

1978

-10J r

50-

40-

30-

20

- 1 0 -

March April

1980

March April

1981

March April

Fig 1 For description see opposite

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

60 -i

40-

20

140 n

120 -

100 -

BO-

S 60

40-

20-

lOO-i

80-

60-

40-

20-

0

Growth of contrasting potato varieties(d) Aberporth

1978

447

1977

(e) Trefloyne1979 1980

1980Hemsby

1981

March April May June March April May June

Fig 1 Weekly number of day-degrees gt 4 degC (a-c) and total monthly rainfall (d-f) in 2 years at Aber-porth (north Pembrokeshire) Trefloyne (south Pembrokeshire) and Hemsby (Norfolk) T Time ofplanting v time of 50 plant emergence

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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446 R L GRIFFITH AND OTHERS

(a) Aberporth

40

30

20

10

0

50

U 40 H

A

pound 30MU

bullvI 20O

I o

I

1977

-10

50

40 -

30-

20-

10-

0 -

-10

March April

(b) Trefloyne

1979

April

1980

May

(c) Hemsby

V

March April

40-

30-

20

i o H

o

50

40-

30-

20

10

OH

1978

-10J r

50-

40-

30-

20

- 1 0 -

March April

1980

March April

1981

March April

Fig 1 For description see opposite

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

60 -i

40-

20

140 n

120 -

100 -

BO-

S 60

40-

20-

lOO-i

80-

60-

40-

20-

0

Growth of contrasting potato varieties(d) Aberporth

1978

447

1977

(e) Trefloyne1979 1980

1980Hemsby

1981

March April May June March April May June

Fig 1 Weekly number of day-degrees gt 4 degC (a-c) and total monthly rainfall (d-f) in 2 years at Aber-porth (north Pembrokeshire) Trefloyne (south Pembrokeshire) and Hemsby (Norfolk) T Time ofplanting v time of 50 plant emergence

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

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Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

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Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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60 -i

40-

20

140 n

120 -

100 -

BO-

S 60

40-

20-

lOO-i

80-

60-

40-

20-

0

Growth of contrasting potato varieties(d) Aberporth

1978

447

1977

(e) Trefloyne1979 1980

1980Hemsby

1981

March April May June March April May June

Fig 1 Weekly number of day-degrees gt 4 degC (a-c) and total monthly rainfall (d-f) in 2 years at Aber-porth (north Pembrokeshire) Trefloyne (south Pembrokeshire) and Hemsby (Norfolk) T Time ofplanting v time of 50 plant emergence

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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448 R L GBIFFITH AND OTHEBS

Table 3 Sprout growth per tuber number of emerged plants and stems Expt 1

Length of longest sprout(mm)

Total sprout length (mm)Number of sprouts gt 3 mmNumber of emerged plants

f000sha) 22 iv 77Number of above-groundstemsplant 30 v 77

Number of mainstemsplant30 v 77

HomeGuard

50-4

73-23 0

43-5

4-9

2-0

Table 4 Tuber yields

Total yield 31 v 77Yield gt 25 mm 31 v 77Total yield 10 vi 77Yield gt 25 mm 10 vi 77Total yield 1 vii 77Yield gt 25 mm 1 vii 77Yield gt 38 mm 1 vii 77Number of tubers 31 v 77

10 vi 771 vii 77

HomeGuard

8 05-2

12-610-721-520113-2

1193890780

ArranComet

27-8

6113-3

41-9

5-2

2-8

MarisBard

21-2

6014-6

311

5-7

5-2

[tha) and number of tubers

ArranComet

11-68-6

17-915-931 330-223-3

1054984790

MarisBard

9-74-8

16-513-929026-817-2

20231257979

PentlandCrown

15-7

26-92-8

28-8

5-3

1-9

(OOOsha)

PentlandCrown

5-72-5

1109 2

25-324-3180

994760696

StormontEnterprise

14-9

38-74-3

22-6

5-8

4-2

Expt 1StormontEnterprise

5-52-8

11-69-2

24823-6160

1153959815

SE

1-71

2-450-26062

0-26

0-34

SE

0-660-541-051-091-971-871-72

1016661371

From the records of temperature in storage andobservation of the end of dormancy the numbers ofday-degrees experienced by the seed of the varietiesin the majority of experiments are shown in Table2 Where no direct observation of the end of dor-mancy was made the timing of this event wasobtained from records of the same variety in otherexperiments

Only at Trefloyne in 1980 was wilting of plantsobserved in prolonged dry weather in July In allother experiments growth was not observed to beaffected by water availability The monthly rain-fall figures and weekly number of day-degreesgt 4 degC (air) are given in Fig 1

RESULTS

Experiment 1 (1977)

The early varieties produced substantially moresprout growth and emerged earlier than the main-crop varieties (Table 3) Final emergence was com-plete in all varieties and there were no effects ofvariety on number of above-ground sterna How-

ever Maris Bard and Stormont Enterprise pro-duced predominantly mainstems and had signifi-cantly more than other varieties (Table 3) PentlandCrown produced substantially more secondarystems than mainstems

At the first harvest (31 May) Arran Comet pro-duced the highest total yield and all early varietiessignificantly outyielded the maincrops (Table 4)Arran Comet had markedly higher saleable yield( gt 25 mm) than all other varieties and the other twoearlies were higher yielding than the maincrops Atthe two subsequent harvests Arran Comet producedthe highest total and graded yields although MarisBard was usually not significantly lower yieldingMaris Bard lost many of its small tubers so thatalthough it had the most tubers throughout harvest-ing the differences between varieties in number oftubers became smaller and the effects on total yieldwere also found in graded yields NonethelessArran Comet produced significantly higher yield inthe gt 38 mm grade than other varieties as a con-sequence of having the lowest number of tubers ofall varieties which resulted in increased mean

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Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

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Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

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Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 449

Table 5 Sprout growth per tuber number of emerged plants (OOOsha)and number of stems per plant Expt 2

Length of longest sprout (mm)Total sprout lengthNumber of sproutsNumber of emerged

7 iv 781 v 78

(mm)gt 3 mmplants

HomeGuard

21-942-02-7

46-648-4

Number of above-ground stemsplant18 v 78

Number of mainstemsplant18 v 78

Date of sampling

4 v 7818 v 7830 v 78

7 vi 7816 vi 78

Total yield30 v 78

Yield gt 25 mm30 v 78

Total yield7 vi 78

Yield gt 25 mm7 vi 78

Total yield16 vi 78

Yield gt 25 mm16 vi 78

Number of tubersgt 25 mm30 v 78

HomeGuard

0-561-492-492-952-60

4-4

2-5

Table 6

ArranComet

0-722-434 0 14-764-84

Arran Maris Maris PentlandComet Bard (OG) Bard (CC) Crown

17-740-93-7

29-548-4

5-9

2-9

Leaf area bull

MariaBard (OG)

0-370-570-801-431-44

38-4100-9

4 1

33-540-1

1-5

1-4

5-6 11117-1 2582-8 3-7

5-6 31-449-1 47-2

5-5 8-1

50 2-4

indices Expt 2

MarisBard (CC)

0 1 81-383-283163-32

PentlandCrown

0-471-633-413 6 33-68

Desiree

11-7211

2-4

30148-8

3-8

2 0

Desiree

0-371-432-792-963-50

Table 7 Tuber yields (tha) and number of tubers (y000sha) Expt 2

HomeGuard

19-2

17-8

24-8

23-8

27-3

26-4

450

ArranComet

20-6

19-3

30-5

29-2

351

341

592

MarisBard (CC)

16-8

14-1

24-7

220

3 2 0

29-8

440

MarisBard (OG)

11-6

10-4

16-8

15-6

21-4

20-4

222

PentlandCrown

16-7

14-6

25-9

24-5

30-9

299

451

Desiree

17-8

15-9

22-4

21-1

28-8

27-4

548

SE

0-701-740-26

2-281-35

0-55

0-38

SE

004701390-2030-2670-353

SE

0-93

0-98

0-95

0-95

1-53

1-49

304

tuber size At the second harvest the yields of themaincrop varieties were not significantly lowerthan Home Guard and at the final harvest the twomaincrops exceeded Home Guard although notsignificantly Thus by the end of June the main-crop varieties appeared to have surpassed theyields of the most popular early variety but notthose of two less widely-grown varieties

15

Experiment 2 (1978)The growth of the once-grown stock of Maris

Bard was quite different from all other varietiesSprout growth commenced in October and byplanting the length of the longest sprout was sub-stantially greater than that of all other varietiesEmergence commenced earlier than other varieties

AGS 103

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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en O

Tab

le 8

Sp

rout

gr

owth

per

tub

er (

9 M

arch

) n

umbe

r of

em

erge

d pl

ants

an

d st

ems

(O

OO

sha

) E

xpt

3

Len

gth

of l

onge

st s

prou

t (m

m)

Tot

al s

prou

t le

ngth

(m

m)

Num

ber

of s

prou

ts gt

3 n

unN

o o

f em

erge

d pl

ants

18 v

79

No

of

abov

e-gr

ound

ste

ms

21

v 7

9N

o o

f m

ain

stem

sp

lan

t21

v

79N

o o

f se

cond

ary

stem

sp

lan

t21

v 7

9N

o o

f ab

ove-

grou

nd s

tem

sp

lan

t21

v

79

Dat

e of

sam

plin

g P

rim

ura

21 v

i 7

9 4-

23

vii

79

4-

716

vii

79

4-7

31 v

ii

79

0-6

c

hang

e pe

ak L

87

to 3

1 v

ii 7

9

Pri

mu

ra

12-2

27-6 3-3

39-8

272 1-

7

2-6

4-3

Juli

ver

2-8

3-9

41

2-3

44

Juli

ver

10

024

-5 3-6

42-3

172 2-

3

0-9

3-2

Spu

nta

19-9

36-8 2-8

25-4

124 1-

8

0-7

2-5

Ren

ova

11-6

24

03

0

22-9

210 1-

3

20

3-3

Dra

ga

5-7

14

02-

5

2-5

124 1-

8

0-5

2-3

Tab

le 9

L

eaf

area

indi

ces

Exp

t 3

Spu

nta

2-4

3-9

3-4

2-8

28

Ren

ova

2-5

3-7

4-3

31

28

Dra

ga

2-9

4-0

6-2

31

40

Ho

me

Gu

ard

20

2-3

2-6

1-5

42

Hom

eG

uard

57-4

60-8 1-3

14-4

176 2-

2

11

3-3

Arr

anC

omet

33-8

38-8 1-4

30-1

199 1

1

2-4

3-5

Pen

tlan

dJa

vel

in

12-7

25-3 30

51-2

189 2

1

1-2

3-3

Arr

an

Fen

tlan

d

F4

61

1

8-9

21-4 3-2

45-3

219 3-

2

10

4-2

Com

et

Jav

elin

F

46

11

30

4-2

3-7

1-8

57

2-7

3-8

3-3

0-4

89

2-8

3-7

3-7

1-8

51

SE

1-30

1-91

0-24

3-66

17-7 0-80

0-32

0-31

SE

0-31

0-33

0-36

03

8

mdash

bull 9 BIT tT r hgt O o s W

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

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Growth of contrasting potato varieties 451but approximately 20 of plants failed to emergeowing to little potato disorder The emergedplants were smaller than those of other varietiesand produced lower leaf area indices (L) Thenumber of tubers was less than in all other varietiesand yield was also less than other varieties Thedata for this seed stock of Maris Bard were ana-lysed with all other varieties and are presented butfurther reference to this stock is limited to theimplications of little potato disorder in assessingvarieties

At planting sprout lengths in Home Guard andcertified Maris Bard Arran Comet and PentlandCrown were less than in Expt 1 (Table 5) As inExpt 1 Arran Comet and Home Guard emergedearlier than the maincrop varieties but Maris Bard(CC) emerged later than the maincrop varietiesThe growth of the leaf surface was extremely rapidin Arran Comet and it had the highest leaf areaindices throughout the experiment (Table 6) ItsL value exceeded 4 before the end of May andthroughout June was significantly greater than allother varieties (Table 6) Initially Home Guardhad larger leaf area indices than the remainingvarieties but by 18 May there were no significantdifferences between these varieties During Juneleaf area indices increased in the maincrops andMaris Bard but decreased in Home Guard and on16 June Pentland Crown had significantly largerL than Home Guard

At the first harvest on 30 May yield was highthe mean yield for the experiment was 17-1 thaand Arran Comet produced the highest total yieldand yield in the grade gt 25 mm and these yieldswere significantly higher than all varieties exceptHome Guard (Table 7) Home Guard was notsignificantly higher yielding in these grades thanany other variety but it produced significantlygreater yields in the grade gt 38 mm than themaincrop varieties At the second harvest (7 June)Arran Comet produced significantly larger yields intotal and in the grade gt 32 mm than all othervarieties There were no significant differences inthese yields between Home Guard Maris Bard andPentland Crown but Desiree produced significantlylower yields than Pentlond Crown Home Guardand Arran Comet did not differ significantly inyield of tubers gt 38 mm but both outyielded othervarieties At the final harvest Arran Comet still hadthe highest total and graded yields but these werenot significantly greater than Pentland Crown andMaris Bard Home Guard and Ddsiree producedsimilar yields which were not significantly differentfrom Pentland Crown and Maris Bard (Table 7)Yield at the final harvest was high in relation tothe length of the growing season Although tuberinitiation was not recorded accurately it is un-likely to have occurred before mid-April in any

variety and therefore a total bulking period of nomore than 8 weeks produced an average rate of4-5 thaweek Rate of bulking may have beenrelated to the size of the leaf surface since the twovarieties with the smallest leaf areas (Home Guardand Desiree) had the lowest rates of bulking

As in Expt 1 Arran Comet produced the highestyields throughout the period of harvesting In con-trast Maris Bard (CC) approached the yield ofArran Comet only at the final harvest and was notsignificantly different from the maincrops

Experiment 3 (1979)There were large differences between varieties in

all aspects of sprout growth (Table 8) Arran Cometand Home Guard produced only one or two sproutsof greater length than other varieties There werefew differences between the remaining varietiesexcept that Draga had very short sprouts Emerg-ence did not seem to be determined by sproutlength at planting as Pentland Javelin and F4611were the first varieties to complete emergence andHome Guard was especially slow This delay inemergence in Home Guard (and Arran Comet) wasassociated with coiled sprouts and in Home Guardsome plants failed to emerge owing to little potatodisorder

There were considerable differences between thevarieties in number of above-ground stems and inthe composition of this population (Table 8) Pri-mura and F4611 produced the most above-groundstems but in the former most were secondary stemsand in the latter main stems Only Home Guardfailed to produce a leaf area index of 3 (Table 9)and the largest peak L values were produced byPrimura (early) and Draga (maincrop) Thesepeak values occurred at different times Primuraproduced its leaf surface very rapidly and its Lexceeded 4 in mid-June Draga was much later inproducing its leaf area and reached its peak in JulyAll varieties decreased in leaf area in the later partof July as a consequence of increasing moisturestress in a prolonged period of dry sunny weatherPrimura and Pentland Javelin were close to com-plete senescence by the end of July while Spuntaand Renova lost relatively little leaf area at thistime

At the first and second harvests Primura andthe four British earlies produced the highest totalyields and most were significantly higher than theother Dutch varieties (Table 10) At the thirdharvest Primura was the highest-yielding varietyand significantly better than all varieties exceptF4611 Pentland Javelin and Renova Betweenthe third and fourth harvests four varieties bulkedlittle (Juliver F4611 Arran Comet and PentlandJavelin) and at the final harvest Primura was still

15-2

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Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

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Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

0 T

uber

yie

lds

(th

a)

Exp

t 3

Tot

al y

ield

Yie

ld gt

25

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

38

mm

Len

gth

of l

onge

stsp

rout

(mm

)T

otal

spr

out

leng

th(m

m)

No

of

spro

uts

gt 3

mm

Dat

eof

sam

plin

g

21 v

i21

vi

3 v

ii

3 v

ii

16 v

ii

16 v

ii

31 v

ii

31 v

ii

Pri

mur

a

7-9

19-3

3-2

79 79 79

79

79

79

79

79

Pri

mur

a

10- 7- 31-

28-

45-

43-

49-

44-

Juliv

er

4-8

15-9

30

2 7 7 9 2 7 8 3

Juliv

er

Spun

ta

Ren

ova

7 5- 23 21 35 33 35 32

2 bull7 1 bull4 0 9 bull5 bull4

Tab

le 1

1

Spun

ta

16

0

30-8

2-7

71

50

25-4

22

1

37-6

35-5

45-4

39

1

7-8

5-7

22-3

20-3

42-6

41-4

48

043

-8

Spro

ut

grow

th p

er t

uber

1

Dra

ga

2-2

11

14-6

12

-5

31-4

30-5

41

13

90

[3 i

ii

80)

Mar

iaA

min

ca

Ren

ova

Dra

ga

Bar

d

16-9

10

-2

25-6

16

-4

21

1-9

2-7

6-3

0-9

15

1

23-6 2-3

Hom

eG

uard

11-5

10-6

26-5

24-9

35-4

33-5

40-2

36

0

Exp

t 4 Arr

anC

omet

18-3

27-1 1-7

Arr

an

Pen

tlan

dC

omet

Ja

veli

n

10-3 98

28-8

27-8

36-9

36-4

39-7

37

1 Pen

tlan

dJa

veli

n

10-3

19-6 2-7

13-9

12-8

31-7

29-9

41

040

-1

40-8

36

0

IF

46

11

11-5

28

1

3-7

F46

11

10 9 28

27

419 9 6 bull0 2

40-3

41 37bull0 bull8

Wil

ja

10-7

21-6 2-6

SE

1 1 1 1 1 1 2 1bull02

bull15

bull72

bull75

bull87

bull93

bull02

bull97

SE

0-92

1-38

0-22

bull t1

Q e i gtbull o a CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

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456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

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Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

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458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

2

Tub

er y

ield

s t

ha)

and

num

ber

of

tube

rs (

OO

Os

ha)

Exp

t 4

To

tal

yie

ld18

v

i 8

0Y

ield

gt

2

5 m

m18

v

i 8

0

To

tal

yie

ld27

v

i 8

0Y

ield

gt

32

mm

27

vi

80

To

tal

yie

ld8

vii

80

Yie

ld

gt

32 m

m8

vii

80

To

tal

yie

ld28

v

ii

80Y

ield

gt

32

mm

28

vii

80

Nu

mb

er o

f tu

ber

s18

v

i 8

0

Pri

mu

ra

Juli

ver

S

pu

nta

A

min

ca

Ren

ov

a

20-7

19

-8

23-9

26

-3

24-3

19-9

27-5

26-2

35-1

33-9

40-3

39-3

655

19-2

25-4

24-6

35

0

34-3

44-4

43-6

558

22-6

31

1

29

1

38-7

36-8

45-7

43-8

710

25-3

31-4

29-7

42

0

40-6

49-8

48-6

739

23-3

31-6

30-2

43-9

42-7

58-1

57-0

676

Dra

ga

9-6

8-3

15-3

13-7

29-3

28-4

47-2

46-7

510

Mar

iaB

ard

24-9

24

0

31

0

30

0

43-5

42-4

59-7

58-7

568

Arr

an

Pen

tlan

dC

omet

Ja

vel

in

F4

61

1

23

0

22-3

29

0

27-4

35-6

34-2

41-8

40-5

478

25-0

24-2

29-6

28-2

37-3

35-8

43-3

41-9

720

28-4

27-3

36-6

35-2

47

1

45-8

59-9

59-2

781

Wil

ja

21-1

20-8

30-4

28-2

38-5

37

0

46-7

45-3

628

SE

1-27

1-23

1-52

1-56

2-31

2-24

2-59

2-57

55

0I S

Tab

le 1

3

Spro

ut g

row

th p

er t

uber

on

29

Feb

ruar

y 19

80 a

nd

num

ber

of

emer

ged

plan

ts

(00

0sh

a)

Exp

t 5

Len

gth

of lo

nges

t sp

rout

(m

m)

Tot

al s

prou

t le

ngth

(m

m)

No

of

spro

uts

gt 3

mm

Num

ber

of e

mer

ged

plan

ts 2

1 i

v 8

0N

umbe

r of

em

erge

d pl

ants

24

iv 8

0

Arr

anC

omet

10

130

-3 4-2

62-3

64-5

Hom

eG

uard

15-8

33

13

059

-263

-6

Mar

iaB

ard

6-8

23

14-

554

-863

-6

Red

Cra

igs

Ro

yal

7-2

20-2 3-6

3-5

12-3

Wil

ja

Est

ima

5-3

14-4 2-9

16-7

43-4

4-7

14

03

026

-839

-5

Mar

isP

iper

8-3

22-5 3-6

57-8

62-3

Pen

tlan

dS

quir

e

10

124

-4 3-7

51-3

55-7

Des

iree

12-2

19-6 2-3

54-0

62-3

SE

0-22

1-07

0-22

4-82

4-34

CO

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Tab

le 1

4 T

uber

yie

lds

(th

a) a

nd n

umbe

r of

tub

ers

(O

OO

sha

) E

xpt

5

Tot

al y

ield

24

vi 8

0Y

ield

gt

32

mm

24 v

i 8

0T

otal

yie

ld8

vii

80

Yie

ld gt

32

mm

8 v

ii 8

0T

otal

yie

ld22

vii

80

Yie

ld gt

32

mm

22 v

ii 8

0N

umbe

r of

tub

ers

24

vi 8

0

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Tot

al y

ield

Yie

ld gt

32

mm

Arr

anC

omet

18-7

13-6

25-6

20-8

29-2

24-9

85

6

Num

ber

of t

ub

ers

Dat

e

Hom

eG

uard

11

1

7-4

15-7

13

0

20-9

18-0

632 Tab

le

ofsa

mpl

ing

8 8 22

22

6 6

22

vi

vi

vi

vi

vii vii

vi

81 81 81 81 81

81

81

Mar

isB

ard

20-3

15-3

31-4

28-2

39-6

36-3

817

15

Tub

er

Est

ima

16-5

13-6

22-3

20-7

29-3

27-9

548

Red

Cra

igs

Roy

al

17-8

12

1

23-3

21-3

32-6

30-8

61

7

yiel

ds (

tha

)

Wil

ja

12-8

4-5

18-3

15

0

21-3

17-4

625

Wil

ja

18-1

14-5

27-2

24-8

35-9

33-9

339

and

num

ber

Mar

isB

ard

14-9 8-9

18-0

14-8

21-2

17-8

669

Est

ima

21-6

18-3

36

1

33-9

47-6

46-3

624

Mar

isP

iper

20-0

15-2

32

1

29

1

41-2

38-7

896

of t

uber

s (

000s

ha)

E

xpt

Van

essa

12-7 9-6

21-5

20-4

21-0

19-5

435

Uls

ter

Scep

tre

11-3 6-5

13-9

11-3

15-3

12-2

529

Pen

tlan

dS

quir

e

18

1

16-3

27-7

25-8

37-4

36

1

501

6 Fox

ton

8-6

01

16-7

10

1

18-9

11-9

919

Des

iree

15-9

12-2

26-4

24-5

32-8

31-3

537

Rec

ord

6-7

2-8

13-6

12-2

17-9

16-4

500

s 0- 0- 1- 1- 1- 2- 52-

SE

0-83

0-73

1-22

1-33

1-42

1-46

30

1

B

66 65 45 55 94 08 4

t1 t OTHE

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 455

gt gtmdash copy -H -H copy - I

oH

the highest-yielding variety although not signifi-cantly higher than Renova and Spunta

Primura had more tubers than most varietiesand the ranking order of varieties for graded yieldswas not the same as for total yields The fourBritish earlies produced similar yields in the gradegt 32 mm at the first harvest and all significantlyout-yielded other varieties (Table 10) AlthoughDraga was significantly lower yielding than othervarieties at the second and third harvests therewere few other significant differences betweenvarieties At the final harvest Primura producedthe highest graded yields and as in total yieldsRenova and Spunta were not significantly different

In this experiment the variety with the highesttotal yield again remained constant throughout(Primura) although the ranking of varieties forsaleable yields was more variable than in Expts 1and 2 Arran Comet produced high early yields butwas not outstanding and no higher yielding thanHome Guard throughout the rest of the season Itsyield relative to most other varieties decreasedwith delay in harvesting and was modest by finalharvest

Experiment 4 (1980)Table 11 shows that sprouts were shorter than in

Expt 3 but the relative values of varieties commonto both experiments were unchanged Arran Comethad much shorter sprouts than in Expt 3 F4611produced the highest total yield at all harvests andwas significantly higher yielding than most varie-ties (Table 12) The ranking order of varieties foryield did not change much during the seasonRenova Aminca and Maris Bard were the closestto F4611 throughout and at the final harvestdifferences between F4611 Renova and MarisBard were small Arran Comet produced signifi-cantly lower yields than F4611 at the first harvestand its yields decreased relative to other varietiesduring the season Draga had very low yieldsthroughout

Experiment 5 (1980)The reduced storage temperatures for the early

varieties reduced their sprout lengths comparedwith previous experiments and only Home Guardproduced substantially longer sprouts than themaincrop varieties (Table 13) The maincrop varie-ties emerged as early as the earlies and several daysin advance of Wilja and Estima The emergence ofRed Craigs Royal was severely delayed by coiledsprouts Final emergence was complete in all varie-ties Despite its slightly later emergence Estimaproduced the highest yields throughout the seasonand differences were significant for all comparisonsat the second and third harvests Maris Piper (main-crop) and Maris Bard (early) produced similar yields

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

456 R L GRIFFITH AND OTHERS

throughout ranked second to Estima and weresignificantly higher yielding than all other varieties(Table 14) The other maincrops were significantlyhigher yielding than Home Guard throughout Therelative yield of Home Guard and Arran Cometdecreased as harvesting was delayed and HomeGuard was the lowest-yielding variety throughoutthe season

Experiment 6 (1981)As in Expt 5 Estima produced the highest total

and saleable yields at all dates of harvesting(Table 15) Differences between other early varie-ties were small although Maris Bard was again thesecond highest yielding at the first harvest Thetwo maincrop varieties were very low yieldinginitially but at the final harvest not significantlylower yielding than any variety except Estima

DISCUSSION

The experiments covered five seasons with a widerange of yields and may be taken as a small butrepresentative sample of the environments in whichcrops are grown for harvesting before the end ofJuly The coefficients of variation (Table 16) wereof similar size as those reported in many otherexperiments and only large differences achieve theusual levels of significance In spite of this it ispossible to detect some remarkable variations indifferences between specific varieties at the differ-ent sites and at the different times of harvestingwithin a site Of the varieties present in mostexperiments Arran Comet ranged from clearly thehighest yielding (Expts 1 and 2) through themoderate (Expts 3 and 4) to the low yielding (Expt5) Its yields in Expts 3-5 decreased relative to theother varieties as harvesting was delayed Therewere however some consistent features in thedata In all experiments the variety with thehighest initial yield maintained its position through-out although the magnitude of the advantageusually decreased with delay in harvesting InExpts 4 and 5 the final harvest was in late July andat this time the highest-yielding varieties had largeleaf surfaces and high bulking rates so it 3eems un-likely that they would lose their advantage muchbefore the end of the growing season The resultssuggest that in general high early yields are notprejudicial to high yields later in the season Theimpression that high early yields are prejudicial tolater yields is the consequence of experimentsusing early varieties such as Home Guard in whichold seed is planted and its inadequate leaf coverreduces bulking rates and yields later in the season

When assessing the differences in effects of treat-ments such as varieties shown by individual experi-ments there is usually confounding of the physical

characteristics of the site seasonal weather andvarious aspects of source and management of theseed Nonetheless consideration of the three earlyvarieties Arran Comet Home Guard and MarisBard which were in most experiments providessome evidence of how these factors interact AtTrefin Arran Comet produced a growth patternwhich was almost ideal for producing high earlytuber yields This was the earliest environmentand although planting was early emergence occur-red quickly as a consequence of long sprouts andsufficiently high soil temperatures The advantagesof using old seed for rapid emergence were noteroded by any subsequent disadvantages in size orlongevity of the leaf surface This growth patternwould ensure the superiority of yields of ArranComet for the whole of the harvesting period Incontrast at Trefloyne and Brancaster in sub-sequent years Arran Comet produced more modestyields At Trefloyne the seed was physiologicallyolder mainly as a consequence of later planting andemergence was delayed as a consequence of coiledsprouts and Arran Comet was lower yielding thanseveral varieties throughout harvesting At Bran-caster quite young seed was similarly affected bycoiled sprouts and emergence was also delayed andyields poor Temperatures after planting weresometimes lower at these sites than at Trefin(Fig 1) and soil conditions rather cloddier on theheavier soils at Trefloyne Of greater significance forNorfolk in 1981 was the period of low temperature(mean soil temperature at 10cm 3-1 degC for 2 days)shortly after planting and the period of cold frostyweather in late April (mean soil temperature at10 cm 4-3 degC for 5 days) just as emergence was be-ginning The available temperature data are notextensive but suggest that small differences in meansoil temperature (1 or 2 degC) may markedly affectemergence and subsequent growth of some potatovarieties In such conditions the potential advan-tages of increased physiological age can be negatedby delay in emergence Arran Comet always pro-duced a complete plant stand and a full leaf coverbut away from Trefin so did several other varietieswhich emerged more or as quickly Jones (1981)showed that the rate of post-planting sprout elonga-tion decreased above a certain sprout length insome varieties and in adverse circumstances (lowtemperature and poor soil conditions) this wasassociated with coiling of stems and little potatodisorder The Brancaster experiment shows that inArran Comet coiling of very short sprouts canoccur Similar more dramatic effects were foundin Home Guard Only at Trefin did this varietyyield nearly as much as the highest-yielding earlyand at this site the old seed emerged quickly andcompletely and produced a full leaf cover Howeverthe leaf area began to decrease in June and the

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

Growth of contrasting potato varieties 457yield of Home Guard decreased relative to the othervarieties as harvesting was delayed As in ArranComet the old seed of Home Guard at Trefloynedid not emerge quickly but additionally failed toproduce a complete plant stand Consequently leafarea indices were lower than on other varieties andyields were initially high but decreased relative toother varieties as harvesting was delayed Theseeffects were even more marked at Brancaster wheredespite using younger seed emergence was de-layed leaf area indices were low and yields werethe lowest at the first harvest and remained sothroughout the season

The results for Maris Bard illustrate the effectsof age directly The home-produced seed in Expt 2was too old and produced incomplete emergence ofsmall plants whilst the certified stock was physio-logically much younger and produced higher yieldsthough not significantly greater than some main-crop varieties In the previous year the Maris Bardseed was older than in Expt 2 and emerged quicklyand produced yields almost as high as Arran CometAt Trefloyne (Expt 4) and Brancaster seed ofgreater age than that planted in Expt 1 emergedquickly and produced high yields throughout as acomplete leaf cover was achieved

It seems clear that the maximum benefit of in-creasing age of seed in some early varieties can beobtained only in environments in which rapid andcomplete emergence occurs and a large leaf surfaceis produced Thus only environments such asTrefin with mild temperatures and light soils arelikely to grow old seed of Arran Comet and par-ticularly Home Guard successfully in most or allseasons At harsher sites the advanced age of seedis likely to be disadvantageous because coiledsprouts andor little potato disorder will delayemergence and reduce leaf cover and thereby re-duce tuber bulking As these varieties have only ashort dormant period considerable ageing of seedwill usually occur by planting In early varietieswith longer dormancies there will be considerablevariation in age (ie sprout length) by planting asa consequence of variation in temperatures andearly yields will usually increase with age as inMaris Bard in these experiments and others atTrefloyne (OBrien et al 1983) The temperaturesduring January and February are crucial to theage of such varieties for if they are low (lt 5 degC)little or no ageing will occur and seed will emergeslowly Such effects will also occur in maincropswhich do not usually have much sprout growth atplanting but which can clearly emerge and yield aswell as earlies at some sites Increasing the age of

seed of such varieties by increasing storage tempera-ture as in Expts 2 and 5 can improve their earlyyields considerably However this is still in-adequate for some varieties such as Record andFoxton which both produced poor early yields inExpt 6

The inference from this interpretation of theresults is that the yield estimates of varieties fromexperiments can be markedly affected by the ageof seed planted and the environmental conditionsafter planting These effects are however moreexplicable than hitherto accepted The consequencesof this analysis for the testing of varieties are thatthere is little justification for attempting an overallscore for earliness and the present maturity classi-fication is not necessarily a good guide to the useful-ness of varieties for specific environments andmanagement practices

For breeders the results suggest that there islittle justification in breeding for earliness throughvarieties which achieve advanced physiological ageby the time of planting since such seed may besuitable in only a minority of sites used for theearly crop The greatest opportunity exists forvarieties which have reasonably long dormantperiods (and consequently do not achieve advancedage before planting) but emerge and expand theirleaf surface more rapidly in the prevailing tempera-tures than existing varieties In this respect theobjectives of early and maincrop breeding are notreally different as illustrated by the success ofF4611 (maincrop x maincrop) in these experi-ments The major limitation to the use of main-crop varieties very early in the season is often theirpropensity to produce larger numbers of tubers butthe character can be altered through selectionConsidering all varieties sprout length at plantingwas not closely related to emergence as found byJones (1981) so there may be some prospect ofproducing varieties which achieve reasonably ad-vanced age with short sprouts and still emergerapidly Such varieties would be suited to mech-anical planting and allow commercial exploitationof the advantages of ageing

The authors thank the collaborating farmersMr I Davies Park Court Trefin Dyfed and Greensof Soham Soham Cambs for the provision of sitesand help in carrying out the experiments and MissJ Amies for help in preparing the manuscript Theseed of the Dutch varieties was supplied by Beesonamp Wiskar Ltd The first two experiments were partof a research programme funded by the PotatoMarketing Board

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available

458 R L GRIFFITH AND OTHERS

REFERENCESALLEN E J (1977) Effects of date of planting on

growth and yield of contrasting potato varieties inPembrokeshire Journal of Agricultural Science Cam-bridge 89 711-735

GUNASENA H P M amp HARRIS P M (1971) The effectof CCC nitrogen and potassium on the growth andyield of two varieties of potatoes Journal of Agricul-tural Science Cambridge 76 33-52

JONES J L (1981) Effect of date of planting on con-trasting potato varieties PhD thesis UniversityCollege of Wales Aberystwyth

OBRIEN P J amp ALLEN E J (1981) The concept andmeasurement of physiological age Proceedings of the

Sth Triennial Conference of the European Associationfor Potato Research Munich pp 64-66

OBRIEN P J ALLEN E J BEAN J N GRIFFITHR L JONES SUSAN A amp JONES J L (1983)Accumulated day-degrees as a measure of physio-logical age and the relationships with growth andyield in early potato varieties Journal of AgriculturalScience Cambridge 101 113-131

RUDEFORTH C C amp BRADLEY R U (1972) Soils landclassification and land use in West and Central WalesSpecial Soil Survey No 6 Soil Survey Rothamsted

SOIL SURVEY RECORD No 24 (1974) 1 Soils in Dyfed II(PembrokeHaverfordwest)

at httpwwwcambridgeorgcoreterms httpdxdoiorg101017S0021859600047420Downloaded from httpwwwcambridgeorgcore Faculty of Classics University of Cambridge on 24 Nov 2016 at 154851 subject to the Cambridge Core terms of use available