An invertebrate survey of Penn Common (concentrating on...

An invertebrate survey of Penn Common

(concentrating on the wet habitats, especially the springs and flushes)

to cover those species and groups that help evaluate its biological

importance and help to inform its habitat management under

the Higher Level Scheme agreement.

Peter Boardman, Ian Cheeseborough, Nigel Jones, Richard Wright

November 2011

Penn Common invertebrate survey – 2011 (Boardman, Cheeseborough, Jones & Wright)

INDEX

i. Summary …………………………………………………………………………….. 3

1. Rationale …………………………………………………………………………………………. 4

2. Methodology…………………………………………………………………………………… 5

3. Results – overview………………………………………………………………………….. 5

4. Results – analysis of invertebrate communities………………………………. 9

5. Results – summary of ISIS analysis…………………………………………………… 10

6. Management Suggestions……………………………………………………………….. 11

7. Discussion……………………………………………………………………………………….. 13

ii. References………………………………………………………………………………. 15

iii. Acknowledgements…………………………………………………………………. 15

Appendix 1 – Full survey data for Penn Common golf course ……………………. 16

Appendix 2 – ISIS outputs for Penn Common …………………………………………….. 31


Summary

An invertebrate survey of Penn Common, near Wolverhampton, recorded 457 species of invertebrates

during 2011.

Of these, the vast majority were common species with only 7 species recorded of national or regional

importance. These were the flies; Spania nigra, Syntormon macula, Sphaerophoria taeniata, and Herina

frondescentiae. The weevil Trachyphloeus asperatus, the spider-hunting wasp Anoplius caviventris, and the

solitary wasp Tiphia minuta.

24 species were noted as new to Staffordshire (V.C. 39).

Data was put through ISIS software to identify species assemblages for the site. As a whole site, significant

species assemblage types (SATs) were identified for scrub edge, rich flower resource, and bark & sapwood

decay assemblages. The most significant broad assemblage type (BATs) noted was that for flowing water

suggesting the wetland habitats are important at the site. Heathland communities were not significant

across the whole site data.

When data was split down into habitat areas the species assemblage types produced significant results for

scrub-heath and moorland species assemblage amongst the dry acid grassland habitats, and the bark and

sapwood decay assemblages amongst the aculeate habitats.

The calcareous flush was found to be of little significance currently.

Penn Common was found to be regionally important for its aculeate hymenoptera assemblage, though much

of this is associated with dead-wood rather than heathland species.

Management suggestions were made to increase the suitability of habitat for heathland hymenoptera,

amongst other habitats across the site.


1. Rationale

1.1 An invertebrate survey was carried out on Penn Common (SO898945), near Wolverhampton, to identify the

most important invertebrate communities at the site and examine how they compare to other similar sites

in the area (Staffordshire and West Midlands primarily but also east Shropshire and north Worcestershire).

1.2 Outputs were to examine how the invertebrate communities can be conserved and, where possible,

enhanced (under management through the Higher Level Stewardship (HLS) agreement).

1.3 Penn Common has historically been recognised as an important site in terms of nature conservation with

historical botanical records suggesting important habitats (particularly base rich flushes and heathland) for

invertebrates at the site. It was noted that grazing had recently ceased but more up to date information

regarding habitat quality and suitability for invertebrate assemblages was lacking, hence the need to assess

the site through survey work.

1.4 The site hosts Penn Common Golf Course, where golf has been played since 1893 (see map 1), and is

intersected roughly down the centre on a north / south plane by a minor road. It sits to the south of Penn, a

suburb of the major conurbation of Wolverhampton, and to the west of Sedgley, and is located in

Staffordshire (V.C. 39).

Map 1 – Penn Common Golf Course (www.maps.google.co.uk.)


2. Methodology

2.1 The survey methodology followed that laid out in ‘Surveying terrestrial and freshwater invertebrates for

conservation evaluation’ (Drake et al, 2007) and results were analysed using the ISIS software, developed by

Natural England, to assess invertebrate communities at the site.

2.2 The surveyors used appropriate methodologies for their particular invertebrate interests; Pete Boardman &

Nigel Jones (diptera – timed sweeping over flushes, damp and marginal habitats, plus netting individual

species of interest, water traps, limited Malaise trapping), Ian Cheeseborough (aculeate hymenoptera –

netting individual species and observation of nesting aggregations and dead wood habitats, water traps,

limited Malaise trapping), Richard Wright (coleoptera – pitfall trapping and sweeping / beating heather and

other vegetation, hand searching).

2.3 Where possible other invertebrate and insect orders were recorded when encountered. These included the

Lepidoptera which were being surveyed by another contractor so little attention was given to the group

during our site visits.

2.4 Visits were timed to assess the peak phenology of the groups studied.

3. Results - overview

3.1 Survey visits were undertaken in April, May, June, July, September and October by the surveyors, after an

initial visit in early April accompanied by the course manager, Paul Mills from Penn Common Golf Course.

3.2 Survey work during 2011 was in some ways hampered by the very dry conditions that occurred through the

year. A hot, dry spring was followed by a generally dull, dry summer and the dry weather continued on into

autumn. Several of the wet or damp areas where survey work was focused were subject to desiccation and

at times insects were hard to come across. Nevertheless a total of 457 species were recorded at the site.

3.3 The table below details the breakdown of invertebrate orders recorded;

Order No of Species

Diptera (true flies) 220 Coleoptera (beetles) 82 Hymenoptera (bees, wasps and ants) 73 Aranae & Opiliones (spiders and harvestman) 33 Hemiptera (true bugs) 21 Lepidoptera (butterflies and moths) 15 Diplopoda (millipedes) 4 Odonata (damselflies and dragonflies) 4 Isopoda (woodlice) 3 Orthoptera (grasshoppers and crickets) 2 Total - 457

Table 1 – summary of invertebrate orders recorded at Penn Common during the 2011 survey


3.4 Of the 457 species recorded few were noted to be of conservation importance except for the following 7

species;

3.5 Diptera (Spaniidae) Spania nigra Meigen, 1830 Liverwort Snipefly. This species is listed as Notable but being

small is often overlooked. Larvae leaf mine species of liverworts and Pellia neesia has been noted to host the

fly (Stubbs & Drake, 2001). It has a widespread distribution within the United Kingdom

(www.data.nbn.org.uk) and appears here at its first Staffordshire site (Jukes, 2005). It has been recorded in

Shropshire from dingle woodland near Ironbridge Gorge and at three sites around the Long Mynd in south

Shropshire (Shropshire Ecological Data Network pers. comm.)

3.6 Diptera (Dolichopodidae) Syntormon macula Parent, 1927. This uncommon long-legged fly is often

associated with dingle woodlands or damp places and has a mostly south-western distribution. It was

originally designated a Red Data Book species but is now recognised as more commonly occurring, so was

given the lower risk (near threatened) status to reflect this (Falk & Crossley, 2005). It appears here at its first

Staffordshire site (Jukes, 2005). It is known from three dingle woodlands in Shropshire (Boardman, 2005 &

Shropshire Ecological Data Network pers. comm.).

3.7 Diptera (Syrphidae) Sphaerophoria taeniata (Meigen, 1822). This hoverfly has a mostly southern distribution

with Kent across to Somerset seeing the majority of records. It is uncommon outside of this range

(www.hoverfly.org.uk), and it appears to be heading north and is recorded here at its first Staffordshire

location (Jukes, 2005).

3.8 Diptera (Ulidiidae) Herina frondescentiae (Linnaeus, 1758). This non-tephritid picture-wing fly has a

widespread distribution nationally but can be considered local in the Midlands. It is associated with damp

grasslands and appears new to Staffordshire at this site (Jukes, 2005).

Herina frondescentiae (photo by Nigel Jones)

3.9 Coleoptera (Curculionidae) Trachyphloeus asperatus Boheman, 1843. This weevil is associated with lowland

calcareous grassland and has a widespread distribution in England, with mainly a maritime distribution in

Wales. It is classed as Notable A, and is the first Staffordshire record since 1907 (Slawson & Brown, 2009).


3.10 Hymenoptera (Pompilidae) Anoplius caviventris (Aurivillius, 1907). This spider-hunting wasp has a mostly

southern distribution (www.data.nbn.org) and is classed as Nationally Notable B by Falk (1991) but ‘very

rare’ by Michael Archer in his more recent review. It appears here as new to Staffordshire (Webb, Bloxham

and Slawson, 2005). It is known from one site in Worcestershire (Trevis, 2004) and a single site in Shropshire

(Ian Cheeseborough pers. comm. & Shropshire Ecological Data Network pers. comm.).

3.11 Hymenoptera (Tiphiidae) Tiphia minuta Vander Linden, 1827. This solitary wasp is a parasite of dung beetles

(Scarabaeidae) and is considered to be regionally scarce (Jukes, 2004).

3.12 As well as these important species several others were noted as new to Staffordshire when checked against

current checklists;

3.13 Diptera;

craneflies (Limoniidae); Dicranomyia affinis, Eloeophila submarmorata, Erioptera squalida.

long-legged flies (Dolichopodidae); Hercostomus parvilamellatus, Teuchophorus calcaratus, T.

monacanthus, T. nigricosta, T. simplex,

snail-killing flies (Sciomyzidae); Pherbellia schoenherri, Tetranocera fuscinervis.

flat-footed flies (Platypezidae); Agathomyia antennata

druid flies (Clusiidae); Clusiodes gentilis

stalk flies (Psilidae); Chamaepsila rosae

shore flies (Ephydridae); Ditrichophora calceata

big-headed flies (Pipunculidae); Jassidophaga villosa

lauxaniid flies (Lauxaniidae); Minettia fasciata

parasitic flies (Tachinidae); Lydella stabulans, Vibrissina debilitata

dung flies (Scathophagidae); Nanna flavipes

anthomyid flies (Anthomyiidae); Anthomyia procellaris

muscid flies (Muscidae); Coenosia infantula

3.14 Hemiptera;

leatherbugs (Coreidae); Coreus marginatus (Dock Bug)

3.15 Coleoptera;

ground beetle (Carabidae); Agonum emarginatum

3.16 Hymenoptera;

solitary wasp (Sphecidae); Psenulus concolor

3.17 As well as assessing the site as a whole, it was broken down to the main areas of interest as detailed in the

tender document, to begin to assess the invertebrate communities present at the site.

3.18 They were as follows (see map 2);

1 – damp grassland, pond etc at SO89709483

2 – damp, rushy grassland with scrub at SO89629479

3 – species-rich dry grassland at SO895294444


4 – acid grassland / heather at SO89459440

5 – wet woodland / flush with calcareous influence at SO89599442

6 – acid grassland / heather at SO89799476

7 – damp woodland at SO90309478

8 – marsh / wet woodland and wet grassland at SO90379448

8a* – open scrub / woodland with bracken at SO903944

9 – woodland flush / pond / open ditch at SO90259456

9a* –cleared patch with dead wood at SO90269452

10 – flower-rich clearing at SO90269452

11 – damp woodland at SO89639441

12 – eastern footpath / sheltered hedgerow at SO90349456

3.19 Other records away from these areas were noted as ‘general site records’ and given a site centroid grid

reference of SO898945.

3.20 Areas 8a and 9a were separated from areas 8 and 9 respectively to take into account good aculeate

hymenoptera habitat including dead wood, which would otherwise potentially biased species assemblage

analysis in the larger parent areas.

3.21 All records are detailed in Appendix of this report within Table 2.

Map two – compartments survey during Penn Common survey 2011


4. Results - Analysis of Communities

4.1 Data was put into the ISIS spreadsheet to identify Species Assembly Types (SATs) and Broad Assemblage

Types (BATs) that may exist at Penn Common. It should be noted that approximately 30 species could not be

used in the ISIS analysis as they registered an “error”. These species are ones excluded from ISIS as they are

ones where sufficient information is known about them, are vagrants, or their habitats do not fit into the

classification.

4.2 All SATs and BATs tables used in this analysis are produced in the Appendix of this report.

4.3 Whole site data - ISIS recognised 11 SATs of which three (scrub edge, rich flower resource and bark &

sapwood decay) suggested strong species assemblies (favourable condition in CSM). Less strong SATs were

those from the dung, open short sward, shingle banks, seepage, reed-fen and pools, scrub-heath and

moorland, bare sand and chalk, and heartwood decay assemblies.

4.4 This suggests that the site as a whole has more value for its scrub / woodland edge, associated dead wood,

and floral resources than for its heathland or wetland features. This may represent a decline in the state of

the habitat but this can only be suggested, rather than proven, without previous evidence of invertebrate

survey work at the site and is based upon the perception that features such as the calcareous flush and

heathland were perhaps once in better condition due to the historical botanical records that exist.

4.5 Whole site data - 8 BATs were identified, of which the flowing water broad assembly was noted as

favourable (in terms of CSM) and achieved the highest rarity score (154). The grassland and scrub matrix

achieved the highest BAT species richness score of 154. These figures suggest that within the site as a whole

the grassland and scrub matrix and the flushes are key features. Other BATs identified were; unshaded early

successional mosaic, permanent wet mire, wood decay, shaded field and ground layer, mineral marsh and

open water, arboreal canopy.

4.6 Calcareous flush at SO89599442 (area 5) - ISIS only recognised a single SAT from this data, a single species

of the shingle bank assembly.

4.7 Calcareous flush at SO89599442 (area 5) - 5 BATs were identified; grassland and scrub matrix, flowing

water, mineral marsh and open water, shaded field and ground layer, and permanent wet mire. No features

of either SATs or BATs reached favourable levels and this suggests that this flush is of limited value for

invertebrates in its current condition, with few specialists identified. It should be noted that part of this area

of the site has only recently been cleared of scrub.

4.8 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - ISIS recognised 8 SATs from

this area but none of them were strong communities. They were; seepage, open short sward, scrub edge,

reed-fen and pools, bark and sapwood decay, rich flower resource, scrub-heath and moorland, bare sand

and chalk. 8 BATs were recognised with the flowing water broad assembly noted as favourable.

4.9 Wet habitats in the east of the site (areas 8 and 9 – not including 8a and 9a) - BATs recognised were;

grassland and scrub matrix, permanent wet mire, flowing water, mineral marsh and open water, shaded

field and ground layer, unshaded early successional mosaic, wood decay and arboreal canopy.


4.10 This analysis suggests no strong species assemblages occur within this wetland area, though it does

recognise that flowing water is important within the broader assemblage of species and is a key feature of

the site.

4.11 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - ISIS showed 5 SATs within these areas (mostly

targeted for aculeate hymenoptera) and one species assembly type stands out as favourable – the bark and

sapwood decay. We had already noted that there was a higher than expected list of dead wood aculeate

hymenoptera, considering tree cover is limited, and few old or veteran trees occur at the site, so it was

pleasing to see ISIS agreeing with our own interpretation. ISIS suggests 26 species within the bark and

sapwood decay assembly. Other SATs listed were; rich flower resource, dung, scrub edge and open short

sward.

4.12 Scrubby, flower rich and drier habitats (areas 8a, 9a and 10) - 5 BATs were identified with three standing

out; unshaded early successional mosaic, wood decay and grassland and scrub matrix. The two remaining

and less strongly represented BATs were arboreal canopy and mineral marsh and open water.

4.13 Dry acid grassland and heather (areas 3, 4 and 6) - ISIS listed 4 SATs within these areas (mostly targeted for

coleoptera) and the scrub heath and moorland assembly was noted as favourable with 10 species within the

assembly type. The remaining three SATs were bare sand and chalk, scrub edge, and open short sward.

4.14 Dry acid grassland and heather (areas 3, 4 and 6) - 7 BATs were identified with the grassland and scrub

matrix assembly standing out far above all others listed. The others were; unshaded early successional

mosaic, mineral marsh and open water, shaded field and ground layer, permanent wet mire, flowing water.

This suggests that this area of the site on the western side is the most important area of acid grassland and

scrubby heath, but bare ground and short sward assemblies are poorly represented.

4.15 Damp grassland (areas 1 and 2) - ISIS was unable to identify any SATs for this habitat.

4.16 Damp grassland (areas 1 and 2) - 5 BATs. Of these the strongest broad assembly was amongst grassland

and scrub matrix species. The permanent wet mire broad assembly was not overly strong but nevertheless

worthy of mention whilst the mineral marsh and open water, shaded field and ground layer, and flowing

water broad assemblies were poorly represented. This data suggests that damp grassland habitats at the

site of the least productivity for species assemblages on the site.

5. Results – Summary of ISIS analysis

5.1 Analysis of the species and broad species assemblages of Penn Common using ISIS suggests that the scrub

matrix assembly, with associated dead wood, flowing water, dry acid grassland and flower rich grassland are

the key current habitat types at the site for invertebrates.


5.2 Heathland containing good quality heather and bare open ground seems in too short a supply to host

important invertebrate assemblages at present levels. Aculeate hymenoptera assemblages are more reliant

upon dead wood than traditionally suitable heathland habitat at the site.

5.3 The calcareous flush in the east of the site is currently not important in terms of species assemblages;

through it does host three broad wetland assemblages and a scrub matrix assemblage. This suggests that as

the flush scrubbed up it became less important for wetland species, though with recent and continued

habitat management may revert to hold more characteristic wetland assemblages of note in the future.

5.4 Other wet habitats at the site were not viewed as strongly important in terms of individual species

assemblages but flowing water as a broad assembly was noted as important.

5.5 Damp grassland at the north of the site is of little importance for species assemblages, which may reflect

the lack of grazing at the site, or simply the dry weather from 2011.

6. Management Suggestions

6.1 Current management of habitats needs to be continued and increased to maintain a mosaic of shady wet,

open wet, flower-rich (low growing and tussocky) grassland, and heathland habitats with heather and bare

ground. For example the aculeate hymenoptera at Penn Common need nesting opportunities (sandy bare

ground or dead wood), and foraging/hunting grounds in proximity to these nesting sites. Keeping the mix of

bare ground, short flower-rich grassland, rank patches of vegetation, scrub and woodland is ideal.

6.2 Wet areas and flushes;

Probably too many of the wet areas are, or have been until recently, scrubbed up to the detriment of

wetland diptera and other groups. These areas should be rotationally cut as has been done in area 5.

The main passage of water from north of south along the eastern boundary of the site (area 8) is

canalised and the grass is mown right up to the edge. Though there is probably not too much that can be

done about the canalization given the proximity to the fairway, some grading down the stream banks (to

make them more gently sloped) and a more relaxed approach to grass cutting may provide more

marginal habitat in this area.

The pond at the bottom of this ditch is becoming choked with vegetation and would benefit from some

clearance.

6.3 Heathland;

The current managed heather areas on the site are just too small to be of much use as heathland

pockets, and are often just a couple of straggly heather plants in the middle of a fairway. Areas should

be expanded and could be seeded with heather seeds from a local source.


More bare ground would benefit nesting aggregations of bees. Bare ground is often seen as an erosion

problem or unsightly on golf courses and seeded with a grass mixture at the first opportunity. Some

decent sized scrapes on marginal areas of the fairways (as large as practicably possible) would provide

potentially a great deal of bare ground across the course. This sort of approach has recently been

undertaken at Highgate Common for the rare solitary bee Andrena nigrispina. The scrapes were

observed on a recent site visit there by two of our surveyors. Several of these bee scrapes are needed

around the site to encourage nesting aggregations providing horizontal and vertical surfaces into which

bees and wasps can burrow. These can cover a variety of aspects, but are probably the single most

important habitat management tool for heathland bees.

Solitary bees Andrena barbilabris and Crossocerus quadrimaculatus were observed using the bunkers for

nesting but only around the unraked sides, therefore whilst bunkers are a key aspect for the golfers at

the site, parts of them need not be regulary raked to maintain more breeding habitat for solitary bees.

This may be a matter of education of golfers rather than management per se.

6.4 Grassland;

The amount of acid grassland and species rich grassland should be increased at the site. This could be

done by relaxing mowing regimes on the fairways. Instead of a standard lozenge shape of a golf hole, the

fairways could be altered so that more of an hour glass shape exists filtering golfers down the middle of

the fairway in areas less commonly used on each hole (for instance between the tee and where the

average drive from that tee lands).

Flower rich areas need cutting and raking, with the arisings removed to restrict nutrient input. Cutting of

flowering areas needs to be timed so that they are cut after flowering (not during) and with areas left for

invertebrates to over-winter in seed heads or tussocks.

Where thick areas of bracken persist, some rotational cutting to open up the structure would add

another dimension to the habitat.

6.5 Dead wood;

ISIS analysis identified dead wood as a key feature of aculeate hymenoptera at the site, therefore as well

as ensuring existing amounts of dead wood are retained, more dead wood should be left available for

aculeate hymenoptera and coleoptera to utilise.

Standing dead wood is more valuable than fallen dead wood. Often standing dead wood is removed as

it is deemed to be deleterious to health and safety, however reducing the height of dead wood by

removing large branches or the upper half of the tree whilst leaving the trunk still standing to maintain

safety is still leaving a valuable resource in place.


Standing dead wood could be further improved by drilling small holes into it with a battery powered drill

and bit. This method has worked well at sites in Shropshire where it has been adopted and used by a

variety of dead wood nesting bees and wasps (Cheeseborough pers. comm.).

The existing dead wood left on the ground is very quickly covered with vegetation making it unsuitable

quickly unsuitable. Fallen dead wood should therefore be kept clear of vegetation or kept higher up of

the ground if this is a problem.

7. Discussion

7.1 Penn Common was not considered to be of high conservation status in its current state. The vast majority of

species encountered were common in terms of nature conservation interest, though some can be

considered heathland, or wetland specialists.

7.2 The strongest species assemblages represented those of scrub edge, flower-rich grassland, and dead wood

assemblages of aculeate hymenoptera. Had heathland habitat been in better condition the species

assemblages would have reflected this with higher scores in the scrub-heath and bare sand assemblages.

7.3 The calcareous flush was found to be of less importance for wetland species than had been hoped based

upon historical botanical records.

7.4 One surprise was the large amount of dead wood aculeate hymenoptera found, considering the lack of

established veteran trees.

7.5 Clearly the impact of a very popular local amenity such as the Golf Course undoubtedly has had an effect

upon the habitat, though this is not to say the impact of more considered habitat management could

improve the existing situation under the Higher Level Scheme.

7.6 It should be noted that our results were affected by the lack of rain through the 2011 season, which led to

the drying out of most of the “wet” areas on the site by the end of the 2011 recording season, and clearly

had a detrimental effect on the numbers of wetland diptera and coleoptera sampled.

7.7 In terms of the aculeate hymenoptera recorded at the site, the species list achieved during the survey

compares favourably with other Staffordshire heathlands, and Penn Common could perhaps be listed as

fourth best Staffordshire heathland site (behind Highgate Common, Gentleshaw Common and Bevins

Birches) (Jukes, 2004) on its aculeate list.

7.8 It is noted that the better West Midlands heathland sites have decent amounts of bare or sparsely

vegetated ground, heather dominated areas and flower-rich heathy verge (Falk, 2010). Penn Common has

some flower-rich areas but not enough heather dominated areas and too little bare or sparsely vegetated

ground to host the level of aculeate hymenoptera that might improve its standing within the West Midlands

league table.


7.9 As already mentioned the amount of dead wood fauna was unexpected and this seems to be boosting the

site aculeate list. This is perhaps masking the lack of heathland specialists found during this survey. Although

above sites like Pelsall Common in the league table, Penn Common does however have fewer species of

national or regional interest than many other of the key West Midlands heathland sites, with only one very

rare species recorded (Anoplius caviventris) and two regionally scarce species (Tiphia minuta and Psenulus

concolor*) (Note – Psenulus concolor is viewed as common in Shropshire (Cheeseborough pers.comm.) but

is absent from the Staffordshire checklist (Webb, Bloxham & Slawson, 2011), and is also absent from Steven

Falk’s West Midlands region review of bee and wasp assemblages (Falk, 2010)). Both Anoplius caviventris

and Psenulus concolor appear to be new to Staffordshire. It should be noted that Anoplius caviventris was

listed as a general site record during our survey and as such was not included in the ISIS analysis for any

individual compartment, though was part of the whole site analysis.

7.10 Most coleoptera noted were common species with only three heathland specialists recorded (Bradycellus

ruficollis, Lochmaea suturalis and Micrelus ericae). This again illustrates that good quality heathland habitat

is currently at a premium on Penn Common. Approximately 27 species of beetle recorded were wetland

specialists, though all were considered common.

7.11 Wetland diptera were well represented but again most species were considered common. As stated earlier

the subsequent drying out of the wet areas, as the 2011 recording season progressed, did have a deleterious

effect upon our survey and towards the end of the season there were few wetland flies around, particularly

craneflies.

7.12 It would be hoped that further habitat management work could improve the wetland areas across the site.

ISIS analysis of the species assemblages of the calcareous flush showed how species associated with scrub

currently dominate the wetland assemblages. The continued removal of scrub here should begin to show a

change in the assemblages over time, as the vegetation responds to more available light.

7.13 Elsewhere, on the eastern side of the site, the flowing water species assemblages were identified by ISIS as

of importance and these should at least be maintained.


References;

Boardman, P.J. (2005); The Red Data Book Invertebrates of Shropshire, a compilation and review of data.

Unpublished report for Shropshire Council.

Drake, C.M, Lott, D.A., Alexander, K.N.A, & Webb, J. (2007); Surveying terrestrial and freshwater invertebrates for

conservation evaluation. Natural England. Sheffield.

Jukes, A. (2004); An evaluation of the invertebrate fauna of Highgate Common in relation to other sites within

Staffordshire and the West Midlands. Unpublished report for Natural England.

Jukes, A. (2005); A provisional atlas of the True Flies (including hoverflies) (Diptera) of Staffordshire. Staffordshire

Ecological Record (web version)

Falk, S.J. (1991); A review of the scarce and threatened bees, wasps and ants of Great Britain. JNCC. Peterborough.

Falk, S.J. (2010); The bee and wasp assemblages (Hymenoptera: Aculeata) of some key heathland sites in the West

Midlands region. British Journal of Entomology and Natural History. Vol. 23 p77-97. BENHS. Dinton.

Falk, S.J. & Crossley, R. (2005); A review of the scarce and threatened flies of Great Britain. JNCC. Peterborough.

Slawson, C., & Brown, E. (2009); A provisional atlas of the Beetles (Coleoptera) of Staffordshire. Staffordshire

Ecological Record (web version).

Stubbs, A.E., & Drake, C.M., (2001); British Soldierflies and their allies: an illustrated guide to their identification and

ecology. BENHS. Dinton Pastures.

Trevis, G. (2004); Aculeate Hymenoptera in Worcestershire. Worcestershire Record No.16 p27-28

Webb, J., Bloxham, M. & Slawson C. (2005); A provisional atlas of the Bees and Wasps (Hymenoptera) of

Staffordshire. Staffordshire Ecological Record (web version).

Websites

www.data.nbn.org.uk – The NBN Gateway

www.hoverfly.org.uk – The Hoverfly Recording Scheme

www.staffs-ecology.org.uk – Staffordshire Ecological Record

Acknowledgements

The authors would like to thank Paul Mills at Penn Common Golf Course for access to the course and showing us

around on our first visit, and Jon Webb at Natural England for supplying ISIS. We would also like to thank for golfers

who were playing the course during our visits and fully expect that our attendance was blamed for more than one

bad shot or lost ball.


APPENDIX 1 – Full survey data for Penn Common Golf Course

Order Taxon Compartment Grid ref Date Determiner

Araneae Alopecosa pulverulenta 3 SO89529444 2011 Richard Wright

Araneae Alopecosa pulverulenta 4 SO89459440 2011 Richard Wright

Araneae Antistea elegans 5 SO89599442 2011 Richard Wright

Araneae Antistea elegans 8 SO90379448 2011 Richard Wright

Araneae Bathyphantes gracilis 6 SO89799476 2011 Richard Wright

Araneae Centromerita bicolor 2 SO89629479 2011 Richard Wright



Araneae Drassyllus pusillus 3 SO89529444 2011 Richard Wright

Araneae Drassyllus pusillus 7 SO90309478 2011 Richard Wright

Araneae Erigone atra 1 SO89709483 2011 Richard Wright

Araneae Erigone dentipalpis 3 SO89529444 2011 Richard Wright

Araneae Erigone dentipalpis 4 SO89459440 2011 Richard Wright

Araneae Gongylidiellum vivum 5 SO89599442 2011 Richard Wright

Araneae Gongylidiellum vivum 8 SO90379448 2011 Richard Wright

Araneae Lepthyphantes tenuis 2 SO89629479 2011 Richard Wright




Araneae Leptorhoptrum robustum 4 SO89459440 2011 Richard Wright

Araneae Leptorhoptrum robustum 6 SO89799476 2011 Richard Wright

Araneae Meta mengei 1 SO89709483 2011 Richard Wright

Araneae Meta segmentata 2 SO89629479 2011 Richard Wright

Araneae Micaria pulicaria 3 SO89529444 2011 Richard Wright



Araneae Micrargus herbigradus 7 SO90309478 2011 Richard Wright

Araneae Neottiura bimaculata 4 SO89459440 2011 Richard Wright

Araneae Oedothorax gibbosus 1 SO89709483 2011 Richard Wright





Araneae Oedothorax retusus 1 SO89709483 2011 Richard Wright

Araneae Oedothorax retusus 5 SO89599442 2011 Richard Wright

Araneae Pachygnatha clercki 1 SO89709483 2011 Richard Wright




Araneae Pachygnatha degeeri 1 SO89709483 2011 Richard Wright




Araneae Pardosa nigriceps 3 SO89529444 2011 Richard Wright

Araneae Pardosa nigriceps 6 SO89799476 2011 Richard Wright

Araneae Pardosa palustris 4 SO89459440 2011 Richard Wright

Araneae Pardosa prativaga 1 SO89709483 2011 Richard Wright






Araneae Pardosa pullata 1 SO89709483 2011 Richard Wright







Araneae Pirata hygrophilus 9 SO90259456 2011 Richard Wright

Araneae Pirata piraticus 1 SO89709483 2011 Richard Wright



Araneae Pocadicnemis pumila 5 SO89599442 2011 Richard Wright

Araneae Pocadicnemis pumila 6 SO89799476 2011 Richard Wright

Araneae Robertus lividus 4 SO89459440 2011 Richard Wright

Araneae Robertus lividus 6 SO89799476 2011 Richard Wright

Araneae Walckenaeria vigilax 5 SO89599442 2011 Richard Wright

Araneae Walckenaeria vigilax 9 SO90259456 2011 Richard Wright

Araneae Xysticus cristatus 3 SO89529444 2011 Richard Wright



Coleoptera Acupalpus dubius 1 SO89709483 2011 Richard Wright







Coleoptera Adalia bipunctata site record SO898945 20/05/2011 Nigel Jones

Coleoptera Adalia bipunctata 9 SO90259456 29/06/2011 Pete Boardman

Coleoptera Agonum emarginatum 5 SO89599442 2011 Richard Wright



Coleoptera Agonum fuliginosum 5 SO89599442 2011 Richard Wright

Coleoptera Agonum fuliginosum 7 SO90309478 2011 Richard Wright

Coleoptera Agonum muelleri 8 SO90379448 2011 Richard Wright

Coleoptera Agonum viduum 5 SO89599442 2011 Richard Wright

Coleoptera Agonum viduum 8 SO90379448 2011 Richard Wright

Coleoptera Agriotes acuminatus 3 SO89529444 2011 Richard Wright

Coleoptera Agriotes lineatus 2 SO89629479 2011 Richard Wright

Coleoptera Agriotes pallidulus 3 SO89529444 2011 Richard Wright

Coleoptera Agriotes sputator 4 SO89459440 2011 Richard Wright

Coleoptera Amara communis 3 SO89529444 2011 Richard Wright



Coleoptera Amara lunicollis 3 SO89529444 2011 Richard Wright



Coleoptera Amara plebeja 1 SO89709483 2011 Richard Wright





Coleoptera Anacaena globulus 1 SO89709483 2011 Richard Wright





Coleoptera Anotylus rugosus 1 SO89709483 2011 Richard Wright








Coleoptera Anthobium unicolor 2 SO89629479 2011 Richard Wright

Coleoptera Anthobium unicolor 7 SO90309478 2011 Richard Wright

Coleoptera Aphodius prodromus 8 SO90379448 2011 Richard Wright

Coleoptera Aphthona euphorbiae 3 SO89529444 2011 Richard Wright

Coleoptera Apion frumentarium 5 SO89599442 20/04/2011 Pete Boardman

Coleoptera Apion frumentarium 7 SO90309478 20/04/2011 Pete Boardman

Coleoptera Athous haemorrhoidalis 7 SO90309478 2011 Richard Wright

Coleoptera Barypeithes pellucidus 2 SO89629479 2011 Richard Wright




Coleoptera Bembidion decorum 5 SO89599442 2011 Richard Wright

Coleoptera Bembidion guttula 5 SO89599442 2011 Richard Wright

Coleoptera Bembidion guttula 6 SO89799476 2011 Richard Wright

Coleoptera Bembidion lampros 3 SO89529444 2011 Richard Wright



Coleoptera Bembidion mannerheimi 1 SO89709483 2011 Richard Wright






Coleoptera Bembidion obtusum 1 SO89709483 2011 Richard Wright





Coleoptera Brachygluta haematica 4 SO89459440 2011 Richard Wright

Coleoptera Brachygluta haematica 8 SO90379448 2011 Richard Wright

Coleoptera Bradycellus ruficollis 4 SO89459440 2011 Richard Wright

Coleoptera Bradycellus ruficollis 6 SO89799476 2011 Richard Wright

Coleoptera Bradycellus verbasci 4 SO89459440 2011 Richard Wright

Coleoptera Calvia quattuordecimguttata 7 SO90029443 20/04/2011 Pete Boardman

Coleoptera Calvia quattuordecimguttata site record SO898945 20/05/2011 Nigel Jones

Coleoptera Cantharis nigra 3 SO89529444 2011 Richard Wright

Coleoptera Cassida rubiginosa 3 SO89529444 2011 Richard Wright

Coleoptera Ceratapion gibbirostre 3 SO89529444 2011 Richard Wright

Coleoptera Chaetocnema concinna 3 SO89529444 2011 Richard Wright

Coleoptera Chaetocnema concinna 4 SO89459440 2011 Richard Wright

Coleoptera Chaetocnema hortensis 3 SO89529444 2011 Richard Wright

Coleoptera Chilocorus renipustulatus site record SO898945 20/05/2011 Nigel Jones

Coleoptera Clivina fossor 8 SO90379448 2011 Richard Wright

Coleoptera Clytus arietis 12 SO90349456 18/04/2011 Ian Cheeseborough

Coleoptera Clytus arietis 8 SO90379448 20/05/2011 Nigel Jones

Coleoptera Coccidula rufa 1 SO89709483 2011 Richard Wright

Coleoptera Coccidula rufa 5 SO89599442 2011 Richard Wright

Coleoptera Coccinella septempunctata 4 SO89459440 2011 Richard Wright

Coleoptera Coccinella septempunctata site record SO899945 18/04/2011 Ian Cheeseborough

Coleoptera Coccinella septempunctata 7 SO90029443 20/04/2011 Pete Boardman

Coleoptera Coccinella septempunctata 7 SO90309478 20/04/2011 Pete Boardman

Coleoptera Coccinella septempunctata site record SO898945 20/05/2011 Nigel Jones

Coleoptera Corticarina minuta 3 SO89529444 2011 Richard Wright

Coleoptera Cortinicara gibbosa 4 SO89459440 2011 Richard Wright



Coleoptera Dryops ernesti 1 SO89709483 2011 Richard Wright





Coleoptera Eutrichapion ervi 3 SO89529444 2011 Richard Wright

Coleoptera Eutrichapion viciae 3 SO89529444 2011 Richard Wright

Coleoptera Exochomus quadripustulatus 7 SO90309478 2011 Richard Wright

Coleoptera Gabrius breviventer 5 SO89599442 2011 Richard Wright

Coleoptera Gabrius breviventer 8 SO90379448 2011 Richard Wright

Coleoptera Gymnetron pascuorum 3 SO89529444 2011 Richard Wright

Coleoptera Gymnetron pascuorum 4 SO89459440 2011 Richard Wright

Coleoptera Harmonia axyridis site record SO898945 20/05/2011 Nigel Jones

Coleoptera Lathrobium geminum 5 SO89599442 2011 Richard Wright

Coleoptera Lathrobium geminum 8 SO90379448 2011 Richard Wright

Coleoptera Leiosoma deflexum 3 SO89529444 2011 Richard Wright

Coleoptera Lesteva sicula 5 SO89599442 2011 Richard Wright

Coleoptera Lesteva sicula 8 SO90379448 2011 Richard Wright

Coleoptera Lochmaea suturalis 4 SO89459440 2011 Richard Wright

Coleoptera Lochmaea suturalis 6 SO89799476 2011 Richard Wright

Coleoptera Malthinus flaveolus 8 SO90379448 29/06/2011 Pete Boardman

Coleoptera Mecinus pyraster 3 SO89529444 2011 Richard Wright

Coleoptera Micrelus ericae 4 SO89459440 2011 Richard Wright

Coleoptera Micrelus ericae 6 SO89799476 2011 Richard Wright

Coleoptera Nargus velox 4 SO89459440 2011 Richard Wright



Coleoptera Nebria brevicollis 3 SO89529444 2011 Richard Wright



Coleoptera Neocrepidodera ferruginea 9 SO90259456 2011 Richard Wright

Coleoptera Neocrepidodera transversa 3 SO89529444 2011 Richard Wright

Coleoptera Notaris acridulus 5 SO89599442 2011 Richard Wright

Coleoptera Notaris acridulus 8 SO90379448 2011 Richard Wright

Coleoptera Ochthebius minimus 9 SO90259456 2011 Richard Wright

Coleoptera Oedemera nobilis 9a SO90269452 20/05/2011 Ian Cheeseborough

Coleoptera Olibrus aeneus 5 SO89599442 2011 Richard Wright

Coleoptera Oxystoma craccae 3 SO89529444 2011 Richard Wright

Coleoptera Paradromius linearis 3 SO89529444 2011 Richard Wright

Coleoptera Perapion curtirostre 1 SO89709483 2011 Richard Wright



Coleoptera Perapion hydrolapathi 1 SO89709483 2011 Richard Wright

Coleoptera Philonthus cognatus 1 SO89709483 2011 Richard Wright





Coleoptera Phyllobius pomaceus 9 SO90259456 20/04/2011 Pete Boardman

Coleoptera Phyllobius roboretanus 7 SO90309478 2011 Richard Wright

Coleoptera Phyllotreta tetrastigma 5 SO89599442 2011 Richard Wright



Coleoptera Poecilus versicolor 3 SO89529444 2011 Richard Wright

Coleoptera Propylea quattuordecimpunctata 7 SO90309478 2011 Richard Wright

Coleoptera Propylea quattuordecimpunctata site record SO899945 18/04/2011 Ian Cheeseborough

Coleoptera Propylea quattuordecimpunctata 8a SO903945 28/07/2011 Ian Cheeseborough

Coleoptera Protapion assimile 3 SO89529444 2011 Richard Wright

Coleoptera Protapion fulvipes 3 SO89529444 2011 Richard Wright

Coleoptera Protapion trifolii 3 SO89529444 2011 Richard Wright

Coleoptera Psyllobora vigintiduopunctata 3 SO89529444 2011 Richard Wright

Coleoptera Pterostichus diligens 5 SO89599442 2011 Richard Wright


Coleoptera Pterostichus diligens 9 SO90259456 2011 Richard Wright

Coleoptera Pterostichus madidus 3 SO89529444 2011 Richard Wright

Coleoptera Pterostichus melanarius 1 SO89709483 2011 Richard Wright




Coleoptera Pterostichus nigrita 5 SO89599442 2011 Richard Wright




Coleoptera Pterostichus strenuus 1 SO89709483 2011 Richard Wright








Coleoptera Pterostichus vernalis 5 SO89599442 2011 Richard Wright



Coleoptera Quedius curtipennis 1 SO89709483 2011 Richard Wright





Coleoptera Quedius fuliginosus 4 SO89459440 2011 Richard Wright

Coleoptera Quedius fuliginosus 9 SO90259456 2011 Richard Wright

Coleoptera Quedius maurorufus 5 SO89599442 2011 Richard Wright



Coleoptera Quedius molochinus 7 SO90309478 2011 Richard Wright

Coleoptera Rhagonycha fulva 3 SO89529444 2011 Richard Wright

Coleoptera Rhagonycha limbata 7 SO90309478 2011 Richard Wright

Coleoptera Rhyzobius litura 1 SO89709483 2011 Richard Wright




Coleoptera Rugilus rufipes 2 SO89629479 2011 Richard Wright

Coleoptera Rutpela maculata 9a SO90269452 20/06/2011 Ian Cheeseborough

Coleoptera Sitona cambricus 2 SO89629479 2011 Richard Wright

Coleoptera Sitona lineatus 3 SO89529444 2011 Richard Wright

Coleoptera Sitona lineatus 9 SO90259456 2011 Richard Wright

Coleoptera Sphaeroderma testaceum 3 SO89529444 2011 Richard Wright

Coleoptera Sphaeroderma testaceum 8 SO90379448 2011 Richard Wright

Coleoptera Stenus clavicornis 1 SO89709483 2011 Richard Wright







Coleoptera Stenus flavipes 2 SO89629479 2011 Richard Wright

Coleoptera Stenus flavipes 5 SO89599442 2011 Richard Wright

Coleoptera Stenus providus 5 SO89599442 2011 Richard Wright

Coleoptera Sunius propinquus 3 SO89529444 2011 Richard Wright

Coleoptera Sunius propinquus 6 SO89799476 2011 Richard Wright

Coleoptera Tachinus rufipes 2 SO89629479 2011 Richard Wright





Coleoptera Tachyporus chrysomelinus 1 SO89709483 2011 Richard Wright



Coleoptera Tachyporus dispar 5 SO89599442 2011 Richard Wright



Coleoptera Tachyporus hypnorum 1 SO89709483 2011 Richard Wright








Coleoptera Tachyporus solutus 2 SO89629479 2011 Richard Wright



Coleoptera Trachyphloeus asperatus 4 SO89459440 2011 Richard Wright

Coleoptera Trachyphloeus asperatus 6 SO89799476 2011 Richard Wright

Coleoptera Trixagus dermestoides 3 SO89529444 2011 Richard Wright



Coleoptera Xantholinus linearis 3 SO89529444 2011 Richard Wright

Coleoptera Xantholinus longiventris 2 SO89629479 2011 Richard Wright

Coleoptera Xantholinus longiventris 8 SO90379448 2011 Richard Wright

Diplopoda Cylindroiulus punctatus 6 SO89799476 2011 Richard Wright

Diplopoda Cylindroiulus punctatus 7 SO90309478 2011 Richard Wright

Diplopoda Ophyiulus pilosus 5 SO89599442 2011 Richard Wright




Diplopoda Polydesmus angustus 1 SO89709483 2011 Richard Wright





Diplopoda Tachypodoiulus niger 1 SO89709483 2011 Richard Wright








Diptera Agathomyia antennata 8 SO90379448 20/05/2011 Nigel Jones

Diptera Anomoia purmunda site record SO898945 28/07/2011 Nigel Jones

Diptera Anthomyia procellaris 9 SO90259456 20/05/2011 Nigel Jones

Diptera Argyra argentina 8 SO90379448 28/07/2011 Nigel Jones

Diptera Beris chalybata 5 SO89599442 20/05/2011 Nigel Jones

Diptera Beris chalybata 7 SO90309478 20/05/2011 Nigel Jones

Diptera Beris morrisii 8 SO90379448 28/07/2011 Nigel Jones

Diptera Beris vallata 8 SO90379448 28/07/2011 Nigel Jones

Diptera Bibio johannis 1 SO89709483 18/04/2011 Nigel Jones

Diptera Bibio johannis 5 SO89599442 18/04/2011 Nigel Jones

Diptera Bibio marci site record SO898945 18/04/2011 Nigel Jones

Diptera Bibio marci 7 SO90029443 20/04/2011 Pete Boardman

Diptera Neolimnomyia nemoralis 5 SO89599442 20/05/2011 Pete Boardman




Diptera Campsicnemus curvipes 8 SO90379448 18/04/2011 Nigel Jones



Diptera Chaetostomella cylindrica 1 SO89709483 20/05/2011 Nigel Jones

Diptera Chaetostomella cylindrica site record SO898945 28/07/2011 Nigel Jones

Diptera Chamaepsila rosae 5 SO89599442 20/05/2011 Nigel Jones

Diptera Chamaepsila rosae 8 SO90379448 20/05/2011 Nigel Jones

Diptera Cheilosia albitarsis 9 SO90259456 20/05/2011 Nigel Jones

Diptera Cheilosia illustrata 9a SO90269452 20/06/2011 Ian Cheeseborough

Diptera Cheilosia illustrata site record SO898945 28/07/2011 Nigel Jones

Diptera Cheilotrichia cinerascens 5 SO89599442 20/04/2011 Pete Boardman




Diptera Chelifera precatoria 5 SO89599442 20/05/2011 Nigel Jones

Diptera Chrysogaster solstitialis site record SO898945 28/07/2011 Nigel Jones

Diptera Chrysopilus cristatus 5 SO89599442 20/05/2011 Nigel Jones

Diptera Chrysopilus cristatus 8 SO90379448 20/05/2011 Nigel Jones

Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman

Diptera Chrysopilus cristatus 9 SO90259456 29/06/2011 Pete Boardman

Diptera Chrysotoxum festivum site record SO898945 28/07/2011 Nigel Jones

Diptera Chrysotus gramineus site record SO898945 28/07/2011 Nigel Jones

Diptera Cleigastra apicalis 9 SO90259456 18/04/2011 Nigel Jones

Diptera Clusiodes albimanus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Clusiodes gentilis 8 SO90379448 20/05/2011 Nigel Jones

Diptera Coenia palustris 8 SO90379448 28/07/2011 Nigel Jones

Diptera Coenosia femoratus site record SO898945 28/07/2011 Nigel Jones

Diptera Coenosia infantula site record SO898945 28/07/2011 Nigel Jones

Diptera Coenosia tigrina site record SO898945 20/05/2011 Nigel Jones

Diptera Coenosia tigrina 9 SO90259456 28/07/2011 Nigel Jones



Diptera Conops quadrifasciatus site record SO898945 28/07/2011 Nigel Jones

Diptera Dasysyrphus venustus 8 SO90379448 20/05/2011 Nigel Jones

Diptera Dicranomyia affinis 5 SO89599442 20/05/2011 Pete Boardman

Diptera Dicranomyia chorea 5 SO89599442 20/04/2011 Pete Boardman



Diptera Dicranota pavida 5 SO89599442 20/05/2011 Pete Boardman

Diptera Dicranota subtilis 5 SO89599442 20/05/2011 Pete Boardman

Diptera Dioctria atricapilla 8 SO90379448 20/05/2011 Nigel Jones

Diptera Dioctria baumhaueri 8 SO90379448 29/06/2011 Pete Boardman

Diptera Ditrichophora calceata 8 SO90379448 28/07/2011 Nigel Jones

Diptera Dolichocephala irrorata 8 SO90379448 18/04/2011 Nigel Jones

Diptera Dolichopus brevipennis 2 SO89629479 20/05/2011 Nigel Jones



Diptera Dolichopus discifer 8 SO90379448 28/07/2011 Nigel Jones

Diptera Dolichopus picipes 5 SO89599442 20/05/2011 Nigel Jones

Diptera Dolichopus plumipes 5 SO89599442 20/05/2011 Nigel Jones





Diptera Dolichopus popularis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Dolichopus popularis 8 SO90379448 28/07/2011 Nigel Jones

Diptera Dolichopus trivialis 5 SO89599442 20/05/2011 Nigel Jones




Diptera Dolichopus ungulatus 5 SO89599442 20/05/2011 Nigel Jones

Diptera Dolichopus ungulatus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Elgiva cucularia 1 SO89709483 20/05/2011 Nigel Jones

Diptera Elgiva solicita 1 SO89709483 20/05/2011 Nigel Jones

Diptera Eloeophila maculata 5 SO89599442 20/05/2011 Pete Boardman

Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Nigel Jones

Diptera Eloeophila submarmorata 5 SO89599442 20/05/2011 Pete Boardman

Diptera Empis aestiva 1 SO89709483 18/04/2011 Nigel Jones

Diptera Empis livida 7 SO90309478 28/07/2011 Nigel Jones

Diptera Empis livida site record SO898945 28/07/2011 Nigel Jones

Diptera Empis nigripes 9 SO90259456 18/04/2011 Nigel Jones



Diptera Empis praevia 5 SO89599442 20/05/2011 Nigel Jones

Diptera Empis stercorea 5 SO89599442 20/05/2011 Nigel Jones

Diptera Empis tessellata site record SO898945 20/05/2011 Nigel Jones

Diptera Epistrophe eligans 8 SO90379448 20/04/2011 Pete Boardman

Diptera Epistrophe eligans site record SO898945 20/05/2011 Nigel Jones

Diptera Epistrophe grossulariae 8 SO90379448 28/07/2011 Nigel Jones

Diptera Episyrphus balteatus site record SO898945 20/05/2011 Nigel Jones

Diptera Episyrphus balteatus 9 SO90259456 29/06/2011 Pete Boardman




Diptera Erioconopa trivialis 9 SO90259456 18/04/2011 Pete Boardman




Diptera Erioptera fusculenta 9 SO90259456 29/06/2011 Pete Boardman

Diptera Erioptera squalida 9 SO90259456 29/06/2011 Pete Boardman

Diptera Eriothrix rufomaculata site record SO898945 28/07/2011 Nigel Jones

Diptera Eristalis arbustorum site record SO898945 20/05/2011 Nigel Jones

Diptera Eristalis intricarius 9 SO90259456 20/04/2011 Pete Boardman

Diptera Eristalis pertinax 8 SO90379448 18/04/2011 Nigel Jones

Diptera Eristalis pertinax 9 SO90259456 20/04/2011 Pete Boardman

Diptera Eristalis tenax site record SO898945 20/05/2011 Nigel Jones

Diptera Eristalis tenax 8 SO90379448 28/07/2011 Nigel Jones

Diptera Eristalis tenax 1 SO89709483 15/09/2011 Pete Boardman

Diptera Eristalis tenax 9 SO90259456 15/09/2011 Pete Boardman

Diptera Eudasyphora cyanella site record SO898945 28/07/2011 Nigel Jones

Diptera Eumerus funeralis site record SO898945 20/06/2011 Nigel Jones

Diptera Eupeodes latifasciatus 1 SO89709483 20/05/2011 Nigel Jones

Diptera Eupeodes luniger site record SO898945 20/06/2011 Nigel Jones

Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones

Diptera Fannia armata 5 SO89599442 20/05/2011 Nigel Jones

Diptera Fannia lustrator 8 SO90379448 20/05/2011 Nigel Jones

Diptera Fannia serena 5 SO89599442 20/05/2011 Nigel Jones

Diptera Fannia similis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Geomyza tripunctata 1 SO89709483 18/04/2011 Nigel Jones

Diptera Gloma fuscipennis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Gloma fuscipennis 8 SO90379448 20/05/2011 Nigel Jones

Diptera Helina impuncta 5 SO89599442 20/05/2011 Nigel Jones

Diptera Helina impuncta 8 SO90379448 28/07/2011 Nigel Jones

Diptera Helina reversio 5 SO89599442 28/07/2011 Nigel Jones

Diptera Helina reversio 9 SO90259456 28/07/2011 Nigel Jones

Diptera Helophilus hybridus 1 SO89709483 28/07/2011 Nigel Jones

Diptera Helophilus pendulus 8 SO90379448 18/04/2011 Nigel Jones


Diptera Helophilus pendulus 8 SO90379448 20/05/2011 Nigel Jones

Diptera Helophilus pendulus site record SO898945 20/05/2011 Nigel Jones

Diptera Helophilus pendulus 9 SO90259456 29/06/2011 Pete Boardman

Diptera Hercostomus aerosus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Hercostomus cupreus 1 SO89709483 20/05/2011 Nigel Jones

Diptera Hercostomus cupreus 5 SO89599442 20/05/2011 Nigel Jones

Diptera Hercostomus germanus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Hercostomus parvilamellatus 5 SO89599442 20/05/2011 Nigel Jones

Diptera Herina frondescentiae 9 SO90259456 20/05/2011 Nigel Jones

Diptera Herina frondescentiae site record SO898945 28/07/2011 Nigel Jones

Diptera Hilara curtisi 8 SO90379448 20/05/2011 Nigel Jones

Diptera Hilara litorea 8 SO90379448 28/07/2011 Nigel Jones

Diptera Hilara lurida 8 SO90379448 28/07/2011 Nigel Jones

Diptera Hybos femoratus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Hydrellia maura 5 SO89599442 20/05/2011 Nigel Jones





Diptera Hydromya dorsalis 8 SO90379448 18/04/2011 Nigel Jones


Diptera Hydromya dorsalis 5 SO89599442 20/04/2011 Pete Boardman


Diptera Hydrotaea cyrtoneurina 5 SO89599442 20/05/2011 Nigel Jones

Diptera Hydrotaea cyrtoneurina 8 SO90379448 28/07/2011 Nigel Jones

Diptera Ilisia maculata 8 SO90379448 20/05/2011 Nigel Jones

Diptera Ilisia maculata 9 SO90259456 20/05/2011 Pete Boardman

Diptera Jassidophaga villosa 9 SO90259456 18/04/2011 Nigel Jones

Diptera Lauxania cylindricornis site record SO898945 20/05/2011 Nigel Jones

Diptera Leptogaster cylindrica site record SO901944 20/06/2011 Ian Cheeseborough

Diptera Leucophora obtusa 8a SO903945 18/04/2011 Nigel Jones

Diptera Limnellia quadrata 9 SO90259456 18/04/2011 Nigel Jones

Diptera Limnia paludicloa 9 SO90259456 28/07/2011 Nigel Jones

Diptera Limnia unguicornis 8 SO90379448 20/05/2011 Nigel Jones

Diptera Limonia flavipes 5 SO89599442 20/05/2011 Pete Boardman

Diptera Limonia macrostigma 8 SO90379448 20/05/2011 Pete Boardman

Diptera Limonia nubeculosa 8 SO90379448 20/05/2011 Pete Boardman

Diptera Limonia nubeculosa 9 SO90259456 15/09/2011 Pete Boardman

Diptera Limonia phragmitidis 8 SO90379448 20/05/2011 Pete Boardman

Diptera Lonchaea chorea 8 SO90379448 20/05/2011 Nigel Jones

Diptera Lonchoptera lutea 5 SO89599442 18/04/2011 Nigel Jones

Diptera Lonchoptera lutea 7 SO90309478 18/04/2011 Nigel Jones

Diptera Loxocera albiseta 8 SO90379448 28/07/2011 Nigel Jones

Diptera Loxocera albiseta 9 SO90259456 28/07/2011 Nigel Jones

Diptera Lydella stabulans site record SO898945 28/07/2011 Nigel Jones

Diptera Machimus cingulatus site record SO898945 28/07/2011 Nigel Jones

Diptera Meigenia mutabilis 8 SO90379448 28/07/2011 Nigel Jones

Diptera Meiosimyza decipiens site record SO898945 28/07/2011 Nigel Jones

Diptera Meiosimyza rorida site record SO898945 28/07/2011 Nigel Jones

Diptera Melanogaster hirtella 9 SO90259456 20/05/2011 Nigel Jones

Diptera Melanostoma mellinum 9 SO90259456 28/07/2011 Nigel Jones

Diptera Melanostoma scalare 5 SO89599442 18/04/2011 Nigel Jones






Diptera Merodon equestris 8 SO90379448 20/05/2011 Nigel Jones

Diptera Mesembrina meridiana 9 SO90259456 15/09/2011 Pete Boardman


Diptera Microchrysa flavicornis 8 SO90379448 28/07/2011 Nigel Jones

Diptera Microchrysa polita 8 SO90379448 20/05/2011 Nigel Jones

Diptera Microphor holosericeus 5 SO89599442 20/05/2011 Nigel Jones

Diptera Minettia fasciata site record SO898945 28/07/2011 Nigel Jones

Diptera Minettia inusta 8 SO90379448 28/07/2011 Nigel Jones

Diptera Minettia longipennis site record SO898945 20/05/2011 Nigel Jones

Diptera Minettia longipennis site record SO898945 28/07/2011 Nigel Jones

Diptera Molophilus cinereifrons 5 SO89599442 20/05/2011 Pete Boardman

Diptera Molophilus obscurus 1 SO89709483 20/05/2011 Pete Boardman

Diptera Molophilus obscurus 9 SO90259456 29/06/2011 Pete Boardman

Diptera Morellia hortorum 8 SO90379448 28/07/2011 Nigel Jones

Diptera Muscina levida 8 SO90379448 28/07/2011 Nigel Jones

Diptera Myathropa florea site record SO898945 20/05/2011 Nigel Jones

Diptera Nanna fasciata 2 SO89629479 18/04/2011 Nigel Jones

Diptera Nanna flavipes 2 SO89629479 18/04/2011 Nigel Jones

Diptera Neoascia podagrica 5 SO89599442 18/04/2011 Nigel Jones

Diptera Neoascia podagrica 5 SO89599442 28/07/2011 Nigel Jones

Diptera Nephrotoma appendiculata 8 SO90379448 20/05/2011 Pete Boardman

Diptera Nephrotoma appendiculata 9 SO90259456 29/06/2011 Pete Boardman

Diptera Nephrotoma quadrifaria 9 SO90259456 29/06/2011 Pete Boardman

Diptera Neria cibaria 8 SO90379448 20/05/2011 Nigel Jones

Diptera Neuroctena anilis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Notiphila maculata 9 SO90259456 28/07/2011 Nigel Jones

Diptera Ocydromia glabricula 5 SO89599442 20/05/2011 Nigel Jones

Diptera Oedalea stigmatella 8 SO90379448 20/05/2011 Nigel Jones

Diptera Opomyza germinationis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Opomyza petrei 8 SO90379448 28/07/2011 Nigel Jones

Diptera Ormosia lineata 11 SO89649435 20/04/2011 Pete Boardman



Diptera Oxycera rara 9 SO90259456 29/06/2011 Pete Boardman

Diptera Pachygaster leachii 8 SO90379448 28/07/2011 Nigel Jones

Diptera Palloptera scutellata 1 SO89709483 18/04/2011 Nigel Jones




Diptera Parydra coarctata 5 SO89599442 20/05/2011 Nigel Jones

Diptera Parydra coarctata 9 SO90259456 28/07/2011 Nigel Jones

Diptera Pelatachina tibialis site record SO898945 28/07/2011 Nigel Jones

Diptera Phaonia errans 8 SO90379448 28/07/2011 Nigel Jones

Diptera Phaonia fuscata site record SO898945 28/07/2011 Nigel Jones

Diptera Phaonia subventa 8 SO90379448 28/07/2011 Nigel Jones

Diptera Phasia obesa site record SO898945 20/05/2011 Nigel Jones

Diptera Phasia pusilla 5 SO89599442 20/05/2011 Nigel Jones

Diptera Pherbellia schoenherri site record SO898945 28/07/2011 Nigel Jones

Diptera Pilaria discicollis 9 SO90259456 29/06/2011 Pete Boardman

Diptera Platycheirus clypeatus 9 SO90259456 28/07/2011 Nigel Jones

Diptera Platycheirus granditarsus 9 SO90259456 28/07/2011 Nigel Jones

Diptera Platycheirus manicatus 9 SO90259456 20/05/2011 Nigel Jones

Diptera Platycheirus peltatus 9 SO90259456 28/07/2011 Nigel Jones

Diptera Platycheirus tarsalis 9 SO90259456 20/05/2011 Nigel Jones

Diptera Platycheirus tarsalis 5 SO89599442 28/07/2011 Nigel Jones

Diptera Platypalpus longicornis 8 SO90379448 20/05/2011 Nigel Jones

Diptera Poecilobothrus nobilitatus 9 SO90259456 29/06/2011 Pete Boardman

Diptera Poecilobothrus nobilitatus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Portevinia maculata 8 SO90379448 20/05/2011 Nigel Jones

Diptera Psila fimetaria site record SO898945 20/05/2011 Nigel Jones

Diptera Ptychoptera albimana 5 SO89599442 20/04/2011 Pete Boardman





Diptera Rhagio lineola 8 SO90379448 28/07/2011 Nigel Jones

Diptera Rhagio scolopaceus 8 SO90379448 20/05/2011 Nigel Jones

Diptera Rhamphomyia albipennis 1 SO89709483 18/04/2011 Nigel Jones

Diptera Rhamphomyia albohirta 5 SO89599442 18/04/2011 Nigel Jones

Diptera Rhamphomyia albohirta 8 SO90379448 20/05/2011 Nigel Jones

Diptera Rhamphomyia nigripennis 5 SO89599442 20/05/2011 Nigel Jones

Diptera Rhamphomyia subcinerascens 8 SO90379448 18/04/2011 Nigel Jones

Diptera Rhamphomyia subcinerascens 9 SO90259456 18/04/2011 Nigel Jones

Diptera Rhamphomyia sulcata 7 SO90309478 18/04/2011 Nigel Jones

Diptera Rhamphomyia umbripennis 5 SO89599442 18/04/2011 Nigel Jones

Diptera Rhamphomyia umbripennis 7 SO90309478 18/04/2011 Nigel Jones

Diptera Rhaphium brevicorne 8 SO90379448 28/07/2011 Nigel Jones

Diptera Rhaphium caliginosum 8 SO90379448 18/04/2011 Nigel Jones

Diptera Rhingia campestris 7 SO90029443 20/04/2011 Pete Boardman

Diptera Rhingia campestris 8 SO90379448 20/05/2011 Nigel Jones

Diptera Sapromyza quadricincta 8 SO90379448 20/05/2011 Nigel Jones

Diptera Sapromyza sexpunctata site record SO898945 28/07/2011 Nigel Jones

Diptera Scaeva pyrastri 8a SO903945 28/07/2011 Ian Cheeseborough

Diptera Scatella tenuicosta 9 SO90259456 28/07/2011 Nigel Jones

Diptera Scathophaga furcata site record SO898945 28/07/2011 Nigel Jones

Diptera Scathophaga lutaria 8 SO90379448 18/04/2011 Nigel Jones

Diptera Scathophaga stercoraria site record SO898945 20/05/2011 Nigel Jones

Diptera Sepedon sphegea 1 SO89709483 18/04/2011 Nigel Jones

Diptera Sepedon sphegea 1 SO89709483 20/04/2011 Pete Boardman

Diptera Sepsis fulgens site record SO898945 28/07/2011 Nigel Jones

Diptera Sepsis punctum site record SO898945 28/07/2011 Nigel Jones

Diptera Sicus ferrugineus site record SO898945 20/06/2011 Nigel Jones

Diptera Siphona geniculata 8 SO90379448 28/07/2011 Nigel Jones

Diptera Spania nigra 5 SO89599442 20/05/2011 Nigel Jones

Diptera Sphaerophoria interrupta site record SO898945 28/07/2011 Nigel Jones

Diptera Sphaerophoria scripta 9 SO90259456 15/09/2011 Pete Boardman

Diptera Sphaerophoria taeniata 2 SO89629479 28/07/2011 Nigel Jones

Diptera Sphegina clunipes 8 SO90379448 20/05/2011 Nigel Jones

Diptera Sybistroma obscurellum 8 SO90379448 28/07/2011 Nigel Jones

Diptera Sympycnus desoutteri 5 SO89599442 20/05/2011 Nigel Jones

Diptera Sympycnus desoutteri 9 SO90259456 20/05/2011 Nigel Jones

Diptera Syntormon macula 8 SO90379448 18/04/2011 Nigel Jones

Diptera Syntormon macula 8 SO90379448 20/05/2011 Nigel Jones

Diptera Syritta pipiens site record SO898945 20/05/2011 Nigel Jones

Diptera Syrphus vitripennis site record SO898945 28/07/2011 Nigel Jones

Diptera Tachina fera 9a SO90269452 20/05/2011 Ian Cheeseborough

Diptera Tachina fera 9 SO90259456 15/09/2011 Pete Boardman

Diptera Tephritis cometa 9 SO90259456 20/05/2011 Nigel Jones

Diptera Terellia ruficauda 8 SO90379448 28/07/2011 Nigel Jones

Diptera Terellia ruficauda 9 SO90259456 28/07/2011 Nigel Jones

Diptera Tetanocera elata site record SO898945 28/07/2011 Nigel Jones

Diptera Tetanocera ferruginea site record SO898945 28/07/2011 Nigel Jones

Diptera Tetanocera fuscinervis 8 SO90379448 20/05/2011 Nigel Jones

Diptera Teuchophorus calcaratus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Teuchophorus monacanthus 9 SO90259456 20/05/2011 Nigel Jones

Diptera Teuchophorus monacanthus 8 SO90379448 28/07/2011 Nigel Jones

Diptera Teuchophorus nigricosta 8 SO90379448 28/07/2011 Nigel Jones

Diptera Teuchophorus simplex 8 SO90379448 28/07/2011 Nigel Jones

Diptera Themira annulipes site record SO898945 28/07/2011 Nigel Jones

Diptera Thricops diaphana 8 SO90379448 28/07/2011 Nigel Jones

Diptera Tipula fascipennis 9 SO90259456 29/06/2011 Pete Boardman

Diptera Tipula lateralis 11 SO89649435 20/04/2011 Pete Boardman


Diptera Tipula lateralis 9 SO90259456 29/06/2011 Pete Boardman

Diptera Tipula paludosa 1 SO89709483 15/09/2011 Pete Boardman



Diptera Tipula paludosa site record SO898945 15/09/2011 Pete Boardman

Diptera Tipula submarmorata site record SO89399444 20/05/2011 Pete Boardman

Diptera Tipula vittata 9 SO90259456 18/04/2011 Pete Boardman





Diptera Trichopsomyia flavitarsis 9 SO90259456 20/05/2011 Nigel Jones

Diptera Tricyphona immaculata 5 SO89599442 20/04/2011 Pete Boardman



Diptera Urophora cardui site record SO898945 28/07/2011 Nigel Jones

Diptera Urophora jaceana 8 SO90379448 29/06/2011 Pete Boardman

Diptera Urophora jaceana site record SO898945 28/07/2011 Nigel Jones

Diptera Vibrissina debilitata site record SO898945 28/07/2011 Nigel Jones

Diptera Volucella bombylans site record SO898945 20/05/2011 Nigel Jones

Diptera Xyphosia miliaria site record SO898945 28/07/2011 Nigel Jones

Hemiptera Acanthosoma haemorrhoidale 1 SO89709483 15/09/2011 Pete Boardman

Hemiptera Capsus ater 3 SO89529444 2011 Richard Wright

Hemiptera Coreus marginatus 9a SO90269452 20/06/2011 Ian Cheeseborough

Hemiptera Deraeocoris lutescens 7 SO90309478 2011 Richard Wright

Hemiptera Elasmostethus interstinctus site record SO898945 20/05/2011 Nigel Jones

Hemiptera Himacerus mirmicoides 4 SO89459440 2011 Richard Wright

Hemiptera Ischnodemus sabuleti 5 SO89599442 2011 Richard Wright

Hemiptera Leptopterna dolabrata 2 SO89629479 2011 Richard Wright

Hemiptera Leptopterna dolabrata 3 SO89529444 2011 Richard Wright

Hemiptera Liocoris tripustulatus 7 SO90309478 2011 Richard Wright

Hemiptera Nabis rugosus 2 SO89629479 2011 Richard Wright






Hemiptera Notostira elongata 3 SO89529444 2011 Richard Wright

Hemiptera Orthotylus ericetorum 4 SO89459440 2011 Richard Wright

Hemiptera Orthotylus ericetorum 6 SO89799476 2011 Richard Wright

Hemiptera Palomena prasina 7 SO90309478 2011 Richard Wright

Hemiptera Palomena prasina site record SO899945 18/04/2011 Ian Cheeseborough

Hemiptera Palomena prasina 9a SO90269452 20/05/2011 Ian Cheeseborough

Hemiptera Piezodorus lituratus site record SO902948 18/04/2011 Ian Cheeseborough

Hemiptera Pithanus maerkelii 9 SO90259456 29/06/2011 Pete Boardman

Hemiptera Scolopostethus decoratus 4 SO89459440 2011 Richard Wright

Hemiptera Scolopostethus decoratus 6 SO89799476 2011 Richard Wright

Hemiptera Stenodema calcarata 3 SO89529444 2011 Richard Wright

Hemiptera Stenotus binotatus 3 SO89529444 2011 Richard Wright

Hemiptera Stygnocoris pedestris 3 SO89529444 2011 Richard Wright

Hemiptera Stygnocoris rusticus 4 SO89459440 2011 Richard Wright



Hemiptera Velia caprai 5 SO89599442 2011 Richard Wright

Hymenoptera Ancistrocerus gazella 8a SO903944 28/07/2011 Ian Cheeseborough

Hymenoptera Andrena barbilabris 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena bicolor site record SO899948 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena bicolor 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Andrena chrysosceles 12 SO90349456 18/04/2011 Ian Cheeseborough


Hymenoptera Andrena cineraria 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena cineraria site record SO899948 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena dorsata 8a SO903944 28/07/2011 Ian Cheeseborough

Hymenoptera Andrena fucata 8a SO903944 20/05/2011 Ian Cheeseborough

Hymenoptera Andrena fulva 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena fulva 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena fulva site record SO899945 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena haemorrhoa 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena haemorrhoa 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena haemorrhoa site record SO899945 18/04/2011 Ian Cheeseborough



Hymenoptera Andrena nigroaenea 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena nigroaenea site record SO901944 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena scotica 8a SO903945 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena scotica site record SO899945 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena scotica site record SO901944 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena subopaca 8a SO903944 18/04/2011 Ian Cheeseborough

Hymenoptera Andrena wilkella 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Andrena wilkella 8a SO903944 28/07/2011 Ian Cheeseborough

Hymenoptera Anoplius caviventris site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Anoplius nigerrimus 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Anthidium manicatum 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Anthophora furcata 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Anthophora furcata 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Argogorytes mystaceus 8a SO903944 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus hortorum site record SO899943 29/06/2011 Ian Cheeseborough

Hymenoptera Bombus hypnorum 9a SO90269452 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus hypnorum site record SO898945 20/05/2011 Nigel Jones

Hymenoptera Bombus hypnorum 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Bombus lapidarius site record SO901944 18/04/2011 Ian Cheeseborough

Hymenoptera Bombus lapidarius site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus lapidarius 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Bombus lapidarius 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Bombus lucorum 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Bombus lucorum 9a SO90269452 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus lucorum 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Bombus lucorum 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Bombus pascuorum 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Bombus pascuorum site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus pascuorum 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Bombus pascuorum 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Bombus pratorum 9a SO90269452 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus pratorum site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus pratorum 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Bombus sylvestris 9a SO90269452 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus sylvestris site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus terrestris 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Bombus terrestris site record SO899945 18/04/2011 Ian Cheeseborough

Hymenoptera Bombus terrestris 9a SO90269452 20/05/2011 Ian Cheeseborough

Hymenoptera Bombus terrestris 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Bombus vestalis site record SO899945 18/04/2011 Ian Cheeseborough

Hymenoptera Crossocerus annulipes 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Crossocerus capitosus 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Crossocerus cetratus 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Crossocerus megacephalus 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Crossocerus nigritus 8a SO903944 20/05/2011 Ian Cheeseborough

Hymenoptera Crossocerus podagricus 9a SO90269452 28/07/2011 Ian Cheeseborough


Hymenoptera Crossocerus quadrimaculatus 8 SO90379448 20/06/2011 Ian Cheeseborough

Hymenoptera Dolichovespula norwegica 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Dolichovespula sylvestris 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Ectemnius cavifrons 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Ectemnius continuus 8a SO903944 20/05/2011 Ian Cheeseborough



Hymenoptera Halictus rubicundus 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Halictus tumulorum 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Hylaeus communis 9 SO90259456 28/07/2011 Nigel Jones

Hymenoptera Hylaeus communis 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Lasioglossum calceatum 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Lasioglossum leucopus 8a SO903945 28/07/2011 Ian Cheeseborough

Hymenoptera Lasioglossum villosulum 8a SO903945 18/04/2011 Ian Cheeseborough

Hymenoptera Lasius flavus 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Lasius niger site record SO899948 18/04/2011 Ian Cheeseborough

Hymenoptera Lindenius albilabris site record SO899943 29/06/2011 Ian Cheeseborough

Hymenoptera Lindenius albilabris 8a SO903944 28/07/2011 Ian Cheeseborough

Hymenoptera Megachile ligniseca 8a SO903945 29/06/2011 Ian Cheeseborough

Hymenoptera Megachile willughbiella 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Megachile willughbiella 10 SO89639441 28/07/2011 Ian Cheeseborough

Hymenoptera Megachile willughbiella 8a SO903944 28/07/2011 Ian Cheeseborough

Hymenoptera Mimumesa dahlbomi 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Myrmosa atra site record SO899943 29/06/2011 Ian Cheeseborough

Hymenoptera Nomada fabriciana 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Nomada flava site record SO901944 18/04/2011 Ian Cheeseborough

Hymenoptera Nomada flava/panzeri 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Nomada goodeniana 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Nomada lathburiana 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Nomada ruficornis 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Nomada ruficornis 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Osmia caerulescens 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Osmia rufa 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Osmia rufa site record SO899945 18/04/2011 Ian Cheeseborough

Hymenoptera Osmia rufa site record SO901944 20/06/2011 Ian Cheeseborough

Hymenoptera Passaloecus gracilis 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Passaloecus singularis 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Pemphredon lugubris 8 SO90379448 20/05/2011 Nigel Jones

Hymenoptera Pemphredon lugubris 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Psenulus concolor 8a SO903944 20/05/2011 Ian Cheeseborough

Hymenoptera Psenulus pallipes 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Rhopalum clavipes 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Rhopalum clavipes site record SO898945 28/07/2011 Nigel Jones

Hymenoptera Sphecodes ephippius 12 SO90349456 18/04/2011 Ian Cheeseborough

Hymenoptera Stigmus solskyi 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Tiphia minuta 8a SO903945 29/06/2011 Ian Cheeseborough

Hymenoptera Trichrysis cyanea 9a SO90269452 20/06/2011 Ian Cheeseborough

Hymenoptera Trypoxylon attenuatum 9a SO90269452 20/05/2011 Ian Cheeseborough



Hymenoptera Trypoxylon clavicerum 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Vespula germanica 7 SO90309478 18/04/2011 Ian Cheeseborough

Hymenoptera Vespula germanica 9a SO90269452 28/07/2011 Ian Cheeseborough

Hymenoptera Vespula rufa site record SO898945 20/05/2011 Ian Cheeseborough

Hymenoptera Vespula rufa 8a SO903944 29/06/2011 Ian Cheeseborough

Hymenoptera Vespula vulgaris 11 SO89649435 18/04/2011 Ian Cheeseborough

Hymenoptera Vespula vulgaris 8a SO903945 29/06/2011 Ian Cheeseborough

Isopoda Oniscus asellus 7 SO90309478 2011 Richard Wright

Isopoda Philoscia muscorum 1 SO89709483 2011 Richard Wright








Isopoda Trichoniscus pusillus 1 SO89709483 2011 Richard Wright




Lepidoptera Aglais urticae 8 SO90379448 20/04/2011 Pete Boardman

Lepidoptera Aglais urticae 9a SO90269452 20/05/2011 Ian Cheeseborough

Lepidoptera Aglais urticae 9 SO90259456 29/06/2011 Pete Boardman

Lepidoptera Anthocharis cardamines site record SO902948 18/04/2011 Ian Cheeseborough

Lepidoptera Anthocharis cardamines 8 SO90379448 20/04/2011 Pete Boardman

Lepidoptera Anthocharis cardamines 9 SO90259456 20/04/2011 Pete Boardman

Lepidoptera Aphantopus hyperantus 9a SO90269452 20/06/2011 Ian Cheeseborough

Lepidoptera Aphantopus hyperantus 9 SO90259456 29/06/2011 Pete Boardman

Lepidoptera Aphantopus hyperantus 8a SO903945 28/07/2011 Ian Cheeseborough

Lepidoptera Callophrys rubi 9a SO90269452 20/05/2011 Ian Cheeseborough

Lepidoptera Coenonympha pamphilus 9a SO90269452 20/05/2011 Ian Cheeseborough

Lepidoptera Coenonympha pamphilus 9a SO90269452 20/06/2011 Ian Cheeseborough

Lepidoptera Inachis io 8 SO90379448 20/04/2011 Pete Boardman

Lepidoptera Inachis io 9a SO90269452 20/06/2011 Ian Cheeseborough

Lepidoptera Inachis io 8a SO903944 28/07/2011 Ian Cheeseborough

Lepidoptera Lycaena phlaeas 9a SO90269452 28/07/2011 Ian Cheeseborough

Lepidoptera Maniola jurtina 9 SO90259456 29/06/2011 Pete Boardman

Lepidoptera Ochlodes venata 9a SO90269452 20/06/2011 Ian Cheeseborough

Lepidoptera Ochlodes venata 9 SO90259456 29/06/2011 Pete Boardman

Lepidoptera Pararge aegeria 7 SO90309478 20/04/2011 Pete Boardman

Lepidoptera Pararge aegeria 9a SO90269452 20/05/2011 Ian Cheeseborough



Lepidoptera Pieris napi 8 SO90379448 20/04/2011 Pete Boardman

Lepidoptera Pieris napi 9a SO90269452 20/05/2011 Ian Cheeseborough

Lepidoptera Pieris rapae 9a SO90269452 20/06/2011 Ian Cheeseborough

Lepidoptera Pieris rapae 9a SO90269452 28/07/2011 Ian Cheeseborough

Lepidoptera Polygonia c-album 8 SO90379448 20/04/2011 Pete Boardman

Lepidoptera Pyronia tithonus 9a SO90269452 28/07/2011 Ian Cheeseborough

Lepidoptera Stigmella aurella 7 SO90029443 20/04/2011 Pete Boardman

Lepidoptera Thymelicus sylvestris 9a SO90269452 20/06/2011 Ian Cheeseborough

Odonata Coenagrion puella 9a SO90269452 20/05/2011 Ian Cheeseborough

Odonata Coenagrion puella 9a SO90269452 20/06/2011 Ian Cheeseborough

Odonata Ischnura elegans 9 SO90259456 29/06/2011 Pete Boardman

Odonata Pyrrhosoma nymphula 9a SO90269452 20/05/2011 Ian Cheeseborough

Odonata Sympetrum striolatum 1 SO89709483 15/09/2011 Pete Boardman

Opiliones Mitopus morio 2 SO89629479 2011 Richard Wright

Opiliones Nemastoma bimaculatum 2 SO89629479 2011 Richard Wright




Opiliones Paroligolophus agrestis 1 SO89709483 2011 Richard Wright



Opiliones Rilaena triangularis 1 SO89709483 2011 Richard Wright



Orthoptera Omocestus viridulus 9a SO90269452 20/06/2011 Ian Cheeseborough

Orthoptera Tetrix subulata 5 SO89599442 2011 Richard Wright




Orthoptera Tetrix subulata 5 SO89599442 20/04/2011 Pete Boardman

Appendix 2 – ISIS outputs for Penn Common

WHOLE SITE

SAT

code SAT name No. spp. Condition

% of national species

pool

Related BAT rarity

score

F001 scrub edge 19 fav 11

F002 rich flower resource 15 fav 6

A212 bark & sapwood decay 23 fav 5 131

F006 dung 3 3

F112 open short sward 4 2 120

W111 shingle banks 1 2 154

W126 seepage 1 2 154

W314 reedfen and pools 2 2 135

F003 scrub-heath & moorland 5 1

F111 bare sand & chalk 5 1 120

A211 heartwood decay 1 1 131

BAT code BAT name

Representation (1-100)

Rarity score Condition BAT species richness IEC

F2 grassland & scrub matrix 34 115 154

F1

unshaded early successional

mosaic 11 120 51

W3 permanent wet mire 10 135 44

W1 flowing water 8 154 fav 35

A2 wood decay 7 131 30 0

F3 shaded field & ground layer 6 130 29

W2 mineral marsh & open water 5 112 25

A1 arboreal canopy 3 13

CALCAREOUS FLUSH

SAT



pool

Related BAT rarity

score

W111 shingle banks 1 2 132

BAT

code BAT name

Representation

(1-100)

Rarity

score Condition BAT species richness IEC


W1 flowing water 19 132 19


F3 shaded field & ground layer 11 11

W3 permanent wet mire 10 10


WET HABITATS IN EAST OF SITE

SAT



pool

Related BAT rarity

score

W126 seepage 2 4 177

F112 open short sward 3 2

F001 scrub edge 2 1

W314 reedfen and pools 1 1 121

A212 bark & sapwood decay 3 1

F002 rich flower resource 1 0

F003 scrub-heath & moorland 1 0

F111 bare sand & chalk 1 0

BAT

code BAT name

Representation

(1-100)

Rarity

score Condition BAT species richness IEC


W3 permanent wet mire 16 121 40

W1 flowing water 12 177 fav 31



F1 unshaded early successional mosaic 3 8

A2 wood decay 3 8 0


DRY ACID GRASSLAND

SAT



pool

Related BAT rarity

score

F003 scrub-heath & moorland 10 fav 3

F111 bare sand & chalk 3 1

F001 scrub edge 1 1

F112 open short sward 1 1

BAT code BAT name




F1

unshaded early successional

mosaic 5 8

W2 mineral marsh & open water 4 6



W1 flowing water 1 1

DAMP GRASSLAND

SAT code SAT name No. spp. Condition

% of national species pool

Related BAT rarity score

BAT code BAT name







W1 flowing water 2 2


ACULEATE HABITATS

SAT code SAT name No. spp. Condition

% of national species pool

Related BAT rarity score

A212 bark & sapwood decay 26 fav 5 119

F002 rich flower resource 8 3

F006 dung 3 3

F001 scrub edge 5 3

F112 open short sward 2 1 104

BAT code BAT name



F1 unshaded early successional mosaic 29 104 28

A2 wood decay 28 119 27 0




An invertebrate survey of Penn Common (concentrating on...

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