CHECKLIST OF THE AMPHIBIANS AND REPTILES OF THE …caddolakedata.us/media/1794/hardy, laurence m...

BULLETINof the

MUSEUM of LIFE SCIENCES

NUMBER 10

CHECKLIST OF THE AMPHIBIANSAND REPTILES OF THE

CADDO LAKE WATERSHEDIN TEXAS AND LOUISIANA

LAURENCE M. HARDY

LOUISIANA STATE UNIVERSITY IN SHREVEPORT1995

BULLETINof the

MUSEUM OF LIFE SCIENCES

Number 10, Pages 1-31August 20, 1995

CHECKLIST OF THE AMPHIBIANSAND REPTILES OF THE

CADDO LAKE WATERSHEDIN TEXAS AND LOUISIANA

Laurence M. HardyMuseum of Life Sciences

Louisiana State University in ShreveportOne University Place

Shreveport, Louisiana 71115-2399

1995 HARDY: AMPHmlANS AND REPTILES OF CADDO LAKE

Checklist of the Amphibians and Reptiles of theCaddo Lake Watershed in Texas and Louisiana

Laurence M. Hardy

INTRODUCTION

3

The Caddo Lake Initiative (Anonymous 1993)and the Caddo Lake Scholars project wasestablished in 1993 to promote awareness of theenvironmental importance of the Caddo Lakeecosystem to the inhabitants and visitors to thisinteresting area. In 1994 the Caddo Lake areawas designated a RAMSAR site -- one of themajor wetland ecosystems in the world. Acomprehensive public event (Grandfriends Day)further educated the local citizenery in theenvirons of the lake about the value of theecosystem as an important component of thehealth of our environment. Dahmer (1989) givesa brief history of the region around Caddo Lake.

The drainage system of the Caddo Lakewatershed, approximately 7,000 square miles(Anonymous 1993), encompasses Caddo Parish ofnorthwestern Louisiana and eleven counties inTexas, including all of Camp and Marion countiesand parts of the following counties: Cass,Franklin, Gregg, Harrison, Hopkins, Morris,Titus, Upshur, and Wood (Fig. 1). The recentlynamed Ramsar Wetlands Area, is a uniqueecosystem that includes Cypress swamps,marshes, grasslands, bottomland hardwoodforests, and pine forests in a relativelyundisturbed condition. The Caddo Lake and Lake0' The Pines dams, as flood control and waterlevel stabilization structures, are the majorencroachments on the original natural ecosystem.Residences, subsistence farming, and lightcommerce have added to the ecological impactduring the last 100 years.

The watershed for the Lake 0' The Pines iscompletely contained within the drainage systemof Caddo Lake; therefore any reference to thewatershed for Caddo Lake also refers to thewatershed for Lake 0' The Pines. The CaddoLake watershed is the area of Caddo Lake from168 to 500 feet in elevation (msl). The

elevation above mean sea level is from 168 feetat the spillway of Caddo Lake to 500 feet nearPickton, Texas. Annual rainfall is approximately43-50 inches (Stephens and Holmes 1989).

The drainage system is within the Austro-riparian Biotic Province (Dice 1943). Accordingto Blair (1950) most of the Caddo Lakewatershed lies within the Austroriparian BioticProvince, which is bounded approximately by aline from western Harris County through westernRed River County. This is approximately thesame as the Piney Woods (Stephens and Holmes1989). The western edge of the watershed iswithin the Texan Biotic Province. This region isin the Post Oak Belt of Stephens and Holmes(1989). The eastern two-thirds of the watershedis in the Pineywoods vegetational area of Gould(1969) and the remainder to the west is in thePost Oak Savannah. Habitats include Cypressswamps in the wetlands to oak-hickory to pineforests in the uplands. Much of the land iscultivated in row and broadcast crops and pinefarms. Oil production occurs throughout most ofthe watershed.

The great biological surveys of the 1800'sconcentrated on areas in the American southwestor in areas to the east of the Mississippi River.There have been very few biological studies donein eastern Texas and northwestern Louisiana,even though early studies recognized thedistinctive eastern deciduous forest as differentfrom the grasslands and deserts to the west. Theecotone between the eastern deciduous forest andthe western grasslands includes the Caddo Lakewatershed and it is thus of great importance tocomplete studies of biodiversity and ecology inorder to more clearly define this major transitionzone. This important transition zone has hadvery little study, primarily due to its remotelocation from large population centers and the

4 HARDY: AMPHIBIANS AND REPTILES OF CADDO LAKE

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relative inaccessibility from educational andresearch institutions. This project is one ofseveral which were initiated to further theknowledge and understanding of the biota of theCaddo Lake region .

The objectives of this project were threefold: 1)to compile a bibliography of the scientificliterature that documents museum specimens,specific localities, or other pertinent informationrelated to the amphibian and reptile fauna of theCaddo Lake watershed; 2) to inventory theexisting voucher specimens of amphibians andreptiles in scientific museum collections; 3) fromthe preceding information, to compile a checklistof the species of amphibians and reptiles known toexist in the Caddo Lake watershed.

The earliest explorers in the watershed of theCaddo Lake were probably the survivors of the LaSalle expedition. La Salle had been killed at aplace southwest of the watershed on 19 March1687. After that date the remainder of the partycontinued to the vicinity of the Caddo Indians andRed River. Their march took them through thecenter of the watershed in the spring of 1687. Nobiological specimens are known from thatexpedition. In the years around 1800-1803 theremight have been surveys or explorations in thewatershed related to the Louisiana Purchase. Thesouthern boundary of the watershed from HopkinsCounty, Texas to Latex in Caddo Parish,Louisiana, as defined by us is the same as part ofthe boundary of the watershed for the LouisianaPurchase. The actual area, measured by me usingthe cut and weight method (Lind 1974) is 2,349.5sq. mi.

MATERIALS AND METHODS

The scientific literature was searched for allpapers that specifically referred to specimens fromlocalities known to be within the boundaries of thewatershed. No specimens from the watershedwere located on the Internet. If only counties,and not specific localities, are cited, only recordsdocumented from Marion and Camp Counties areincluded herein. Only Camp and MarionCounties are completely included in the watershed.All known museum collections in Louisiana were

examined for specimens. Also, the majorcollections in Texas, Arkansas, and Oklahomawere studied. Several large, comprehensivecollections and many smaller collections,.including known private collections, in easternTexas, southern Arkansas, and southeasternOklahoma were also examined. A total of 66collections were studied (Table 1).

RESULTS

Burt (1936) is the first scientific report on anyof the amphibians or reptiles of the Caddo Lakewatershed. The first collection was by A. R.Cahn on 27 March 1920. A few collections hadbeen assembled up to 16 April 1993.

Blair (1950) reported two species of snakes(Carphophis vermis, Regina rigida), sevenspecies of salamanders (Necturus beyeri,Amphiuma tridactylum, Ambystoma maculatum,A. talpoideum, A. opacum, Desmognathusfuscus, Eurycea quadridigitatus), and twospecies of frogs (Pseudacris crucifer, Ranapalustris) that reached their western distribu-tionallimits in the Austroriparian Province.

This paper and Ingold (1995) are the firstchecklists based on comprehensive studies ofmuseum specimens and the scientific literature ofthe entire Caddo Lake watershed.

The herpetofauna of the Caddo Lake watershedthat can be confirmed by voucher specimens inscientific museum collections or by specificcitations in the scientific literature consists of 85species, including 27 species of amphibians and58 species of reptiles. An additional sevenspecies are probably present, but are rare andwere not documented in this study.

For each species, specimens are listed by parishor county with the specific locality followed bythe date of collection and the museum number (inparentheses). Specimens identified by "nv"following the museum number were not examinedand verified. Literature records are listed onlywhen the locality is known to be from within thewatershed. Some of the place names cited areincluded in Figure 2. Place names used inlocalities are used in their original spelling.Directions have been abbreviated (i.e. N=north).

6HARDY: AMPHffilANS AND REPTILES OF CADDO LAKE

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Table I. -- Locations and acronyms of the collections of amphibians and reptiles examined for specimens from theCaddo Lake watershed.

7

Collection

Private CollectionAmerican Museum of Natural HistoryArkansas State UniversityAustin College

. Brazosport Museum Natural ScienceBryce C. Brown (in Strecker Mus.)Centenary CollegeChris McallisterCincinnati Museum Natural HistoryCleveland Museum Natural HistoryDallas Museum of Natural HistoryEast Texas Baptist UniversityEast Texas State UniversityEast Texas State UniversityField Museum of Natural HistoryFt. Worth Museum of Science HistoryHardin-Simmons UniversityHenderson State UniversityHouston Museum of Natural ScienceLouisiana State UniversityLouisiana State UniversityLouisiana State UniversityLouisiana Tech UniversityMcNeese State UniversityMichigan State UniversityMidwestern UniversityMilwaukee Public MuseumMississippi Museum Natural ScienceNavarro CollegeNorth Texas State UniversityNortheastern Louisiana UniversityNorthwestern State UniversityOhio State University MuseumOklahoma State UniversityPanola CollegeAcad. of Nat. Sci. of PhiladelphiaPrairie View A&M UniversityRandall MontgomeryRice UniversityRobert A. Vines Science CenterSan Angelo State UniversitySmithsonian InstitutionSouthern Arkansas UniversitySouthern Methodist University

City

Daingerfield, TXNew York, NYJonesboro, TXSherman, TXLake Jackson, TXWaco, TXShreveport, LADeSoto, TXCincinnati,OHCleveland, OHDallas, TXMarshall, TXTexarkana, TXCommerce, TXChicago,ILFort Worth, TXAbilene, TXArkadelphia, ARHouston, TXAlexandria, LABaton Rouge, LAShreveport, LARuston, LALake Charles, LAEast Lansing, MIWichita Falls, TxMilwaukee, WIJackson, MSCorcicana, TXDenton, TXMonroe, LANatchitoches, LAColumbus, OHStillwater, OKPanola, TXPhiladelphia, PAPrairie View, TXJefferson, TXHouston, TXHouston, TXSan Angelo, TXWashington, DCMagnolia, ARDallas, TX

Acronym

AMNHASU-CACBMNSBCBCCCM-DUCCCMNHDMNHETBUETSU- TETSU-CFMNHFWMHSUHSUARHMNSLSUALSUMZLSUSLTUMSUMSUMMUMPMMMNSNCNTSUNLUNSUOSMOSUSPCANSPPVAMRM-JRURA VSCSASUUSNMSAUSMU

8HARDY: AMPHIBIANS AND REPTILES OF CADDO LAKE

Table 1. -- Continued.

1995

Collection

Stephen F. Austin UniversityStrecker MuseumTerry HibbittsTexarkana CollegeTexas A&M UniversityTexas Southern UniversityTexas Tech UniversityTrinity UniversityTulane UniversityUniversity of Arkansas ,FayettevilleUniversity of Arkansas, Little RockUniversity of Arkansas, MonticelloUniversity of Central ArkansasUniversity of HoustonUniversity of MemphisUniversity of MichiganUniversity of New OrleansUniversity of OklahomaUniversity of Texas at ArlingtonUniversity of Texas at AustinUniversity of Texas at TylerWelder Wildlife Refuge

City

Nacogdoches, TXWaco, TXDallas, TXTexarkana, TXCollege Station, TXHouston, TXLubbock, TXSan Antonio, TXNew Orleans, LAFayetteville, ARLittle Rock, ARMonticello, ARConway, ARHouston, TXMemphis, TNAnn Arbor, MINew Orleans, LANorman, OKArlington, TXAustin, TXTyler, TXSinton, TX

Acronym

SFAUSMTH-DTCTCWCTSUTTUTU-SATUARKARKLRARKMUCAUHUMUMMZUNOLAUOMZUTATNHCUTTWWR

LIST OF SPECIES

Class AmphibiaOrder Caudata

AMPIllUMIDAE

Amphiuma tridaetylum (2) -- TX: Harrison:Woodlawn 3.8 mi. N, 04/23/1950 (TNHC9885); nr. Jefferson (Brown 1950). Marion:(Dixon 1987); Carter's Lake, near Jefferson(Sanders 1948). LA: Caddo: 1 mi. N.Vivian in creek, (NWSU 336).

SIRENIDAE

Siren intermedia (7) -- TX: Lake Lone Star, LoneStar,TX, 06/06/1956 (TCWC 17525). Cass:8.4 mi. W Linden, 06/11/1956 (TU 17839).Morris: Lake Lone Star, Lone Star, TX,06/06/1956 (TCWC 17526). Titus: Texas

Utilities Monticello Lignite Mining, WinfieldN section, 10/25/1992 (UTA A-39593); TexasUtilities Monticello Lignite Mining, WinfieldN section, 10/25/1992 (UTA A-39594); TexasUtilities Monticello Lignite Mining, WinfieldN section, 10/25/1992 (UTA A-39595).Wood: Lake Lydia, 02/08/1969 (TCWC29054).

PROTEIDAE

Neeturus beyersi (1) -- TX: Wood: Cedar Lake,1 mi. S Golden, 04/15/1951 (FWM 4196, nv).

SALAMANDRIDAE

Notopthalmus virideseens (5) -- TX: Harrison:S. side of Back Lake, 2.3 mi. N, 1.3 mi. WUncertain, 04/16/1973 (FWM 8489, nv); S.side of Back Lake, 2.3 mi. N, 1.3 mi. W

1995HARDY: AMPHffiIANS AND REPTILES OF CADDO LAKE

9

Uncertain, 04/16/1973 (FWM 8490, nv); S.side of Back Lake, 2.3 mi. N, 1.3 mi. WUncertain, 04/16/1973 (FWM 8515, nv); 2.3mi. N & 1.3 mi. W of Uncertain, 04/16/1973(FWM 8491, nv); 20 mi. E Jefferson (Smithand Sanders 1952). Marion: Carter Lake, E ofJefferson (Burt 1938b); (Dixon 1987). MOrris:2.4 mi. SE of US Hwy 259, TX Hwy 11/49 jctin Daingerfield, 07/11/1993 (FWM 11045, nv);1/2 mi. W Cason (Smith and Sanders 1952).

AMBYSTOMATIDAE

Ambystoma maculatum (2) -- TX: Cass: Woods 1mi. N. on rd. 2.5 mi. NE of Avinger on Hwy155, 11/09/1974 (NLU 37908, nv); 3 mi. WDouglassville (Smith and Sanders 1952).

Ambystoma opacum (13) -- TX: Lake Caddo,OS/27/1920 (AMNH A-6950). Cass: 39 Ian.E Mount Pleasant, Van Terry Ranch,11/06/1988 (UTA A-30680). Harrison: TX:Harrison: 16 1/2 mi. SE Jefferson, 04/01/1934(UMMZ 97311); 4 mi. N of Karnack,04/21/1962 (TCWC 17530); 4 mi. N.ofKarnack, 04/21/1962 (TCWC 17529); CaddoLake State Park, 04/21/1952 (TNHC 28416);Lake Caddo (Brown 1950). Marion: 2 mi.NW, of Karnack, 03/21/1970 (NLU 53181,nv); 2 mi. NW, of Karnack, 03/21/1970 (NLU53182, nv). Upshur: 2.2 mi. NE of Gilmer,03/11/1967 (SM 9372); 3 mi. NE Gilmer,Little Cypress Bayou, 03/08/1969 (UTA A-1551); 3 mi. NE Gilmer, Little Cypress Bayou,03/08/1969 (UTA A-1552). LA: Caddo:Camp on Caddo Lake near Trees, LA,04/08/1966 (NLU 10014, nv); 5.5 mi. SVivian, 04/18/1973 (FWM 8583).

Ambystoma talpoideum (3) -- TX: Marion:(Dixon 1987); Caddo Lake, Doggerty Creekmouth to Cypress Slough, 06/02/1984 (TCWC63359); Jefferson, 0.4 Ian E downtown, alongSt. Hwy. 49, 04/09/1987 (UTA A-24315).LA: Caddo: T16N, R13W, 03/23/1975 (UTAA-5181).

Ambystoma texanum (4) -- TX: Harrison: 7

mi. E of Karnack, 04/21/1962 (TCWC17533); Caddo Lake State Park, 04/21/1962(TNHC 28415). Marion: (Dixon 1987); 5 mi.W Grey, 04/10/1955 (TCWC 13244). LA:Caddo: 5.5 mi. S Vivian, 04/18/1973 (FWM8582).

Ambystoma tigrinum (2) -- TX: Camp: (Dixon1987); Harrison: Little Cypress Bayou at FM134, 4.9 mi. N, 6.8 mi.W Karnack,04/17/1973 (FWM 8548). Titus: Pond nearairport, Mt Pleasant, 06/22/1951 (FWM4140).

PLETHODONTIDAE

Desmognathus auriculatus (16) -- TX: LakeCaddo, OS/27/1920 (AMNH A-12877).Harrison: 3 mi. E Jefferson, 04/25/1960(TCWC 13847); Lake Caddo (Brown 1950).Wood: 11 mi. NE of Hawkins, 03/26/1969 .(SM 12155-68).

Eurycea quadridigitatus (44) -- TX: Lake Caddo,05/03/1920 (AMNH A-6949); Caddo Lake,mouth of Doggerty Creek to Cypress slough,06/02/1984 (TCWC 63361); Caddo Lake,mouth of Doggerty Creek to Cypress slough,06/02/1984 (TCWC 63362-63364). Camp:(Dixon 1987); Near small stream passingbeneath FM 557, 2.6 mi. NW FM 2796,03/29/1992 (UTA A-40282-40287); FerndaleLake, 8 mi. E of Pittsburg, 04/07/1951(FWM 4213, nv). Cass: Hudges [sic;probably Hughes] Spring (SM [BCB 4032-4034, nvJ); Avinger (SM [BCB 5802, nvJ); 2mi. E of Avinger, 04/27/1963 (SM 5802); 5mi. of Hugh [sic; probably Hughes] Springs,04/28/1950 (SM 7479-7480); 16 mi. N ofHugh [sic; probably Hughes] Springs,04/23/1950 (SM 4034); 3 mi. WDouglassville (Smith and Sanders 1952).Harrison: H. R. Taylor House, Harleton,04/07/1962 (TCWC 17599-17600); CaddoState Park, 04/21/1962 (TCWC 17601);Karnack, Caddo Lake State Pk, 04/21/1962(TNHC 28411-28414); Karnack, Caddo LakeState Pk, 06/17/1952 (TNHC 17891-17895,


1995

17898); 7 mi. SE of Jefferson (DMNH 1893,nv); Lake Caddo (Brown 1950). Marion:(Dixon 1987); 16.5 mi. SE Jefferson (Brown1950). Morris: Dangerfield State Park,06/26/1984 (TCWC 63360); 2.4 mi. SE of USHwy 259, TX Hwy 11/49 jct in Dangerfield,07/11/1993 (FWM 11044, nv). Upshur:Beside FM 2796, 2.1 Near small mi. SW FM557, 03/29/1992 (UTA A-40275-40281);stream by Verbena Rd., 0.5 mi. S FM 557,03/29/1992 (UTA A-40324).

Plethodon glutinosus (0) -- TX: Upshur: (Raunand GeIhbach1972).

Order AnuraPELOBATIDAE

Scaphiopus holbrooki (8) -- TX: Cass: 2.2 mi.N.ofjct. 2683 on Hwy 43,11/09/1974 (NLU37905). Harrison: 4.9 mi. N, 6.8 mi. WKarnack, 04/17/1973 (FWM 8550); 4.9 mi. N,6.8 mi. W Karnack (Little Cypress Bayou atFM 134), 04/17/1973 (FWM 8549); 1.6 mi.NE , Karnack, Caddo Lake State Pk,04/21/1962 (TNHC 29031); FM 134 NWKarnak. 1.7 mi. N, 2.6 mi. W Karnak,04/16/1973 (FWM 8522, nv). Marion:(Dixon 1987); Excelsior House, Jefferson,05/11/1968 (SM 9310). LA: Caddo: 10.2 mi.E Leigh (Harrison Co.,TX), 04/17/1973 (FWM8564, nv); 1 mi. E Mooringsport, 05/03/1952(TCWC 9312).

BUFONIDAE

Bufo americanus (3) -- TX: Morris: 2.4 mi. SEof US Hwy 259, TX Hwy 11/49 jct inDaingerfield, 07/11/1993 (FWM 11048, nv);2.4 mi. SE of US Hwy 259, TX Hwy 11/49jctin Daingerfield, 07/12/1993 (FWM 11049,nv). Titus: 0.8 mi. E jct. Co. rd. NW 3 onCo. rd. NW 18, 06/11/1991 (UTA A-37934).

Bufo valliceps (0) -- Conant and Collins (1991).

Bufo woodhousei (155) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12880-12885, 12894,

nv). Camp: (Dixon 1987); Franklin: 0.6mi. W jct. FM 2723 on FM 3122,05/13/1991 (UTA A-37733); 0.5 mi. E jct.FM 21 on Co. rd. 4145, 05/13/1991 (UTAA-37734). Harrison: Roads of Pine Harborsubdivision off Hwy 729, 11/09/1974 (NLU37911-37920, nv); FM 134 NW Karnak. 1.7mi. N, 2.6 mi. W Karnak, 04/16/1973(FWM 8523); 2.0 mi. NW TX 43 via FM134. 1.7 mi. N, 2.6 mi. W, Karnak,04/17/1973 (FWM 8540); 1 mi. S, 0.4 mi. EKarnack (ctr),4.0 mi. S FM 2198 via FM134, 04/17/1973 (FWM 8556); 2.9 mi. S,0.9 mi. E Karnack, 04/17/1973 (FWM8557); 3.5 mi. E Leigh; 1.1 mi. W LAborder via FM 1999, 04/17/1973 (FWM8558); 3.5 mi. E Leigh; 1.9 W LA bordervia FM 1999, 04/17/1973 (FWM 8559); 3.6mi. E Leigh; 1.8 W LA border via FM1999, 04/17/1973 (FWM 8560-8561); 5.3mi. E Leigh; 0.1 mi. W LA border via FM1999, 04/17/1973 (FWM 8562); 0.8 mi. N,0.5 mi. W Leigh, 04/18/1973 (FWM 8595-8603); Lake Caddo, OS/27/1920 (AMNH A-12886, nv); Caddo Lake (SM [BCB 2730-34, nv], 2730-2734); Caddo Lake State Park(SM [BCB 6297-6300, nv]); Caddo LakeState Park, 06/02/1984 (TCWC 63266-63268); Caddo Lake State Park, 1.6 mi. NEof Karnack, 04/21/1962 (TNHC 29032-29033, 29135); Caddo ST Park, 06/28/1961(DMNH 3491, nv); Karnack, Caddo LakeState Park, 06/18/1952 (TNHC 18163-18164); 1 mi. W of Harleton, 04/19/1984(TCWC 17733-17734); 11 mi. NE ofKarnack, 04/15/1960 (TCWC 14167-14168);8.1 mi. NW Karnack, 04/21/1982 (TCWC17728-17732); 1.7 mi. N, 2.6 mi. WKarnack, via FM 134, 04/16/1973 (FWM8524); 2.3 mi. N, 1.3 mi. W Uncertain,04/16/1973 (FWM 8492-8497); Vicinity ofCaddo Lake St. Pk., 04/20/1962 (SM 4526-4527); Vicinity of Caddo Lake St. Pk., N ofMarshall, 0.4/20/1962 (SM 4528-4529); 2mi. NE of Caddo Lake St. Pk., 04/22/1962(SM 6297-6300); Caddo ST Park 2 mi. N 1mi. E Karmack, 06/27/1960 (DMNH 3490);10 mi. NE of Marshall, 08/10/1951 (TNHC

1995

HARDY: AMPHIBIANS AND REPTILES OF CADDO LAKE

(Dixon 1987); Ferndale Lake, 8 mi. EPittsburg, 04/07/1951 (FWM 4102-4103).Gregg: 14 mi. N of Longview, 04/27/1963(SM 6337, nv). Harrison: Caddo Lake (SM2738-2741 [BCB 2738-2741, nv]); CaddoLake State Park (SM [BCB 6336, nv]);Caddo Lake St. Pk, 04/27/1963 (SM 6336);Caddo Lake State Park (UTA A-28220);Jefferson City (SM [BCB 2591-2592, nv]);Caddo Lake State park, 1.6 mi. NE Karnack,04/21/1962 (TNHC 35857-35860); KarnaCk,Caddo Lake State Park, 06/18/1952 (TNHC17673-17674); Karnack, 3 mi. W,06/18/1952 (TNHC 17650-17672); N ofKarnack, border of Caddo Lake in state park,09/09/1950 (UM [MVP 7500-7504]); 8 mi.SE of Jefferson City, 04/21/1950 (SM 2591-2592); Vicinity of Caddo Lake St. Pk., N ofMarshall, 04/20/1962 (SM 4517-4518,4520_4525); 7 mi. SE of Jefferson, 04/21/1950(DMNH 2074, nv); 2 mi. N, 1 mi. EKarnack, Caddo State Park, 06/27/1960(DMNH 2477, nv); 16.0 mi. NE Marshall(Caddo Lake), 06/14/1957 (UTA A-1266,11116-11155); Caddo Lake, Uncertain,06/04/1985 (UTA A-18089); Lake Caddo(Brown 1950). Marion: (Dixon 1987); 3 mi.N Smithland, 08/03/1949 (TU 16269).Morris: Daingerfield (SM [BCB 6349-6350,nv]); 5 mi. N of Daingerfield, 04/27/1963(SM 6349-6350); Rocky Branch (SM [BCB6417-6418, nv]); 3 mi. N of Rocky Branch,04/27/1963 (SM 6417-6418); From a smallroadside canal beside US 259 at jct w/ 1-30,10/30/1992 (UTA A-39565). Titus: HorseCreek, 0.5 mi. S Hwy. 1-30, 10/14/1991(TCWC 69250-69251); Monticello Lignitemining area, Winfield North: pond AR-24,07/11/1991 (UTA A-37763-37764); 7 mi. WMt.Pleasant, 04/21/1950 (SM 9696-9697);Hwy 49 at inter Hwy 11, 11/09/1974 (NLU37932-37934, nv). Upshur: Little CypressCreek, 2.2 mi. from Gilmer, 03/11/1967(SM 7839). LA: Caddo: Vivian (SM[BCB 2895, 2897, 3078, 3097-3098, 3115,3117, 3119-3120, nv]); Caddo Lake,06/23/1947 (TU 328, nv); Caddo Lake,06/24/1947 (TU 377, nv); E

11911); Lake Caddo (Brown 1950). Marion:(Dixon 1987); Caddo Lake, 04/20/1962 (SM6288-6290); Caddo Lake (SM [BCB 6288-6290, nv]); Jefferson, 08/26/1950 (LSUMZ52031-52040); FM 2682 E of 1-43, 07/20/1979(UTA A-34354). Morris: Rocky Branch (SM[BCB 6088, nv]); Rocky Branch (SM [BCB6291-6292, 6294, nv]); 2 mi. N of RockyBranch, Hwy 338, 04/27/1963 (SM 6088); 1.5mi. S of Rocky Branch, 04/27/1963 (SM 6291-6292); 5 mi. S of Rocky Branch, 04/27/1963(SM 6294); 1 mi. S of Naples, 04/27/1963(SM 6293); 2.3 mi. N of Dangerfield,04/27/1963 (SM 6295). Titus: 1 mi. EWinfield, OS/28/1974 (DMNH 2143, nv); 2 mi.NE Winfield, OS/28/1974 (DMNH 2144, nv); 3mi. NE Winfield, OS/29/1974 (DMNH 2142,nv); On Co. Rd. NW 13 xx rd. mi. N jct. FM1734, 07/11/1991 (UTA A-37742-37743, nv);0.1 mi. N jct. FM 1734 on CR NW 15,03/04/1992 (UTA A-37926); 0.3 mi. N jct.FM 1734 on CR NW 15, 03/04/1992 (UTA A-37927); 0.9 mi. E jct. Co. rd. NW 3 on Co.rd. NW 18, 06/11/1991 (UTA A-37929); 1.3mi. E jct. Co. rd. NW 3 on Co. rd. NW 18,06/11/1991 (UTA A-37930). Upshur: LakeO'The Pines, 14 mi. NE of Gilmer,04/26/1963 (SM 6182); Gilmer (SM [BCB6182, nv]); 22 mi. NE of Gilmer, 04/11/1967(SM 8594); Glenwood (KU 17152). Wood: 15mi. SE of Detriot, 05/03/1969 (SM 8492, nv);LA: Caddo Lake, 06/29/1947 (TU 706, 708).Caddo: Caddo Lake, 06/22/1947 (TU 4495);Caddo Lake, 06/23/1947 (TU 309, 4492, nv);Caddo Lake, 06/24/1947 (TU 349-352, 358-359, 4478, 4480, 4493-4494, 4497, nv); CaddoLake, 06/25/1947 (TU 394); Caddo Lake,06/30/1947 (TU 670); Vivian, 06/27/1947 (TV4479,4482, nv); 8 mi. E Leigh (Harrison Co.,TX), 04/17/1973 (FWM 8563); 10.2 mi. ELeigh (Harrison CO.,TX), 04/17/1973 (FWM8565); 5.5 mi. E Leigh (Harrison CO.,TX),04/17/1973 (FWM 8568-8569); Caddo Lakedam, 09/09/1973 (LSUS 3097). Caddo Lake,07/04/1972 (NWSU 4324).

HYLIDAEAcris crepitans (143) -- TX: Lake Caddo,

OS/27/1920 (AMNH A-12887-12890). Camp:

11


191

Caddo Lake at Caddo Lake State Park nr. OilCity, 04/18/1973 (FWM 8586-8588, nv);Jeems Bayou at Hwy 2, 1.9 mi. W Hwy 1,11/04/1967 (NLU 13825-13828, nv).

Hyla chrysoscelis (38) --TX: Cass: Hwy 43 2 mi.N. of jct Hwy 2683, 11/09/1974 (NLU 37972,nv). Franklin: 1.5 mi. E of 115 on 3122,06/24/1984 (TCWC 63206). Gregg: 14 mi. Nof Longview, 04/27/1963 (SM 6376, nv, 6425,nv); Harrison: Caddo St. Park, 06/29/1961(DMNH 3397-3398, nv); Vicinity of CaddoLake State Park, N of Marshall, 04/20/1962(SM 4307-4308);1.7 mi. N, 2.6 mi. WKarnack (FM 134 NW Karnack), 04/16/1973(FWM 8525-8537, 8542-8543); 2.0 mi. NWTX 43 via FM 134. 1.7 mi. N, 2.6 mi. W,Karnak, 04/17/1973 (FWM 8541); 4.9 mi. N,6.8 mi. W Karnack (Little Cypress Bayou atFM 134), 04/17/1973 (FWM 8551, nv); 0.8mi. N, 0.5 mi. W Leigh, 04/18/1973 (FWM8604-8606). Marion: (Dixon 1987); two mi.E of Jefferson, 08/03/1949 (TU 16258).Morris: Rocky Branch (SM [BCB 6419-21,nv]); 3.5 mi. S of Rocky Branch, 04/27/1963(SM 6419-6421). Titus: 1.0 mi. E jct. Co.rd. NW 3 on Co. rd. NW 18, 06/11/1991(UTA A-37932-37933); Upshur: 5.8 mi. WDiana on Rt 154, (KU 145696). LA: Caddo:Jeems Bayou at Hwy. 2, 1.9 mi. W. Hwy. 1,11/04/1967 (NLU 13762);9.9 mi. E Leigh(Harrison Co.,TX), 04/17/1973 (FWM 8566);2.4 mi. E, 0.5 mi. S Mooringsport,04/29/1986 (LSUS 5345). This is a compositeaccount for Hyla chrysoscelis (diploid) and H.versicolor (tetraploid). These are cryptic,sibling species that are indistinguishable frommorphological characteristics. According toRalin (1968) H. versicolor might be present inthe western part of the watershed and H.chrysoscelis in the eastern part.

Hyla cinerea (140) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12896); Caddo LakeState Park, 02/06/1984 (TCWC 63210); 1.5mi.E. of Hwy 115 on 3122, 06/24/1984 (TCWC63213); Dangerfield State Park, 06/26/1984(TCWC 63214). Franklin: 1.5mi. E. of Hwy

shore of Caddo Lake at Caddo Lake State P3I04/18/1973 (FWM 8584-8585, nv); E 115 I

3122, 06/24/1984 (TCWC 63212). Gr~Edwards Survy, Orr Farm, 10/01/1938 (S6665, 7017-7033). Harrison: 2 mi. N 1 nE Karnack, 06/27/1960 (DMNH 2876, nv); 2mi. N, 1.3 mi. W Uncertain, 04/16/19:(FWM 8498); 1.2 mi. NE FM 2198 via diroad.S side Back Lake, 04/16/1993 (FWI8520); Caddo Lake State Park, 02/06/19!(TCWC 63209); Caddo Lake State Pari06/01/1984 (TCWC 63211); Karnack, 3 mi. 1Karnack, 06/18/1952 (TNHC 18034-18098,Karnack at Caddo Lake State Pk., 06/18/195(TNHC 18099); 16.0 mi. NE Marshall (CaddLake), 06/14/1957 (UTA A-129); 5.0 mi. !Marshall, 07/02/1961 (UTA A-546, 547, nv)16.0 mi. NE Marshall (Caddo Lake)06/14/1957 (UTA A-62-72, 73, nv, 74-75)Lake Caddo (Brown 1950). Titus: 2 mi. NIWinfield, OS/28/1974 (DMNH 2145, nv);Monticello Lignite mining area, WinfieICNorth, (UTA A-37616); 0.5 mi. N jct. FM1734 on Co. rd. NW 13, 05/16/1991 (UTA A37737); 0.1 mi. W Co. rd. NW 15 on Co. rd.NW 6,05/16/1991 (UTA A-37738); 0.3 mi. Njet. FM 1734 on CR NW 13, 03/04/1992(UTA A-37914); 0.8 mi. E jct. Co. rd. NW 3on Co. rd. NW 18, 06/11/1991 (UTA A37935); Texas Utilities Monticello LigniteMining, Winfield N section, 08/12/1992 (UTAA-39591-39592). LA: Caddo: Caddo Lake,04/05/1969 (LTU 3118); Caddo Lake,06/25/1947 (TU 389, p.v, 2579); Caddo Lake,06/23/1947 (TU 318-327, 330-336); CaddoLake, 06/24/1947 (TU 384).

Hyla squirella (1) -- TX: Harrison: 2 mi. N, Imi. E Karnack 06/27/1960 (DMNH 3361, nv).

Pseudacris clarki -- Conant and Collins (1991).

Pseudacris crucifer (1) -- TX: Harrison: 20 mi.E Jefferson (Smith and Sanders 1952).Marion: (Dixon 1987); Upshur: 8 mi. N ofGilmer, 03/11/1971 (SM 11629).

Pseudacris streckeri (1) -- TX: Gregg: 9 mi. N


13

of Longview, 04/14/1960 (TCWC 13902).Morris: 1/2 mi. W Cason (Smith and Sanders1952).

Pseudacris triseriata (15) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12878-12879). Cass: 9mi. NW of Jafferson, 03/14/1971 (SM11381). Gregg: 8 mi. N Longview,04/13/1960 (TCWC 13903). Harrison: 7 mi.E of Karnack, 04/21/1962 (TCWC 17861-17863); 1.6 mi. NE in Caddo State Pk.,04/21/1952 (TNHC 29028); Lake Caddo(Brown 1950). Marion: (Dixon 1987); 4 mi.N of Karnack, 04/17/1955 (TCWC 13657).Morris: 1 mi. N of Rocky Branch, 04/27/1963(SM 5921); 3 mi. S of Naples, 04/27/1963 (SM5922); 1.5 mi. N of Rocky Branch, 04/27/1963(SM 5923). Upshur: Little Cyprus Creek, 2.2mi. NE of Gilmer, 03/11/1967 (SM 7982-7983). LA: Caddo: Oil City, 02/20/1965(LSUMZ 17394).

MICROHYLIDAE

Gastrophryne carolinensis (10) -- TX: Camp:(Dixon 1987); Cass: Lone Star (SM [BCB6506, nv]); 2 mi.NE of Lone Star, DangerfieldPk., 04/27/1963 (SM 6506); 3 mi. SDouglasville (Nelson 1972); 3 mi. WDouglassville (Smith and Sanders 1952).Harrison: 2.3 mi.N & 1.3 mi.W of Uncertain,04/16/1973 (FWM 8499, nv); 1.7 mi. N, 2.6mi. W Karnack (2.0 mL NW TX 43 via(FWM 8544, nv); Caddo Lake (SM 2737);Caddo Lake (SM [BCB 2737, nv]); Karnack,Caddo Lake State, 06/18/1952 (TNHC 17503);Caddo Lake, Caddo Lake State Park nearKarnack, and 15 mi. N Marshall (Nelson 1972).Morris: Douglasville (SM [BCB 6507-6508,nv]). Titus: 1.0 mi. W jct. Co. rd. NW 3 onCo. rd. NW 18,06/11/1991 (UTA A-37928).

Gastrophryne olivacea (0) -- Dixon (1987).Nelson (1972) lists the locality, 1 mi. NDouglasville [sic], Morris Co., and cites thisfrom Bryce Brown (no specimen numbers weregiven). Douglassville is in Cass County and isout of the watershed. [Our study of the

Strecker Museum collection revealed twospecimens (BCB 6507-6508) with the localityDouglasville, Morris Co. Neither were locatedfor verification, but they were listed by themuseum as G. carolinensis.]

RANIDAE

Rana areo/ata (0) -- Conant and Collins (1991).

Rana catesbeiana (39) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12897). Cass: 5 mi.W. of Atlanta on Hwy 59, 03/20/1971 (LTU3868). Franklin: White Oak Creek on Hwy37,06/25/1984 (TCWC 63226). Gregg: PrinceBrothers Lake, 10/08/1938 (SM 3354, nv,3357, nv, 4519, nv); Prince Brothers Lake, OrrFarm, 10/08/1938 (SM 5388, nv, 5792, nv,6707, nv). Harrison: 3 mi. SW of Karnack,06/20/1952 (TNHC 18175-18183); 5 mi. ELeigh, 3.1 mi.E FM 9 via FM 1999,04/18/1973 (FWM 8592); 0.6 mi. N, 0.4 mi.W Leigh (0.8 mLS FM 2682 via FM 134),04/17/1973 (FWM 8655); 15 mi. S Marshall,09/09/1950. (UM [MVP 7505], nv); CaddoLake, 06/10/1961 (UTA A-419-420); LakeCaddo (Brown 1950). Marion: (Dixon 1987);Morris: 2 mi.S Naples, 04/27/1963 (SM 5271-5272). Titus: 2 mi. W of Mt. Pleasant,04/27/1969 (SM 10867-10868); MonticelloLignite mining area, Winfield N, 07/30/1991(UTA A-37617); 0.1 mi. N jct. FM 1734 onCR NW 15, 03/04/1992 (UTA A-37905-37906); Texas Utilities Monticello LigniteMining, Winfield N section, 04/11/1991 (UTAA-38006-38007). Upshur: 2.2 mi.NE ofGilmer, 03/11/1967 (SM 8936-8937). Wood:15 mLSE of Detriot, 05/03/1969 (SM 10871,nv). LA: Caddo: Caddo Lake, 06/23/1947(TU 308); Caddo Lake, 06/24/1947 (TU 348);Caddo Lake, 06/24/1947 (TU 383); CaddoLake, 06/25/1947 (TU 396).

Rana clamitans (58) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12891, 12893).Camp: 7 km SW of Pittsburg, 04/28/1987(TCWC 66167). Cass: Beside Timber Branchat jct. with St. Hwy. 43, 07/24/1993 (UTA A-

95 1995HARDY: AMPHffilANS AND REPTILES OF CADDO LAKE

15

Caddo Lake, 06/23/1947 (TU 317); CaddoLake, 06/24/1947 (TU 347, 360); Caddo Lake,06/25/1947 (TU 385-388); Caddo Lake,06/30/1947 (TU 628).

Rana palustris (5) -- TX: Cass: Woods 1 mi.N. on dirt rd. 2.5 mi. NE of Avinger on Hwy155, 11/09/1974 (NLU 37922). Harrison: 20mi. E Jefferson (Smith and Sanders 1952).Marion: (Dixon 1987); Upshur: 0.5 mi. EDiana on Rt. 154 (KU 145764); 6 mi. NEDiana, 04/15/1960 (TCWC 13959); 1.9 mi. NCypress Creek on rd. connecting 154 & 852,04/09/1968 (UTA A-41705-41706).

Rana utricularia (43) -- TX: Lake Caddo,OS/27/1920 (AMNH A-12892); Camp:Ferndale Lake, 04/07/1951 (FWM 4655-4656);Near small stream passing beneath FM 557, 2.6mi. NW FM 2796, 03/29/1992 (UTA A-40322). Franklin: 0.8 mi. W Titus countyline on NE 2170, 09/21/1991 (UTA A-37732).Harrison: Karnack, Caddo Lake State Pk06/18/1952 (TNHC 18205-18206). 4.8 km. WCaddo Lake State Park on FM 134, Big CypressBayou, 06/04/1985 (UTA A-18126); 4.9 mi.N, 6.8 mi. W Karnack, 04/17/1973 (FWM8552, nv); 2.8 mi. N, 3.8 mi.W Karnack (NWof Karnack via FM 134), 04/16/1973 (FWM8539). Caddo Lake (SM 2736); Caddo Lake(SM [BCB 2736, nv]); Lake Caddo (Brown1950). Hopkins: Club Lake Greenville,10/17/1952 (DMNH 3570, nv). Marion:Caddo Lake, 07/21/1979 (UTA A-34450);(Dixon 1987). Morris: 3 mi. N Daingerfield,04/27/1963 (SM 5158); 2.4 mi.SE of US Hwy259, TX Hwy 11/49 jct in Daingerfield,07/12/1993 (FWM 11047, nv). Titus: 3 mi.S Mount Pleasant, 03/14/1971 (SM 11609);Monticello Lignite mining area, WinfieldNorth, 07/30/1991 (UTA A-37618). 0.2 mi. Njct. CR NW 18 on NW 13, 03/04/1992 (UTAA-37917). 0.5 mi. S jct. CR NW 36 on NW13,03/04/1992 (UTA A-37918). 3.4 mi. S jct.Co. road NW 36 on Co. road NW 13,03/04/1992 (UTA A-37919). 1.3 mi. S jct.NW 15 on Co. road NW 18, 03/04/1992 (UTAA-37920). Texas Utilities Monticello Lignite

Mining,Winfield N section, 04/10/1991 (UTAA-37978). Wood: 7 mi. NE Quitman,03/23/1951 (FWM 4657-4658). Cheaton'sFarm, 07/22/1949 (FWM; 4634). 7mi. NEQuitman, 03/23/1951 (FWM; 4659); 15 mi.SEof Detroit, 05/03/1969 (SM 9134, nv).Upshur: Maxwell Ranch, 3.5 mi.SERhonesboro, 03/14/1971 (SM 13015, 13024-13026). Little Cypress Creek, 2.2 mi.NE ofGilmer, 03/11/1967 (SM 13111-13112); nearsmall stream by Verbena Rd., 0.5 mi. S FM557,03/29/1992 (UTA A-40323). LA: Caddo:Caddo Lake, 06/30/1947 (TU 709-716); 2.4mi. E, 0.5 mi. S Mooringsport, 04/29/1986(LSUS 5342). Caddo Lake, 07/04/1972(NWSU 4376). Hillis (1981) and Dixon (1987)document only one species of the Rana pipienscomplex in the watershed. According to theInternational Code of Zoological Nomenclaturethe name Rana utricularia is the valid name ofthis taxon; however, a petition is underconsideration to conserve Rana sphenocephalafor this taxon. Rana utricularia is used heresince that petition has not yet been decided andR. utricularia is in general use (i.e. Conant andCollins 1991).

ReptiliaOrder TestudinataCHEL YDRIDAE

Chelydra serpentina (2) -- TX: Camp: 7.6 kmSE Leesburg, 08/23/1986 (TCWC 66168).Harrison: 10 mi. N Marshall (Smith andSanders 1952). Titus: 2 mi. E Mt. Pleasant(Brown 1950); 3 mi. NW Mt. Pleasant,04/13/1968 (TU 18937); 4 mi. E Mt. Pleasant(Brown 1950).

Macroclemys temmincki (2) -- TX: Marion:(Dixon 1987); Cypress Bayou, Wend CaddoLake, 06/03/1984 (TCWC 63940). Wood:Rock Falls Hunting and Fishing Club,06/02/1986 (UTA R-16714, nv).

KINOSTERNIDAE

Kinostemon subrubrum (21) -- TX: Lake Caddo,


1995

OS/27/1920 (AMNH R-16986). Camp:(Dixon 1987); 4 mi. W Hwy. 557 from Campcounty line, 06/04/1984 (TCWC S-493);Ferndale Lake, 11.2 km SW Pittsburg,06/04/1979 (UTA R-I1185-11187). Franklin:NW comer Lake Franklin, 07/11/1990 (UTAR-28814). Harrison: 5 mi.E of Marshall,04/23/1968 (DMNH 3838, nv); Harleton,04/19/1962 (TCWC 18409-18410). Marion:(Dixon 1987); Morris: 1 mi. N Omaha,04/19/1957 (TCWC 14576). Titus: 5 mi. NWof Mount Pleasant; N. A. Mattingly Farm,04/18/1969 (SM 8889); Monticello lignitemining area, Winfield N, pond B-12,07/11/1991 (UTA R-32385); 2.5 mi. E of jctCR NW 15 on FM 1734,05/13/1991 (UTA R-32681). Upshur: 2.2 mi. NE of Golmer,04/11/1967 (SM 7921). Wood: 1.0 mi.E FM510, 07/27/1975 (UTA R-30632). LA:Caddo: Caddo Lake Oil City, 07/03/1972(NWSU 4758); Caddo Lake, 06/30/1947 (TU669-700); Caddo Lake, 07/01/1947 (TU 745);Oil City, 05/31/1967 (LSUMZ 23650); one mi.NE Trees on La. 2, 05/31/1965 (LSUMZ23649).

Stemotherus carinatus (18) -- TX: Camp:Ferndale Lake, 11.2 km SW Pittsburgh,06/04/1979 (UTA R-11188-11191). Franklin:S side Lake Bob Sandlin at FM 3007,07/11/1990 (UTA R-28815). Harrison: 5 mi.NE of Karnack at Caddo Lake, 08/01/1950(TNHC 11914-11915); Cypress Creek 2 mi.NECaddo State Park, 10/14/1967 (UTA R-1295).Marion: (Dixon 1987); Caddo Lake,07/21/1979 (UTA R-30627). Upshur: 12 Km.SSW Pittsburg, 04/27/1987 (TCWC 66171).LA: Caddo: Caddo Lake, 06/30/1947 (TU682); Caddo Lake, 06/25/1947 (TU 461);Caddo Lake, 06/27/1947 (TU 506-507); CaddoLake, 06/28/1947 (TU 634); Caddo Lake,06/30/1947 (TU 681, nv, 683).

Stemotherus odoratus (91) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16985). Camp: (Dixon1987); Ferndale Lake, 7 mi. SW Pittsburg,04/05/1979 (TCWC 66212-66213); FerndaleLake, 11.2 km SW Pittsburgh, 06/04/1979

(UTA R-I1192-11218). Franklin: White OakCr., on Hwy. 34, 06/26/1984 (TCWC 62609).Harrison: Caddo Lake, 04/20/1962 (TCWC18407); Caddo Lake, Big Cypress Bayou,06/02/1984 (TCWC 62585-62606, 62677-62696); Lake Caddo (Brown 1950). Marion:(Dixon 1987); Caddo Lake, Cypress Bayou,and Carter Lake, 06/03/1984 (TCWC 62607-62608, 62697-62698); Caddo Lake,07/21/1979 (UTA R-30630); LA: CaddoLake, 06/30/1947 (TU 680). Caddo: CaddoLake Oil City, 07/02/1972 (NWSU 4763);Caddo Lake, 06/25/1947 (TU 459-460);Caddo Lake, 06/27/1947 (TU 504-505, 541);Caddo Lake, 06/28/1947 (TU 637); CaddoLake, 06/30/1947 (TU 659-660,679, 736).

EMYDIDAE

Chrysemys picta (0) -- TX: MarionlHarrison:Caddo Lake (Dixon 1987).

Deirochelys reticularia (1) -- TX: Harrison: 5mi. NE Karnack at Caddo Lake, 08/01/1950(TNHC 11912). Marion: (Dixon 1987).

Graptemys kohni (2) -- TX: Camp: (Dixon1987); Ferndale Lake, 11.2 km SW Pittsburg,06/04/1979 (UTA R-30049). Marion: (Dixon1987). LA: Caddo: Caddo Lake, 06/30/1947(TU 698).

Graptemys pseudogeographica (4) -- TX: LakeCaddo, OS/27/1920 (AMNH R-16967,16984). Harrison: 5 mi. NE of Karnack atCaddo Lake, 08/01/1950 (TNHC 11913, nv);Lake Caddo (Brown 1950). LA: Caddo:Caddo Lake (TU 405).

Pseudemys concinna (24) -- TX: Camp: (Dixon1987); Ferndale Lake, ca 11 km SW Pittsburg,04/05/1979 (UTA R-3OO28, 30039, nv);Franklin: 1.3 mi. S PurIey on Hwy. 37,06/25/1985 (TCWC 63650). Gregg: 10 mi.S Longview, 03/19/1960 (TCWC 13965).Harrison: Caddo Lake, 04/15/1960 (TCWC13966); Caddo Lake, Big Cypress Bayou,

1995 HARDY: AMPHIBIANS AND REPTILES OF CADDO LAKE 17

IJ

1J

06/02/1984 (TCWC 62632); Big Cypre$sCreek, vicinity of Caddo Lake State Park,03/18/1992 (UTA R-32727-32728); CaddoLake, 04/21/1962 (TCWC 18464). Marion:(Dixon 1987); 1 mi. WNW Hwy. 43 bridge onN side Big Cypress Bayou, 06/02/1984 (TCWCS-490, nv). Titus: US Hwy 67, 0.3 km W jctFM 1734, 06/11/1991 (UTA R-30197);Monticello Lignite Mining Area (TX Utilities)pond AR-32, 10/12/1992 (UTA R-34117, nv).LA: Caddo: 2 mi. W jct. LA 1 and LA2 onLA 2, 05/31/1967 (LSUMZ 41112); CaddoLake, 06/28/1947 (TU 632-633); Caddo Lake,06/30/1947 (TU 647, 657-658, 661-662, 677,734-735); 3.7 mi. E Texas line on LA 2,05/30/1968 (LSUMZ 41104).

Terrapene carolina (9) -- TX: Camp: 1 mi. Wfrom Camp county line on Hwy. 557,04/21/1962 (TCWC S-500). Harrison: 6 mi.S are City, 04/27/1963 (SM 6143); CaddoState Park, 04/21/1962 (TCWC 18433); 4 mi.N Uncertain, 04/21/1962 (TCWC 18435);Karnack,Caddo Lake State Pk., 06/18/1952(TNHC 17575); 6 mi. N of Karnack,04/23/1950 (TNHC 9724). Marion: 8 mi. SEJefferson, 04/27/1963 (SM 6104); 0.3 mi. WJet. 729 & 155 on 729, 06/04/1984 (TCWC62641); (Dixon 1987). Titus: 3 mi. E ofMount Pleasant, 05/01/1969 (SM 8874); ca. 5km NE Winfield,TX Utilities Mining Land(Winfield Nor), 10/03/1990 (UTA R-28186, );LA: Caddo: Caddo Parrish, Louisiana,07/07/1967 (NLU 19315).

Terrapene ornata (4) -- TX: Camp: (Dixon1987). Harrison: are City (SM [BCB 6143,nv]). Hopkins: 1-30, 2.7 mi. E jct FM 275,06/13/1993 (UTA R-35342). Marion: (Dixon1987); Jefferson (SM [BCB 6104, nv]). Titus:FM 1734, 0.5 km N jct County Rd NW 13,05/18/1991 (UTA R-30144).

Trachemys scripta (157) -- TX: Marrill Lake,06/14/1939 (SM 6729, nv); Lake Caddo,OS/27/1920 (AMNH R-16968, 16982-16983).Camp: 11 km SW Pittsburg, Ferndale Lake06/04/1979 (UTA R-22598-22605, nv);

Ferndale Lake, ca 11 km SW Pittsburg04/05/1979 (UTA R-30038, nv); Cason,06/26/1971 (UTA R-30668, nv). Gregg: 9mi. N Longview, 04/14/1960 (TCWC 13972).Harrison: Caddo Lake, 04/20/1962 (TCWC18451-18453); Caddo Lake, Big Cypress Bayou06/02/1984 (TCWC 62718, 62740-62742);Caddo Lake State Park, Mill Pond (TCWC68554, nv); Mill Pond, Caddo Lake State Pk.,32-4132N,94-1O-45N, . 07/14/1988 (TNHC50867-508, 50862-50863); Caddo Lake, BigCypress Bayou, 06/02/1984 (TCWC 62734-62739); N of Marshall, vincinity of Caddo LakeSt. Pk., 04/20/1962 (SM 4311); Big CypressCreek, Vicinity of Caddo Lake State Park,03/18/1992 (UTA R-32684); Lake Caddo(Brown 1950). Hopkins: 1.0 mi. SW HarpersCrossing, 07/20/1975 (UTA R-30563, nv); 1.0mi.SW Harpers Crossing, archeological dig,07/19/1975 (UTA R-30645, nv); 1.0 mi. SWHarpers Crossing, archeological dig,07/20/1975 (UTA R-30646, nv). Marion:(Dixon 1987); Caddo Lake, Cypress Bayou &Carter Lake, 06/02/1984 (TCWC 62729-62730); N of Uncertain, Wend Caddo Lake,(UTA R-17066-17067); Cypress Bayou, 5 mi.SE of Jefferson, 04/27/1963 (SM 3511).Morris: Dangerfield St. Park, 06/26/1984(TCWC 62717). Titus: White Oak Cr. onHwy. 271, 06/02/1984 (TCWC 62728); TxUtilities Monticello Lignite mining site,Winfield N, 04/11/1991 (UTA R-32703). LA:Caddo: Caddo Lake, 06/25/1947 (TU 1488-1489, 1491, 1495-1496, 1498, 1500, 1503,1505, 1507, 1512, 1516-1517, 1520, 1529,1531-1532, 1538, 1547, 1569-1572, 1582-1584); Caddo Lake, 06/27/1947 (TU 508, nv,509-511, 513-520, 523-533, 534-535, nv, 536-538, 539, nv, 540); Caddo Lake, 06/30/1947(TV 625,648-456, 663-668, 671-676, 688-697,707, 717-733); Caddo Lake, 06/28/1947 (TV638-640); Caddo Lake, 06/26/1947 (TU 463-464).

TRIONYCIllDAE

Apalone mutica (0) -- Conant and Collins (1991).


Apalone spinifera (25) -- TX: Titus: White OakCr., Hwy. 271, 06/25/1984 (TCWC 62703).LA: Caddo: Caddo Lake, 06/27/1947 (TU469-472, 474-480, 483-489); Vivian BlackBayou, 05/16/1960 (NWSU 482); Caddo Lake,06/25/1947 (TU 381,397-399).

Order SquamataSuborder SauriaIGUANIDAE

Anolis carolinensis (49) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16973). Camp:(Dixon 1987); Logging rd. 1 mi. S Pittsburg,off Hwy. 271, 06/04/1984 (TCWC 62902); 4mi. W of County line on Hwy. 157,06/04/1984 (TCWC 62903). Cass: 5 mi. Wof Atlanta on Hwy 59, 04/10/1971 (LTU40110). Gregg: 15 mi. N Kilgore, 10/10/1959(SFA 899). Harrison: 2.3 mi. N & 1.3 mi.W of Uncertain, 04/16/1973 (FWM 8500); 2.6mi. N, 2.0 mi. W Elysian Fields, 04/19/1973(FWM 8607-8608); 20 mi. NE Marshall,Caddo Lake State Park, 03/25/1951 (LSUMZ[MJF 354], ny); Caddo Lake State Park, (SM[BCB 3444-3445], ny); 5 mi. E of Marshall,04/26/1975 (SM 11796 ny, 11800, ny); CaddoLake, (SM 2713-2715); 2.5 mi. E of CaddoLake State Park, 04/22/1962 (SM 3444); 4.5mi. E of Caddo Lake State Park, 04/21/1962(SM 3445); Caddo Pk., 04/27/1963 (SM 5958);6 mi. N Karnack, 04/29/1955 (TCWC 15199);H. R. Taylor house, Harleton, 03/17/1962(TCWC 17989-17991); Caddo Lake State Park,06/02/1984 (TCWC 62904); Karnack, CaddoLake State Park, 04/21/1962 (TNHC 17582);Caddo Lake State Park, 04/21/1962 (TNHC28421-28423); Caddo Lake State Park, 1.6 mi.of Karnack, 04/21/1962 (TNHC 33357-33358);16 mi. SE Jefferson, 04/01/1934 (UMMZ78214(2»; Lake Caddo (Brown 1950).Marion: (Dixon 1987); Caddo Lake (SM [BCB3446], ny); Caddo Lake, mouth of Doggarty Lnto Cypress Slough, 06/02/1984 (TCWC 62900-62901); 16 mi. SE Jefferson (Brown 1950).Morris: 1 mi. N Omaha, Titus Co.,04/19/1957 (TCWC 15196). Upshur: Circle D

Ranch, 4 mi. NE of Gilmer, 04/08/1974 (SM11785); Lake O'The Pines, 20 mi. NE ofGilmer, 04/27/1963 (SM 5955); Little CypressCreek, 2.2 mi. NE of Gilmer, 03/11/1967 (SM8395). Wood: 4 mi. N. E. Mineola,06/11/1960 (SFA 1225); LA: Kleinpeter,(AMNH R-57980). Caddo: Kneipp's camp,Caddo Lake, near Trees, La., 04/08/1966(NLU 10200-10202, ny); 0.3 mi. W LA 1 onMcCloud-State line hwy, 03/11/1966 (LSUMZ41450); Vivian, (SM, [BCB 630, 643], ny); 1mi. east of Mooringsport, 05/03/1952 (TCWC8901-8902).

Phrynosoma comutum (0) -- Conant and Collins(1991).

Sceloporus olivaceus (0) -- TX:Greeg: (Dixon1987). Harrison: Marshall, near Caddo Lake(Axtell 1992).

Sceloporus undulatus (93) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16972); 5 mi. NJefferson, 05/02/1931 (UOMZ 12534). Camp:(Dixon 1987); 1 mi. W on Hwy. 557 fromcounty line, 06/04/1984 (TCWC 62970). Cass:5 mi. W. of Atlanta on Hwy 59, 04/10/1971(LTU 40105); Woods off Hwy 59, 1/2 mi. S.of Black Cypress, 11/09/1974 (NLU 37907,ny); Daingerfield St. Park (SM [BCB 4725-4726, ny); Hughes Springs (SM [BCB 1211,ny, 1338, ny, 1343, ny]); 9 mi. N HughSprings, 04/23/1950 (SM 1211); 15 mi. NHughes Springs, 04/23/1950 (SM 1338, 1345-1346); 16 mi. N Hughes Springs, 04/23/1950(SM 1343); 4 mi. W Atlanta, 04/15/1960 (SM3188, 3352); Dangerfield State PK.,04/27/1963 (SM 4725-4726); 8 mi. N Linden(Burt 1937). Gregg: 15 mi. N Kilgore,10/17/1959 (SFA 902); 1.5 mi. SE Greggton,02/14/1937 (SM 6625, ny); Harrison: 2.3 mi.N & 1.3 mi. W of Uncertain, 04/16/1973(FWM 8501-8514); Caddo St. Park (SM [BCB5071, ny, 5557, ny]); Caddo Lake (SM 2720-2724, 2726-2729); Caddo Lake (SM [BCB2720-2724, ny, 2726-2729, ny]); 1.6 mi. NE,Karnack, Caddo Lake State Pk., 04/21/1962(TNHC 33469-33470); N of Marshall,

1995 HARDY; AMPHIDIANS AND REPTILES OF CADDO LAKE 19

04/20/1962 (SM 2847, 3129-3130, 3276); 4.5Caddo, OS/27/1920 (AMNH R-16973). Camp:(Dixon 1987); Logging rd. 1 mi. S Pittsburg,off Hwy. 271, 06/04/1984 (TCWC 62902); 4mi. W of County line on Hwy. 157,06/04/1984 (TCWC 62903). Cass: 5 mi. Wof Atlanta on Hwy 59, 04/10/1971 (LTU40110). Gregg: 15 mi. N Kilgore, 10/10/1959(SFA 899). Harrison: 2.3 mi. N & 1.3 mi.W of Uncertain, 04/16/1973 (FWM 8500); 2.6mi. N, 2.0 mi. W Elysian Fields, 04/19/1973(FWM 8607-8608); 20 mi. NE Marshall,Caddo Lake State Park, 03/25/1951 (LSUMZ[MJF 354], ny); Caddo Lake State Park, (SM[BCB 3444-3445], ny); 5 mi. E of Marshall,04/26/1975 (SM 11796 ny, 11800, ny); CaddoLake, (SM 2713-2715); 2.5 mi. E of CaddoLake St. Pk., 04/22/1962 (SM 3444); 4.5 mi. Eof Caddo Lake St. Pk., 04/21/1962 (SM 3445);Caddo Pk., 04/27/1963 (SM 5958); 6 mi. NKarnack, 04/29/1955 (TCWC 15199); H. R.Taylor house, Harleton, 03/17/1962 (TCWC17989-17991); Caddo Lake State Park,06/02/1984 (TCWC 62904); Karnack, CaddoLake State Park, 04/21/1962 (TNHC 17582);Caddo Lake State Park, 04/21/1962 (TNHC28421-28423); Caddo Lake State Park, 1.6 mi.of Karnack, 04/21/1962 (TNHC 33357-33358);16 mi. SE Jefferson, 04/01/1934 (UMMZ78214(2»; Lake Caddo (Brown 1950).Marion: (Dixon 1987); Caddo Lake (SM [BCB3446], ny); Caddo Lake, mouth of Doggarty Lnto Cypress Slough, 06/02/1984 (TCWC 62900-62901); 16 mi. SE Jefferson (Brown 1950).Morris: 1 mi. N Omaha, Titus Co.,04/19/1957 (TCWC 15196). Upshur: Circle DRanch, 4 mi. NE of Gilmer, 04/08/1974 (SM11785); Lake O'The Pines, 20 mi. NE ofGilmer, 04/27/1963 (SM 5955); Little CypressCreek, 2.2 mi. NE of Gilmer, 03/11/1967 (SM8395). Wood: 4 mi. N. E. Mineola,06/11/1960 (SFA 1225); LA: Kleinpeter,(AMNH R-57980). Caddo: Kneipp's camp,Caddo Lake, near Trees, La., 04/08/1966(NLU 10200-10202, ny); 0.3 mi. W LA 1 onMcCloud-State line hwy, 03/11/1966 (LSUMZ41450); Vivian, (SM, [BCB 630, 643], ny); 1mi. east of Mooringsport, 05/03/1952 (TCWC

8901-8902).

Phrynosoma cornuturn (0) -- Conant and Collins(1991).

Sceloporus o/ivaceus (0) -- TX:Greeg: (Dixon1987). Harrison: Marshall, near Caddo Lake(Axtell 1992).

Sceloporus undulatus (93) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16972); 5 mi. NJefferson, 05/02/1931 (UOMZ 12534).Camp: (Dixon 1987); 1 mi. Won Hwy. 557from co. line, 06/04/1984 (TCWC 62970).Cass: 5 mi. W. Atlanta on Hwy 59,04/10/1971 (LTU 40105); Woods off Hwy 59,1/2 mi. S. Black Cypress, 11/09/1974 (NLU37907, ny); Daingerfield St. Park (SM [BCB4725-4726, ny); Hughes Springs (SM [BCB1211, ny, 1338, ny, 1343, ny]); 9 mi. NHugh Springs, 04/23/1950 (SM 1211); 15 mi.N Hughes Springs, 04/23/1950 (SM 1338,1345-1346); 16 mi N Hughes Springs,04/23/1950 (SM 1343); 4 mi. W Atlanta,04/15/1960 (SM 3188, 3352); DangerfieldState PK., 04/27/1963 (SM 4725-4726); 8 mi.N Linden (Burt 1937). Gregg: 15 mi. NKilgore, 10/17/1959 (SFA 902); 1.5 mi. SEGreggton, 02/14/1937 (SM 6625, ny);Harrison: 2.3 mi. N, 1.3 mi. W ofUncertain, 04/16/1973 (FWM 8501-8514);Caddo St. Pk (SM [BCB 5071, ny, 5557, ny]);Caddo Lake (SM 2720-2724, 2726-2729);Caddo Lake (SM [BCB 2720-2724, ny, 2726-2729, ny]); 1.6 mi. NE, Karnack, Caddo LakeSt. Pk., 04/21/62 (TNHC 33469-33470); NMarshall, 04/20/62 (SM 2847, 3129-3130,3276); 4.5 mi E Caddo Lake St. Pk., 04/21/62(SM 5071); 2.5 mi. E of Caddo Lake St. Pk.,04/22/1962 (SM 5557); 1/4 mi. E Harleton,04/14/6262 (TCWC 18068-18072); CaddoLake St. Pk., Karnack, 06/18/1952 (TNHC17520); 16 mi. SE Jefferson, 04/01/1934(UMMZ 78217); 2 mi. N Marshall,04/22/1961 (UTA R-658-660); Lake Caddo(Brown 1950). Marion: (Dixon 1987); 1.2mi. S, 0.6 mi. W Smithland (8.4 mi. N FM134 via TX 43), 04/18/1973 (FWM 8573-


1995

8574); Little Cypress Bayou, 06/11/1934(UMMZ 77646); 3 mi. N Jefferson (Brown1950). Titus: 1 mi. N Omehe, 04/19/1957(TCWC 14999); Mt. Pleasant (Smith andSanders 1952). Upshur: 3.9 mi. SRhonesboro, 03/11/1967 (SM 12834, ny,12859, ny, 12896, ny, 12913, ny, 12934, nv);3.9 mi. N Rhonesboro, 03/11/1967 (SM 11949,nv); Circle J Ranch, 4 mi. ME of Gilmer,04/08/1974 (SM 11986); Little Cypress Creek,2,2 mi. NE of Gilmer, 03/11/1967 (SM12903); 5 mi. SE Clayton, 04/15/1960 (TCWC14008); 5 mi. SE are City, 04/15/1960(TCWC 14010); 5 mi. E Gilmer (Smith andSanders 1952). LA: Caddo: 3.5 mi. S, 1.5mi. W Vivian (2.9 mi. E Tx line via LA 2),04/18/1973 (FWM 8578); Kneipp's camp,Caddo lake, near trees, La., 04/08/1966 (NLU10276-10279,nv).

TEDDAE

Cnemidophorus gularis (3) -- TX: Harrison:Caddo Lake State Pk., 04/27/1963 (SM 5460,nv); Caddo Lake State Pk., 04/27/1963 (SM7620); 2 mi. N Marshall, OS/26/1961(UTA R-662, nv). Morris: Daingerfield, (FMNH23685); Daingerfield (Brown 1950).

Cnemidophorus sexlineatus (15) -- TX: Cass:Douglasville, (SM [BCB 5309,. 5311, 5329,nv]); Linden, (SM [BCB 5308, nv]); JimsBayou 5 mi. S Douglasville, 06/11/1934(UMMZ 77642 [2 specimens]); 5 mi. W.Atlanta on Hwy 59, 04/10/1971 (LTU 40108).Harrison: 2 mi. N, 1 mi. E Karnack in CaddoState Park, 06/28/1960 (DMNH 3003, nv); S.side Back Lake, 2.3 mi. N, 1.3 mi. WUncertain, 04/16/1973 (FWM 8516); CaddoLake, 07/24/1951 (UTA R-384, nv); 2 mi. NMarshall, OS/26/1961(UTA R-663). Marion:(Dixon 1987); Jefferson, 08/26/1950 (LSUMZ50226); Little Cypress Bayou, 06/11/1934(UMMZ 77643(2»; Black Cypress Cr 1 mi. NJefferson, 06/11/1934 (UMMZ 77644); LittleCypress Bayou (Brown 1950); 1 mi. NJefferson (Brown 1950). Morris: 17 mi. SLone Star, 04/28/1963 (SM 5330). Upshur: 5

mi. E Gilmer (Smith and Sanders 1952). LA:Caddo: Caddo Lake, 06/25/1947 (TU 462).

SCINCIDAE

Eumeces anthracinus (12) -- TX: Camp: (Dixon1987); 4 mi. W county line on Hwy. 557,06/04/1984 (TCWC 62905); Ferndale Lake, 8mi. E Pittsburg (Webb and Packard 1961).Cass: 5 mi. NE Avinger, 04/27/1963 (SM4314). Harrison: 20 mi. NE Marshall,03/25/1951 (LSUMZ [MJF 114, 355], nv); 1/4mi. E Harleton, 04/20/1962 (TCWC 18183-18185); Karnack, Caddo Lake State Pk.,04/21/1952 (TNHC 17607); Caddo Lake StatePk., 1.6 mi. NE of Karnack, 04/21/1952(TNHC 33382-33383). Marion: (Dixon1987). Morris: 2.4 mi. ESE Daingerfield, then1.5 mi. WSW on Park Rd. 17, 03/15/1992(TCWC 69291). Wood: Big Sandy Creek atHwy 154, 04/18/1987 (UTA R-28870).

Eumeces jaciatus (59) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16979-16981).Camp: (Dixon 1987); Ferndale Lake, 8 mi. EPittsburg, 04/07/1951 (FWM 4012); 1 mi. Won Hwy 557 from Camp county line,06/04/1984 (TCWC 62849). Cass: Avinger(SM [BCB 5471, nv]); are City (SM [BCB2133-2135, nv]); 5 mi. NE Avinger, Rt. 155,04/27/1963 (SM 5159, 5162, 5471); 13 mi.NE Avinger, 04/27/1963 (SM 5164). Gregg:9 mi. N Longview, Hwy. 26, 04/13/1960(TCWC 14026); 9 mi. N Longview,04/14/1960 (TCWC 14030). Harrison:Caddo Lake (FMNH 116852, FWM 4413);2.3 mi. N, 1.3 mi. W of Uncertain, S of BackLake, 04/16/1973 (FWM 8518); 20 mi. NEMarshall, Caddo Lake State Park, 03/25/1951(LSUMZ, [MJF 356], nv); Harleton (SM,[BCB 5475, nv]); 4 mi. S of Harleton,04/28/1963 (SM 5158, 5475); Harleton,04/07/1962 (TCWC 18154-18155); 7 mi. EKarnack, 04/21/1962 (TCWC 18157);4 mi. NKarnack, 04/29/1955 (TCWC 20711);Karnack, Caddo Lake State Pk., 06/18/1952(TNHC 17605-17606); Caddo State Park,10/14/1967 (UTA R-1293-1294);Caddo Lake,

1995 HARDY: AMPHmIANS AND REPTILES OF CADDO LAKE 21

(UTA R-239); 15 mi. E of Marshall, CaddoLake Bayou, 06/10/1961 (UTA R-909); LakeCaddo (Brown 1950). Marion: (Dixon 1987);Caddo Lake, Doggerty Cr., mouth of CypressSlough, 06/02/1984 (TCWC 62848); 16 mi.SE Jefferson, 04/01/1934 (UMMZ 78215).Morris: 1.8 mi. SSE of US 259, TX Hwy 11/9jct in Daingerfield, 07/12/1993 (FWM 11058,nv); Lone Star (SM, [BCB 5468, nv]); 17 mi. Sof Lone Star, 04/28/1963 (SM 5468); 1 mi. NOmaha (Titus Co.), 04/19/1957 (TCWC20405); Dangerfield St. Park, 06/26/1984(TCWC 62850); V2 mi. W Cason (Smith andSanders 1952). Upshur: 1 mi. N Ore City,04/23/1950 (SM 2133-2134); Ore City (SM,[BCB 5493, nv]); 3 mi. N Ore City,04/23/1950 (SM 1249); 2 mi. NW Ore City,04/23/1950 (SM 2135); 8 mi. SE Ore City,04/28/1950 (SM 5167); 5 mi. SE Ore City,04/28/1963 (SM 5467); 8 mi. SE Ore City,04/28/1963 (SM 5493); Little Cypress Creek,2.2 mi. NE Gilmer, 03/11/1967 (SM 8513-8514); 5 mi. NE Ore City near Lake O'thePines, 04/15/1960 (TCWC 14027). LA:Caddo: 0.3 mi. W. La 1 on McCloud stateline rd, 03/12/1966 (LSUMZ 32616); CaddoLake (under log) near Mooringsport, La,11/14/1964 (NLU 9232, nv); 1 mi. east ofMooringsport, 05/03/1952 (TCWC 8963, nv);Caddo Lake, 06/24/1947 (TU 338-341); CaddoLake, 06/30/1947 (TU 627,629).

Eumeces laticeps (2) -- TX: Harrison: Karnack,Caddo Lake State Pk., 06/19/1952 (TNHC17603). Titus: White Oak Cr. on Hwy. 271,06/26/1984 (TCWC 62844).

Eumeces septentrionalis (4) -- TX: Cass: Avinger(SM [BCB 4314, nv]); 7 mi. E Hughes Springs(Brown 1951a). Harrison: Harleton, (SM[BCB 5527, nv]); 4 mi. S of Harleton,04/28/1063 (SM 5527). Titus: 3 mi. S Mt.Pleasant (Smith and Sanders 1952). LA:Caddo: 1 mi. east of Mooringsport,05/03/1952 (TCWC 8966).

Scincella laterale (87) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16974-16978); 4 mi.

07/28/1951 NE of Avinger, 04/27/1963 (SM4402). Camp: (Dixon 1987); 8 mi. E ofPittsburg, Ferndale Lake, 04/07/1951 (FWM4215). Cass: Woods 1 mi. N. on rd. 2.5 mi.NE of Avinger on HWY 165, 11/09/1974(NLU 37965-37966, nv); Woods off HWY 591/2 mi. S. of Black Cypress Bayou,11/09/1974 (NLU 37971, nv); 4 mi. NE ofAvinger, 04/27/1963 (SM 4401). Gregg:Greggton (SM [BCB 6532, nv]); 1 mi. SE ofGreggton, 02/14/1937 (SM 6532, nv); 9 mi.N Longview, 04/13/1960 (TCWC 14043-14044); 0.5 mi. W Aledo H.S., 04/01/1961(TCWC 20354, nv). Harrison: Caddo Lake(SM [BCB 2716-2717, nv]); Caddo Park (SM[BCB 4453, nv]); S side Back Lake, 2.3 mi.N, 1.3 mi. W Uncertain, 04/16/1973 (FWM8517, nv); 1.2 mi. NE FM 2198 via dirtroad.S side Back Lake, 04/16/1973 (FWM8521, nv); 2.0 mi. NW TX 43 via FM 134.1.7 mi. N, 2.6 mi. W, Karnak, 04/17/1973(FWM 8546, nv); 3.6 mi. N, 5.7 mi. WKarnack, 04/17/1973 (FWM 8547, nv); 2.6mi. N, 2.0 mi. W Elysian Fields, 04/19/1973(FWM 8609, ny, 8620, nv); 20 mi. NEMarshall, Caddo Lake State Park, 03/25/1951(LSUMZ [MJF 357-359], nv); Jefferson (SM[BCB 4575, nv]); Caddo Lake State Park (SM[BCB 4574, nv]); Caddo Lake (SM 2716-2717); Caddo Lake State Park, Sawmill Pond,04/05/1985 (UTA R-14064-14066); N ofMarshall, 04/20/1962 (SM 3449-3462); CaddoLake State Pk, 04/27/1963 (SM 4453-4454,4574); 3.5 mi. S of Jefferson, 04/27/1963 (SM4575); 6 mi. W Karnack, 04/29/1955 (TCWC15002); 1/4 mi. E Harleton, 04/14/1962(TCWC 18136-18139); 7 mi. E Karnack,04/21/1962 (TCWC 18140); Caddo Lake StatePark, 06/02/1984 (TCWC 62856); Pyburn,06/18/1952 (TNHC 17598); Karnack, CaddoLake State Pk, 06/18/1952 (TNHC 17599);Karnack, Caddo Lake State Pk, 06/18/1952(TNHC 17600); Karnack, Caddo Lake StatePk, 06/18/1952 (TNHC 17601); Caddo LakeState Pk, Karnack, 04/21/1961 (TNHC28410); Caddo Lake State Pk. 1.6 mi. NE ofKarnack, 04/21/1961 (TNHC 29027); CaddoLake State Pk. 1.6 mi. NE of Karnack,


04/21/1961 (TNHC 33364); Caddo Lake StatePk. 1.6 mi. NE of Karnack, 04/21/1961(TNHC 33365); Lake Caddo (Brown 1950).Marion: (Dixon 1987); 3.4 mi. S, 1 mi. WSmithland, 04/18/1973 (FWM 8571, nv); 3.4mi. S, 1 mi. W Smithland, 04/18/1973 (FWM8572, nv); Caddo Lake, Doggerty Cr. mouth toCypress Slough, 06/02/1984 (TCWC 62855); 1mi. S Jefferson, 09/09/1950 (UM-MVP 7498);16 mi. SE Jefferson (Brown 1950). Morris:Daingerfield St. Park, 06/26/1984 (TCWC62862); Daingerfield St. Park, 06/26/1984(TCWC 62863); 1/2 mi. W Cason (Smith andSanders 1952). Titus: 1 mi. N Omaha,04/19/1957 (TCWC 15010); 1 mi. N Omaha,04/19/1957 (TCWC 15011); White Oak Cr. onHwy. 271, 06/26/1984 (TCWC 62858);Monticello Lignite Mining Area (TX Utilities)Winfield N, 04/14/1992 (UTA R-32628). LA:Kleinpeter (AMNH R-57989); Mt. Pleasant(Smith and Sanders 1952). Caddo: CaddoLake, 06/24/1947 (TU 370); 3.5 mi. S, 1.5 mi.W Vivian, 04/18/1973 (FWM 8579, nv); 3.5mi. S, 1.3 mi. W Vivian (2.9 mi. E TX line viaLA 2), 04/18/1973 (FWM 8580, nv); 1 mi. EMooringsport, 05/03/1952 (TCWC 8958);Caddo Lake, 06/23/1947 (TU 316); CaddoLake, 06/24/1947 (TU 337, 5219).

Suborder SerpentesCOLUBRIDAE

Carphophis vermis (1) -- TX: Harrison:Marshall, 10 mi. NE Marshall, 06/18/1952(TNHC 17552, nv).

Cemophora coccinea (3) -- TX: Harrison: 2 mi.E, 1 mi. N Karnack, Caddo State Park,06/28/1960 (DMNH 3971, nv); 2.0 mi. N ofHwy 43 on Hwy 134, 06/20/1977 (UTA R-6682); near Caddo Lake, 4.5 mi. N Karnack(Raun 1974). Marion: (Dixon 1987).Morris: Dangerfield [sic] State Park, (TCWC63015).

Coluber constrictor (16) -- TX: Cass:Douglasville (SM [BCB 1895], nv); 3 mi. NHughes Springs, 04/23/1950 (SM 1896).

Harrison: 2 mi. E, 1 mi. N Karnack, CaddoLake Park, 06/28/1960 (DMNH 2850, nv); 8mi. S of Karnack, 04/29/1955 (TCWC 14927);Harleton High School, Harleton, 04/11/1962(TCWC 18280); 1/4 mi. E of harleton,04/20/1962 (TCWC 18281); Caddo Lake StatePark, 06/02/1984 (TCWC 63044); 3.6 mi. SWof Karnack on Hwy 43, 04/21/1962 (TNHC33927, nv); 2 mi. N Marshall, 10/19/1961(UTA R-419). Marion: (Dixon 1987); 1.2 mi.S, 0.6 mi. W Smithland, 04/18/1973 (FWM8575, nv). Morris: Dangerfield State Park,(TCWC 63045); Omaha, 2 mi. E of Omaha,(TNHC 10106). Titus: Monticello LigniteMining Area, Winfield N Pond AR-32,06/22/1992 (UTA R-32661); 0.25 mi. W of jct271 on FM 1734,05/16/1991 (UTA R-32670);11/2 mi. E Cookville (Smith and Sanders 1952).Upshur: 1 mi. N of Gilmer, 04/23/1950(DMNH 1840, nv). Wood: 18 mi. N. E.Quitman, 06/29/1966 (SFA 2459).

Diadophis punctatus (4) -- TX: Marion: (Dixon1987). Morris: 4 mi. W Omaha on U.S. 67,04/24/1950 (FWM 4163, nv). Wood: CoitRd., (FWM 7495, nv); Coit Rd., (FWM7496, nv). LA: Caddo: 0.3 mi. W. LA 1 onMcCloud-State Line Rd, 04/02/1966 (LSUMZ42737).

Elaphe guttata (0) --Conant and Collins (1991).A recent survey of the geographic distributionof Texas and Louisiana specimens (Raymondand Hardy 1983) demonstrated a conspicuousvoid of this species for the entire watershed.

Elaphe obsoleta (7) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16989). Camp: (Dixon1987). Harrison: Karnack, Caddo Lake StatePk, 06/17/1952 (TNHC 17867). Marion:(Dixon 1987). Morris: 3.2 km WDangerfield, 11/12/1978 (UTA R-14644).Titus: Monticello Lignite mine, Winfield N, Sof jct NW15 & FM 1734, 05/31/1992 (UTAR-32519); county rd nr Winfield N miningarea (TX utilities, 05/16/91 (UTA R-32662);Monticello Lignite Mining Area (TX Utilities)Winfield N, 07/23/92 (UTA R 33945).

1995 HARDY: AMPHffilANS AND REPTILES OF CADDO LAKE 23

Upshur: 14 km NE New Diana via ZeninRoad, 12/04/1987 (UTA R-19432).

Faranda abacura (3) -- TX: Camp: (Dixon1987); Ferndale Lake, 8 mi. E of Pittsburg,04/07/1951 (FWM 4437, nv). Cass: Linden(Brown 1950). Harrison: 0.4 mi. E Leigh(1.5 mi. W FM 9 via FM 1999), 04/18/1973(FWM 8594). LA: Caddo: Rodessa, 1 mi. Sof, 06/10/1935 (USNM 100591). Smith's(1938) map seems to include the watershed buthe gave no specific localities.

Heterodon nasicus (1) -- TX: Titus: at jet Co.Rd. NW 13 and Fm 1734, 05/15/1991 (UTAR-32669).

Heterodon platyrhinos (1) -- TX: Marion: (Dixon1987). Morris: Dangerfield State Park,(TCWC 63041).

Lampropeltis calligaster (2) -- TX: Camp:(Dixon 1987). Harrison: Harelton,04/07/2962 (TCWC 18332). Marion: (Dixon1987). Upshur: 3 mi. SE of Rhonesboro nearFM RD 49, Maxwell Ranch, 03/14/1971(DMNH 2083, nv).

Lampropeltis getula (6) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16990). Harrison: 1/4mi. E of Harleton, 04/20/1962 (TCWC 18336);Harleton, 04/21/1962 (TCWC 18337); LakeCaddo (Brown 1950). MarioYJ: (Dixon1987); Jefferson, 08/26/1950 (LSUMZ22948). Upshur: Little Cypress Creek, 2.2mi. NE of Gilmer, 04/11/1967 (SM 8807).LA: Caddo: Oil city, Caddo lake area,07/04/1972 (NWSU 4276).

Lampropeltis triangulum (2) -- TX: Harrison: 5mi. E of Marshall, 04/12/1975 (SM 12130);Caddo Lake, 13.8 mi. NW of Karnack,04/21/1962 (TCWC 18340).

Masticophis flagellum (16) -- TX: Cass: HughesSprings (SM [BCB 1900, 2120, nv)); 3 mi. Nof Hughes Springs, 04/23/1950 (SM 1900); 18mi. from Hugh Springs, 04/23/1950 (SM 2120,

nv); 1 mi. S of Avinger, 04/23/1963 (SM6158); 6 mi. S, 2.5 mi. W Atlanta on 43,08/21/1970 (TCWC 67043); 5 mi. E HughesSprings on 11, OS/24/1970 (TCWC 67044).Harrison: Smyrna Cemetery, Harleton,03/11/1962 (TCWC 18286, nv); 3.8 mi. NWofWoodlawn, 04/22/1950 (TNHC 9714, nv).Marion: (Dixon 1987); 2.6 mi. S, 10 mi. EJefferson (3.1 mi. N FM134 via TX 43),04/18/1973 (FWM 8570); 1 mi. SE Jefferson,09/09/1950 (UM). Titus: 0.8 mi. N ofjunction FM 1734 on Co. rd. NW 15,10/22/1991 (UTA R-32657). Upshur: Gilmer,04/17/1938 (SM 6511); Ore City (SM [BCB687, nv)); 3 mi. N of Ore City, 04/23/1950(SM 687); Gilmer (Brown 1950). Wood:Golden, 11/08/1964 (UTA R-2089).

Masticophis taeniatus (1) -- TX: Camp Bullis,07/01/1922 (AMNH R-22743).

Nerodia cyciopion (7) -- TX: Harrison: % mi.NE of Caddo Lake, Karnack, 08/01/1950(TNHC 11916). Marion: (Dixon 1987); N ofUncertain, Wend Caddo Lake (UTA R-15879);Near Uncertain (Caddo Lake) (UTA R-17976-17977). LA: Caddo: Caddo Lake,06/25/1947 (TU 390,391, ny, 401, nv).

Nerodia erythrogaster (13) -- TX: Camp: (Dixon1987). Cass: 4 mi. W of Atlanta, 04/30/1960(SM 3035). Franklin: 0.8 mi. W of jet co. rd4145 on FM 21, 05/13/1991 (UTA R-32644).Harrison: Caddo State Park, 06/29/1961(DMNH 4078, nv); Karnack, Caddo Lake StatePk., 06/18/1952 (TNHC 17569 [2 specimens] -17570). Hopkins: Peerless, 06/10/1975 (UTAR-30493). Marion: (Dixon 1987). Titus:TX Utilities MonticelloLignite Mining SiteWinfield North, 04/12/1991 (UTA R-32680);TX Utilities Monticello Lignite Mining area,Winfield Mining, 01/17/1993 (UTA R-34626-34627); TX Utilities Monticello Lignite Miningarea, Winfield North" 10/24/1991 (UTA R-34691); 0.7 rd mi. S of jet Co Rd NW 3 on CoRd NW 18, 09/24/1992 (UTA R-35033).Upshur: Lake 0' the Pines, 14 mi. NE ofGilmer, 04/26/1963 (SM 4987).

24HARDY: AMPHffiIANS AND REPTILES OF CADDO LAKE

1995

Nerodia fasdata (12) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16988). Camp:(Dixon 1987). Gregg: Warren City,05/11/1957 (TCWC 15380). Harrison:Harelton, 04/21/1962 (TCWC 18235);Karnack, Caddo Lake State Pk., 06/18/1952(TNHC 17566, 17567 [2 specimens], 17568);Caddo Lake St. Pk. (TNHC 26752); CaddoLake State Pk., 04/22/1962 (TNHC 28401 [2specimens]); 5 mi. E of Karnack, 04/08/1955(TCWC 15374); Lake Caddo (Brown 1950).Titus: TX Utilities Monticello Lignite miningarea Winfield North, 05/12/1992 (UTA R-32518); 0.2 mi. W of jct co. rd. NW 15 on Co.Rd NW 18,04/13/1992 (UTA R-32668). LA:Caddo: Caddo Lake, 06/26/1947 (TU 465).

Nerodia rhombifer (25) -- TX: Lake Caddo,OS/27/1920 (AMNH R-16987). Cass: 4 mi. SLinden, 06/10/1949 (UMMZ 99744).Harrison: 8.1 mi. NW Karnack, 04/21/1962(TCWC 18229); 3 mi. W of Karnack,06/18/1952 (TNHC 17560-17563); Lake Caddo(Brown 1950). Marion: (Dixon 1987); LakeO'the Pines, 8 mi. N Harleton, 04/07/1962(TCWC 18222). Morris: Dangerfield StatePark, 06/01/1983 (TCWC 63003). Titus: TXUtilities Monticello Lignite Mining area,Winfield Mining, 01/17/1993 (UTA R-34625);1 mi. N of Omobe (TCWC 14536); 0.5 mi. Sjct Co. rd. NW 4 and US 271, 04/13/1992(UTA R-32635); Monticello Lignite miningarea, Winfield N site, pond AR-32, 05/15/1991(UTA R-32663); 0.7 mi. S of jct CR NW 13 onFM 1734,06/18/1991 (UTA R-32676); 0.6 mi.N of jct CR NW 6 on CR NW 13, 04/13/1992(UTA R-32677); 0.1 mi. N ofjct NW 6 on NW13, 04/13/1992 (UTA R-32678); 0.6 mi. N ofjct FM 1734 on CR NW 13, 05/14/1991 (UTAR-32679, nv); Monticello Lignite Mining area(TX Utilities) pond AR15, 09/28/1992 (UTAR-33953); Monticello Lignite Mining area (TXUtilities) pond AR40, 11/04/1992 (UTA R-33954); TX Utilities Monticello Lignite Miningarea, Winfield Mining, 01/17/1993 (UTA R-34623-34624, 34650); TU Monticello Lignitemining area, Winfield North, ca. 50 m W pondAR 29, 11/18/1992 (UTA R-34698). Upshur:

9.8 mi. S ofjunct. ofHwy 155 & Hwy 271, onHwy 271, 06/10/1084 (TCWC 63010). LA:Caddo: Caddo Lake, 06/25/1947 (TU 393).

Opheodrys aestivus (10) -- TX: Harrison: 2 mi.N, 1 mi. E Karnack in Caddo State Park,06/27/1960 (DMNH 3077, nv); Caddo StatePark (SM [BCB4196], nv); 4.5 mi. E of CaddoState Park, 04/21/1962 (SM 4195). Hopkins:1 mi. E Posey, 06/14/1975 (UTA R-30395).Morris: Daingerfield, (FMNH 23686; Brown1950). Titus: 1.1 mi. N of junction FM 1734on Co. rd. NW 15, 10/22/1991 (UTA R-32631); 0.2 mi. W of jct co. rd. NW 17 on co.rd. NW 18, 05/15/1991 (UTA R-32666); 0.3mi. W of Co. Rd. NW 15 on Co. Rd. NW 6,05/16/1991 (UTA R-32667); 0.3 rd mi. N ofjct FM 1734 on Co rd NW 15, 10/25/1992(UTA R-33950). LA: Caddo: Caddo Dam,05/05/1969 (LSUS 1541).

Regina grahami (3) -- LA: Caddo: Oil City,Wade St., 05/13/1981 (LSUS 5245); 2.4 mi. E,0.5 mi. S Mooringsport, 04/23/1986 (LSUS5344); Caddo Lake, 06/20/1947 (TU 624).

Regina rigida (3) -- TX: Cass: Jims Bayou 5mi. S Douglasville, 06/11/1934 (UMMZ77656). Harrison: 1.6 mi. NE , Karnack,Caddo Lake State Pk., 04/21/1962 (TNHC29029). Marion: (Dixon 1987). LA:Caddo: 3.5 mi. S, 1.5 mi. W Vivian,04/18/1973 (FWM 8581, nv).

Storeria dekayi (19) -- TX: Hare1ton,04/14/1962 (TCWC 18244). Camp: (Dixon1987); Ferndale Lake, 8 mi. E of Pittsburg,04/07/1951 (FWM 4271, nv). Cass: 2.3 mi.SE of Linden on Hwy 125, 11/09/1974 (NLU37903, nv); 3.1 mi. SE of Linden on Hwy125, 11/09/1974 (NLU 37906, nv); 7 mi. SWof Linden, 04/27/1963 (SM 4740). Gregg: 9mi. N of Longview, 04/14/1960 (TCWC14094). Harrison: 1 mi. NW Karnack, FRRd. 134, 03/25/1951 (LSUMZ); 1 mi. NWKarnack, FR 134, 03/24/1951 (LSUMZ); 1/4mi. E of Harelton, 04/14/1962 (TCWC18243). Titus: 0.3 mi. W of jct CR NW 15

1995 HARDY: AMPHffiIANS AND REPTILES OF CADDO LAKE 25

on CR NW 6,03/04/1992 (UTA R-32686); 0.2mi. N of jct FM 1734 on Co. rd. NW 15,04/13/1992 (UTA R-32634); 0.3 mi. N of JctFM 1734 on CR NW 15,03/04/1992 (UTA R-32685); 1.0 mi. N of jct CR NW 6 on CR NW15, 03/04/1992 (UTA R-32687); TUMonticello Lignite Mining Area, Winfield Nwoods around, 09/23/1991 (UTA R-32688).Upshur: 6 mi. N of Winona, 03/11/1967 (SM9081, nv). LA: Caddo: 1 mi. S (by rd.)Mooringsport, 07/01/1970 (LSUS 1608); 1 mi.NW Vivian, 09/27/1975 (LSUS 3495); 2 blocksE Latex state line road, 04/17/1974 (LSUS3516); 1616 N. Pine Street, Vivian, 07/01/1992(LSUS 5676).

Storeria occipitomaculata (7) -- TX: Cass: Woods1 mi. N. on dirt rd. 2.5 mi. NE of Avinger onHwy 155, 11/09/1974 (NLU 27924,nv). Harrison: Karnack,Caddo Lake StatePk., 06/17/1852 (TNHC 17556-17557). Morris: 2.4 mi. ESE Daingerfield,then 1.5 mi. WSW on Park Rd. 17,03/15/1992(TCWC 69301). Wood: roadside park on FM2088,6 km W FM 2869, 04/18/1987 (UTA R-28904). LA: Caddo: 7 mi. SSW (by LA.169) Mooringsport, 07/01/1970 (LSUS 933);Caddo: 3 mi. E (via Blanchard-Latex Road)Texas-Louisiana St. Line, 07/28/1970 (LSUS936).

Tantilla gracilis (12) -- TX: Cass: 5 mi. NEAvinger, 04/27/1950 (SM 5088); 17 mi. N ofHugh Springs, 04/23/1950 (SM 1521, nv); 6mi. SW of Linden, 04/27/1963 (SM 5657);Hughes Springs (SM [BCB 1521, nv]); OreCity (SM [BCB 1522, nv]). Harrison: 7 mi.SE Jefferson, 04/21/1950 (DMNH 1843, nv);Karnack,Caddo Lake State Pk., 06/18/1952(TNHC 17551, yy); 16 mi. SE Jefferson,04/01/1934 (UMMZ 78229 [this specimen wascited by Kim, et al. 1949 from Marion Countybut the location is actually in HarrisonCounty]). Marion: (Dixon 1987); 3 mi. NJefferson (Burt 1936b; Brown 1950). Upshur:GeImer [sic, Gilmer], (SM [BCB 1523, nv]);Gilmer 3 mi. N Jefferson (Burt 1936b); 1 mi.N of Ore City, 04/23/1950 (SM 1522); 2 mi. N

of Gilmer, 04/21/1950 (SM 1523); Gilmer(Brown 1950). LA: Caddo: Caddo Lake,06/23/1947 (TU 10271).

Thamnophis proximus (14) -- LA: 1 mi. NKarnack, 04/29/1955 (TCWC 14911). Cass:Jims Bayou 5 mi. S Douglasville, 06/11/1934(UMMZ 77659). Harrison: 1/4 mi. E ofHarelton, 04/14/1962 (TCWC 18266); S. sideof Back Lake, 2.3 mi. N, 1.3 mi. WUncertain, 04/16/1973 (FWM 8519); 4 mi. NEof Karnack, 06/19/1952 (TNHC 17548-17549);2 mi. N, 1 mi. E Karnack in Caddo State Park,06/28/1960 (DMNH 3053, nv ); 7 mls SE ofJefferson, 04/21/1950 (DMNH 1822, nv);Caddo Lake, 04/15/1960 (TCWC 14105).Marion: (Dixon 1987). Titus: 2 mi. E Mt.Pleasant 05/02/1931 (UMMZ 70372); TXUtilities Monticello Lignite Mining area,Winfield North, 01/17/1993 (UTA R-34651-34652). Upshur: Little Cypress Creek, 2.2 mi.NE of Gilmer, 04/11/1967 (SM 10727). LA:Caddo: Caddo Dam site, 05/07/1969 (LSUS1660).

Tropidoclonion lineatum (7) -- TX: Harrison: 16mi. SE Jefferson, 04/01/1934 (UMMZ 78230[5 specimens]). Marion: (Dixon 1987);04/01/1934 (UMMZ 174460); 04/01/1934(UMMZ 174461); 16 mi. SE Jefferson (Brown1950).

Virginia striatula (25) -- TX: Camp: (Dixon1987); Ferndale Lake, 8 mi. E of Pittsburg,04/07/1951 (FWM 4326-4327, nv). Cass:Hughes Springs (SM [BCB 1703, nv]). 2 mi.N of Hugh Springs, 04/23/1950 (SM 1703).Harrison: Caddo Lake (SM 2742, SM [BCB2742, nv]). 1 mi. N of Karnack, 04/29/1955(TCWC 15230); Karnack, 04/29/1955 (TCWC15229); 3.0 mi. W of Uncertain, 03/19/1992(UTA R-32682); Harleton, 03/17/1962 (KU133594-133595); 0.5 mi. E Harleton,03/20/1962 (KU 133596-133597); 5 mi. NWKarnack (KU 133575-133576). Hopkins:Club Lake, Greenville, 10/13/1952 (DMNH3104, nv). Marion: (Dixon 1987); BigCypress Bayou at U.S. 59 in Jefferson,


04/17/1973 (FWM 8553-8555); 12.2 mi. SLinden (Smith and Sanders 1952). Morris:Douglasville (SM [BCB 3667, nv]); lh mi. WCason (Smith and Sanders 1952). Upshur:Gilmer (SM [BCB 1570, nv]); 8 mi. W ofGilmer, 04/23/1950 (SM 1570, 1587); Gilmer(Brown 1950). LA: Caddo: 0.3 mi. W LA 1on McCloud- State Line Hwy, 03/12/1966(LSUMZ 41449); 7.8 mi. E Leigh (HarrisonCo., TX), 04/17/1973 (FWM 8567).

Virginia valerea (0)-- Conant and Collins (1991).

VIPERIDAE

Agkistrodon contortrix (16) -- TX: Camp: 2.4km. N Leesburg, 05/05/1987 (TCWC 66172,nv). Harrison: NW Karnak via FM 134. 1.8mi. N, 2.8 mi. W Karnak, 04/16/1973 (FWM8538); 2.6 mi. N, 2.0 mi. W Elysian Fields,04/19/1973 (FWM 8610); 3 mi. NE ofKarnack, 04/21/1962 (TCWC 17395);Karnack,caddo Lake State Park, 06/19/1952 (TNHC17571); Caddo Lake State Park, 1.6 mi. NEKarnack, 04/21/1952 (TNHC 33883).Marion: (Dixon 1987); 1.2 mi. S, 0.6 mi. WSmithland, 04/18/1973 (FWM 8576-8577).Titus: Mount Pleasant, 06/02/1935 (SM6063); 0.2 rd mi. E of jct Co rd NW 17 on Cord NW 18,07/30/1991 (UTA R-32299); 0.8 rdmi. W of jct Co rd NW 15 on Co rd NW 18,09/24/1992 (UTA R-33952). Upshur: LittleCypress Creek, 2.2 mi. NE of Gilmer,04/11/1967 (SM 7982). Wood: FM 312, 0.9mi. S jct SH 154,04/10/1990 (UTA R-32200:).LA: Gentilly, (AMNH R-121380). Caddo:5.5 mi. S, 1.5 mi. W Mooringsport,04/18/1973 (FWM 8591); Caddo Lake,04/05/1969 (LTU 5383).

Agkistrodon piscivorus (19) -- TX: White OakCreek, 06/25/1984 (TCWC 63368). Camp:(Dixon 1987); Ferndale Lake, 8 mi. EPittsburg, 04/07/1951 (FWM 4165, nv).Cass: Dangerfield, 08/21/1971 (LTU 40102);Jims Bayou 5 mi. S Douglasville, 06/11/1934(UMMZ 77689). Harrison: 1.1 mi. E Leigh,0.7 mi. E FM 9 via FM 1999, 04/18/1973

(FWM 8593); Lake Caddo, OS/27/1920-06/03/1920 (AMNH R-16991-16992); CaddoLake, (SM 2710); Caddo Lake (SM [BCB2710, nv]); Karnack, Caddo Lake State Park,06/18/1952 (TNHC 17537); 1.0 mi. N ofMarshall, behindradio KMHTbroadcast tower,10/22/1961 (UTA R-I017); Lake Caddo(Brown 1950). Marion: (Dixon 1987); 16 mi.SEJefferson(Brown 1950). Titus: 3 mi. E ofjct Co. Rd NW 15 on FM 1734, 10/23/1991(UTA R-32630); TX Utilities MonticelloLignite Mining area, Winfield N" 06/19/1992(UTA R-34124). Upshur: Gilmer, (UTA R-17008). Wood: 17mi. N 7 mi. E Hwy 154&312, 09/06/1965 (DMNH 2200, nv). LA:Caddo: Caddo Lake, 06/25/1947 (TU 382);Caddo Lake, 06/30/1947 (TU 630); CaddoLake, 06/30/1947 (TU 631, nv).

Crotalus horridus (1) -- TX: Cass: ca. 3 km SMarietta, on oil lease, 05/04/1987 (UTA R-30066, nv).

Sistrurus miliarius (9) -- TX: Harrison: 14.5 kmNE Marshall [9 mi. NE Marshall on tag] (UTAR-14615). Marion: (Dixon 1987); StallcupHome Place 7 1/2 Mi. NW Smithland,11/14/1965 (DMNH 1566, nv); Caddo Lake,(DMNH 3887, nv); N of Jefferson, 09/08/1984(UTA R-15670). Morris: 2.4 mi. SE of USHwy 259, TX Hwy 11/49 jct in Daingerfield,07/12/1993 (FWM 11051, nv); DangerfieldState Park (TCWC 62982, nv). Wood: 7.0mi. E Winnsboro (TCWC 52045); Holly LakeRanch, 10/01/1984 (UTA R-15669). LA:Caddo: 5.3-5.5 Mi. N jct. Blanchard-Furrh&Greenwood-Mooringsport rds., 09/22/1976(LSUS4021).

ELAPIDAE

Micrurus fulvius (5) -- TX: Upshur: 14 km NLognview, off St Hwy 300, Glenwood Acres,10/08/1984 (UTA R-15725, 15734).Harrison: 3.5 mi. E Clayton, 12/09/1966(NLU 13372); 1.6 mi. NE, Karnack, CaddoState Park, 04/21/1962 (TNHC 29036).Morris: Daingerfield (FMNH 23687; Brown

1995HARDY: AMPHffilANS AND REPTILES OF CADDO LAKE 27

1950).

Order CrocodiliaCROCODYLIDAE

Alligator mississippiensis (2) -- TX: Caddo Lake(TCWC 21400-21401). Camp: (Dixon1987). Marion: (Dixon 1987).

DISCUSSION

The objective of this project was to summarizethe known herpetofauna of the Caddo Lakedrainage system based on surveys of the literatureand museum collections. While neither surveysare complete they are thorough enough to presenta reasonably accurate list of the herpetofauna and,of more importance, they assist in identifying rarespecies or those that have changed in localabundance or in geographic distribution during thepast century.

A total of 1748 specimens have been located inthe following 20 collections, ranked by decreasingholdings.

SM 364UTA 279TU 261TCWC 238TNHC 206FWMSH 155AMNH 51NLU 49DMNH 37LSUMZ 32UMMZ 18LSUS 14UM 10KU 9LTU 8NWSU 7SFA 4FMNH 4OU 1USNM 1

Eighty five species have been confirmed,

including 27 (32 %) species of amphibians and 58(68%) species of reptiles. Frogs comprise 56% ofthe amphibian fauna and 18 % of the totalherpetofauna. Snakes make up the largest reptilegroup with 32 (60%) species, followed by 14(24%) species of turtles, 11 (19%) species oflizards, and the alligator. Snakes comprise 38 %of the total herpetofauna.

The following four species have knowngeographic distributions that definitely include allor part of the watershed and thus are probablypresent; however, no specimens nor literaturecitations have been found that cite specificlocations within the watershed:

Rana areolataApalone muticaPhrynosoma comutumVirginia valerea

Another four species have publisheddistributions that closely approximate thewatershed. Eventually, the following species willprobably be documented in the watershed unlesstheir geographic distributions are presentlyshrinking, unknown to biologists:

Pseudacris clarkiGastrophryne olivaceaBufo vallicepsElaphe guttata

Including these eight species as likely membersof the herpetofauna, the total number of species ineach group is: 12 salamanders, 19 frogs (31amphibians), 15 turtles, 12 lizards, 34 snakes and1 crocodilian (62 reptiles) for a total of 93.

SPECIES NOT VERIFIED

The following species are expected in thewatershed because of the known geographicdistribution; however, no specimens nor literaturerecords are known for specific localities.

Pseudacris clarkiGastrophryne olivaceaBufo valliceps


Rana areolataApalone muticaPhrynosoma comutumVirginia valereaElaphe guttata

SPECIES EXCLUDED

Bufo speciosus has been reported from thewatershed; however, it is excluded from the faunain this study until more supporting evidence ispresented. The single specimen is from TX:Harrison: 2 mi. N 1 mi. E Karnack, Caddo StatePark (DMNH 2482,nv). This specimen has beentransferred to the University of Texas at Arlingtonand was unavailable for examination. The localityis well outside of the previously knowndistribution for Bufo speciosus and is within thedistribution of Bufo woodhousei.

The herpetofauna of the Caddo Lake watershedexists in the distinctive ecotone between theeastern deciduous forest and the westerngrasslands. The transition is most evident in thenarrow extension of the Texan Biotic Provincebetween the Austroriparian Biotic Province to theeast and the Kansan Biotic Province to the west.Compared to the herpetofauna of northwesternLouisiana, the Caddo Lake watershedherpetofauna has more species of both amphibiansand reptiles. This somewhat surprising result isprobably due to the greater habitat diversitywithin the watershed and to the presence ofspecies whose geographic distributions lieprimarily to the west of the watershed. The frogadditions of Gastrophryne olivacea andPseudacris clarki represent more western speciesin the watershed. Two more species ofsalamanders (Necturus beyeri and Plethodonglutinosus) are found in the western part of thearea. Among reptiles, the turtle fauna is the sameas that of northwestern Louisiana. HoweverSceloporus olivaceous and Cnemidophorus gularisrepresent western faunal components. Threeadditional species of snakes, Heterodon nasicus,Masticophis taeniatus, and Tropidocloniumlineatum are all additions from the west.

Of the 93 species that probably comprise theherpetofauna, 16 have the western edge of their

geographic distribution and ten have the easternedge of their distributions in the watershed.Another two have their southern edge and threehave their northern edge in the watershed. Thus,31 species (33%) realize their geographic limits inthe watershed. The 16 species of amphibians ofthis group represent 17 percent of the totalherpetofauna, the 15 species of reptiles, 16percent. This reinforces the importance of thisregion as an ecotone between major biogeographicregions.

Future workers should try to confirm thepresence of the following species in thewatershed:

Pseudacris clarkiGastrophryne olivaceaBufo vallicepsBufo speciosusRana areolataApalone muticaPhrynosoma comutumVirginia valereaElaphe guttata

In addition, three other species havedistributions that approach the watershed but atthis time are not known to be present within it.Kinostemon flavescens approaches to within 100miles or so from the west. Arizona elegans andHypsiglene torquata both approach from thesouthwest to within 50-100 miles.

ACKNOWLEDGEMENTS

This project was funded by grant number 1448-0004-94-952 (principal investigators James L.Ingold and Laurence M. Hardy) from the NationalBiological Service, Animal Ecology Branch,National Wetlands Research Center. Forcontinuous encouragement and administrativeassistance we thank Dr. Carroll Cordes, Chief,Animal Ecology Branch of the National WetlandsResearch Center.

For permission to examnine specimens in theircare, providing lists of specimens available,providing working space in their institutions, or

1995 HARDY: AMPHffiIANS AND REPTILES OF CADDO LAKE 29

otherwise attending to our needs while vlsltmgtheir collections we thank the following persons:Gail Manning (Dallas Museum of NaturalHistory), Drs. Max Nickerson and BobHenderson (Milwaukee Public Museum), Dr.Arnold G. Kluge and Greg Snider (University ofMichigan, Museum of Zoology), Dr. Harold K.Voris and Alan Resitar (Field Museum of NaturalHistory), Drs. Douglas Rossman and DavidGood (Louisiana State University, Museum ofZoology), Drs. Robert Cashner and Robert A.Thomas (University of New Orleans), Drs.Harold Dundee and Hank Bart (TulaneUniversity), Drs. Charles J. Cole, Linda Ford,and Ms. Irene Palser (American Museum ofNatural History), Drs. Roy W. McDiarmid andSteve W. Gotte (United States NationalMuseum), Ted Daeschler (Academy of NaturalSciences of Philadelphia), Wesley Hathaway (FortWorth Museum of Science and History), Dr.Jonathan Campbell and Carol Stuart (Universityof Texas at Arlington), Barbara Peterman and JimPeterson for information on localities and othercollections (Dallas Museum of Natural History),Dr. Nancy Glover McCartney (University ofArkansas, Fayetteville), Dr. Lew Densmore(Texas Tech University), Dr. William E.Duellman, John Simmons, and Joe Mendelson(University of Kansas), Dr. Tony Echelle andGoff Luttrell (Oklahoma State University), Dr.Ned Strength and Mr. Stone (San Angelo StateUniversity), Drs. Laurie Vitt, Gary Schnell, andMs. Vicki Byre (University of Oklahoma), Dr.Calvin Smith and David Lintz (Strecker Museum,Baylor University), Dr. Jim Dixon and Ms.Katherine Vaughn (Texas Cooperative WildlifeCollection, Texas A. & M. University), Dr.David Cannatella and Carol Malcolm (Universityof Texas at Austin), Dr. Charles Knight(Mississippi Museum of Natural Science), Drs.Donald Beaver, Chris Carmichael and Ms. LauraAbraczinskas (Michigan State UniversityMuseum), Dr. Robert Kennedy (CincinatiMuseum of Natural Science), Mr. John Condit(Ohio State University Museum), Dr. Tim Matson(Cleveland Museum Natural History), Mr. RandyBeavers (Robert A. Vines Science Center).

I especially appreciated the tireless and

enthusiastic assistance of Jeff Hering andChristina Fowler for helping with all aspects ofthis project.

LITERATURE CITED

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30

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31

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