An Integrated Geoecosystem-Remote Sensing Approach To Aspen Ecosystem Management In Northern Lower...

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An Integrated Geoecosystem-Remote Sensing Approach To Aspen Ecosystem Management In Northern Lower Michigan School of Natural Resources & Environment McIntire-Stennis Program Burt Barnes, Kathleen Bergen Graduate Students: Ephraim Zimmerman Kara Moore, Catherine Yanca October 28, 2002 - PowerPoint PPT Presentation

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  • An Integrated Geoecosystem-Remote Sensing Approach To Aspen Ecosystem ManagementIn Northern Lower Michigan

    School of Natural Resources & EnvironmentMcIntire-Stennis Program

    Burt Barnes, Kathleen Bergen

    Graduate Students: Ephraim Zimmerman Kara Moore, Catherine Yanca

    October 28, 2002

    CollaboratorTom CrowUSFS North Central Research Station

    McIntyre-Stennis/Ecosystem Managment Initiative

  • RationaleFollowing massive logging and post-logging fires in northern Lower Michigan in the mid-to-late 19th Century, the original pine and hemlock-northern hardwood forests were changed to stands primarily dominated by bigtooth and trembling aspens.

    Now, 80-100 years later, aspens are declining and are being replaced by other species in patterns determined by site conditions and forest history.

    These changes are not well understood, yet have profound ecological, social, and economic implications.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • GoalDemonstrate the applicability of the landscape ecosystem approach coupled with remote sensing/GIS methods to develop a model of the spatial and successional status of the aspen resource for ecosystem management.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • ObjectivesDetermine the successional pathways for selected ecosystems at the 4000-ha University of Michigan biological Station (UMBS), northern Lower Michigan

    Characterize the role of red maple, northern red oak, and eastern white pine in forming 21st Century forest communities for selected landforms and ecosystems.

    Model the decline of the aspen resource using an ecosystem approach at multiple spatial scales coupled with remote sensing and GIS methods.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Approach

    An ecosystem map of the area and permanent sample plots were used to determine successional change at multiple spatial scales.

    The composition, structure, and successional trends were determined for 7 aspen- dominated ecosystem types

    The occurrence and successional role of species replacing aspen (red maple, northern red oak, and eastern white pine) in diverse ecosystems were determined

    McIntyre-Stennis/Ecosystem Managment Initiative

  • ResultsAspen

    On moraine landforms, aspens are being replaced by northern-hardwood species.

    In contrast, aspens on outwash plain landforms are being replaced by white and red pines and red maple, depending on ecosystem type.

    Different trends of occurrence and succession for red maple, northern red oak, and white pine for a diverse set of ecosystem types was observed and described.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Different Landscape Ecosystems Types, Different Successional PathwaysDryMesicWet-Mesic

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Red MapleSuccess of red maple is highly ecosystem-dependent

    Contrary to the prevailing view, red maple will not become dominant in the long run on dry and dry-mesic ecosystems at UMBS although many of its clones may persist indefinitely.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • 0.010.020.030.040.050.060.070.080.090.0142425363741424445Ecosystem TypeRelative Density (%)Dominant OverstorySubdominant OverstoryDominant Overstory Subdominant Overstory Understory Comparison of red maple relative density in three forest layers for selected ecosystem types, outwash physiographic system, UMBS, Cheboygan and Emmet Co., northern Lower Michigan.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Northern Red OakRed oak understory density was considerably lower than overstory density in most ecosystems types due to:limited light under dense northern hardwood canopy in mesic ecosystems severe site conditions, lack of fire, and heavy deer browsing in dry ecosystem types

    Higher red oak density in outwash-over moraine ecosystem type open, patchy aspen canopy sandy-loam bands in C horizon

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Comparison of northern red oak density in three forest layers for selected ecosystem types, outwash physiographic system, UMBS, Cheboygan and Emmet Co., northern Lower Michigan.

    McIntyre-Stennis/Ecosystem Managment Initiative

    Chart3

    250

    143.703703703725.1851851852

    184.444444444477.7777777778

    177.777777777884.4444444444

    151.75438596492.6315789474

    55.55555555560

    9.523809523860.3174603175

    33.3333333333426.6666666667

    094.4444444444

    48.888888888971.1111111111

    74.444444444427.7777777778

    54.3293.8

    Overstory

    Understory

    Ecosystem Type

    Density (stems/ha)

    OS_US_ecogroups

    Table 3.2. Comparison of northern red oak overstory layer basal area, relative dominance, density,

    and relative density for upland ecosystem groups within outwash landform types UMBS, Cheboygan and Emmet Co., northern Michigan.

    DensityRelative DensityBasal AreaRelative Dom.

    (stems/ha)(%)

    xsdxsdxsdxsd

    GroupEcosystem GroupPlots(n)groupgroupgroup

    2Climatically moderate, heavily burned high-30163.0105.7227.416.127.74.4238.221.0

    elevation, outwash plains and dunes (36, 37, 74)

    4Climatically moderate, calcareous, high-elevation,32112.7155.4411.312.745.16.4417.816.4

    outwash plain (44,45)

    6Lake moderated, glacial lake beach features1061.757.267.26.862.52.079.819.4

    (76 -77)

    7Ice margin terraces(high terraces 96-97)954.389.076.712.172.44.569.77.8

    5Deep outwash over moraine landforms(59, 60)1423.809523809549.663089577953.55144855147.270253363951.412422223.542926200355.260304697412.8339334251

    Climatically extreme, low-elevation

    1outwash plain (Pellston Plain) (1,4)1010.031.611.85.710.82.615.015.7

    3Climatically moderate, Lake-moderated125.613.830.92.330.10.230.81.8

    high-elevation, outwash plain (40,41)

    Total117431.020.1

    a, b, and c indicate significant differences between ecosystem groups;

    values followed by the same letter are not statistically significant (p>0.10).

    Table 3.2. Comparison of northern red oak understory layer basal area, relative dominance, density,

    and relative density for upland ecosystem groups within outwash landform types UMBS, Cheboygan and Emmet Co., northern Michigan.

    DensityRelative Density

    (stems/ha)

    xsdxsd

    GroupEcosystem GroupPlots (n)

    5Deep outwash over moraine landforms14331.7353.530.530.4

    (59, 60)

    7Ice margin terraces(high terraces 96-97)993.8157.07.011.8

    2Climatically moderate, heavily burned high-3052.6128.86.816.3

    elevation, outwash plains and dunes (36, 37, 74)

    6Lake moderated, glacial lake1049.494.75.410.5

    beach features (76 -77)

    3Climatically moderate, Lake-moderated1235.2121.92.27.7

    outwash plain (40,41)

    1Climatically extreme, low-elevation105.015.80.20.6

    outwash plain (Pellston Plain) (1,4)

    4Climatically moderate, calcareous, high-elevation,321.68.80.10.7

    outwash plain (44, 45)

    Total94143.7847222222

    a and b indicate significant differences between physiographic systems;

    values followed by the same letter are not statistically significant (p>0.10).

    OS_US_ecogroups

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    stems/ha

    group

    Stems/ha

    Red oak OS density (ecosystem groups)

    OS_us_physiographic

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    stems/ha

    Group

    Stems/ha

    Red oak understory Density (ecosystem groups)

    ALL_oak_OS_outwash_types

    Table 3.2. Comparison of northern red oak overstory layer basal area, relative dominance, density,

    and relative density for upland moraine and outwash plain landform types UMBS, Cheboygan and Emmet Co., northern Michigan.

    DensityRdenBasal AreaRdom

    (stems/ha)(%)(%)

    xsdxsdxsdxsd

    GroupPhysiographic SystemPlots (n)GroupGroup

    1Outwash Plain (1,36,37,40,41,44,45,11191.0a119.312.6a15.24.2a5.318.3a0.9

    59,60,74,76,77,96,97)

    2Interlobate Moraine (109, 113)2510.2b23.11.1b2.30.5b1.90.9b2.5

    Total136101.24.8

    a and b indicate significant differences between physiographic systems;

    values followed by the same letter are not statistically significant (p>0.10).

    Table 3.2. Comparison of northern red oak understory layer basal area, relative dominance, density,

    and relative density for upland moraine and outwash plain landform types UMBS, Cheboygan and Emmet Co., northern Michigan.

    DensityRden

    (stems/ha)(%)

    xsdxsd

    GroupEcosystem GroupPlots (n)

    1Outwash Plain (1,36,37,40,41,42,44,45,11172.3723723724a183.6627885527.0262130335a45.5058446027

    59,60,74,76,77,96,97)

    2Interlobate Moraine (109, 113)2519.5555555556a17.11858106332.1230351622a5.8906333993

    Total13691.9

    There are no significant differences in red oak understory density or relative density between physiographic systems;

    values followed by the same letter are not statistically significant (p>0.10).

    Sheet2

    Table 3.4. Comparison of northern red oak overstory layer density and relative density on selected upland landscape ecosystem types

    UMBS, Cheboygan and Emmet Co., northern Lower Michigan.

    DensityRdenBasal AreaRdom

    (stems/ha)(%)(%)

    Ecosystem TypePlots (n)xsdTypexsdTypexsdTypexsd

    3710184.497.73731.814.7748.88.03744.917.8

    745177.8198.13625.816.3378.33.17438.027.2OS

    4419151.8188.97423.219.7367.03.83633.821.1density

    3615143.770.74414.914.8446.27.74419.817.4136374041444559607476779697

    77574.471.5779.58.8453.53.74515.015.11**nd---**nd------------------

    97461.171.1967.115.9962.65.9112.424.836***nd********

    451355.654.1976.37.3772.52.69611.525.737******nd****

    96548.9109.3456.06.0762.41.57710.49.840nd**ndnd***nd*

    76548.942.7595.08.3972.22.8768.96.441**nd---***nd---

    591033.356.7764.93.812.14.2977.710.544********

    1425.050.014.59.1592.04.1597.414.945nd**ndndndnd

    4079.517.5401.62.8400.20.3401.32.259nd*nd*nd*

    4150.00.0410.00.0410.00.0410.00.060---------------

    6040.00.0600.00.0600.00.0600.00.074***---

    76*nd---

    77nd---

    Total11196---

    a, and b indicate significant differences between ecosystem groups; values followed by the same letter are not statistically significant (p>0.10).97

    * indicates test between marked pair had significance at p0.10).60------------------------------------nd60------------------------------------nd

    * indicates test between marked pair had significance at p0.10).

    * indicates test between marked pair had no significance at p0.10).

    * indicates test between marked pair had significance at p0.1045---------------------**nd****45---------------------**nd*nd*

    59------------------------*****59------------------------*****

    60------------------------------------*60------------------------------------*

    74------------------------------ndndnd74------------------------------ndndnd

    76---------------------------------**76---------------------------------ndnd

    77------------------------------------*77------------------------------------nd

    96---------------------------------------96---------------------------------------

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    stems/ha

    Type

    Stems/ha

    Red Oak Undestory Density

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    relden

    Type

    % composition

    Red oak understory relative density

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    density

    type

    stems/ ha

    red oak understory density

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    0

    relden

    type

    % composition

    Red oak understory relative density

    Table 3.29. Comparison of vegetation density and relative density in on the climatically extreme.

    low-elevation, outwash plain (Pellston Plain) ecosystem type 1, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=4)

    Speciesxsdxsdxsdxsd

    Acer rubrum12.525.02.14.20.20.41.52.9

    Pinus strobus37.575.06.312.51.12.28.817.5

    Populus grandidentata109.7138.822.530.72.33.217.824.2

    Populus tremuloides236.1157.253.920.85.62.543.27.6

    Prunus serotina16.733.310.721.41.22.416.432.8

    Quercus rubra25.050.04.59.12.14.212.424.8

    Total437.5100.012.4100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea37.575.01.73.4

    Acer rubrum241.7245.523.220.6

    Acer saccharinum62.5125.02.85.7

    Amelanchier spp.166.7333.318.837.5

    Fagus grandifolia------------

    Pinus resinosa------------

    Pinus strobus261.1493.014.721.4

    Populus balsamifera------------

    Populus grandidentata95.8109.010.811.6

    Populus tremuloides190.3149.528.121.2

    Quercus rubra------------

    Total1055.6100.0

    Table 3.29. Comparison of vegetation density and relative density in

    the climatically morderate high-elevation, outwash plain ecosystem type 36, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=15)

    Speciesxsdxsdxsdxsd

    Acer rubrum37.046.55.56.60.50.72.22.6

    Betula papyrifera4.412.50.82.20.10.20.30.9

    Pinus resinosa133.393.921.916.74.63.620.617.2

    Pinus strobus80.055.614.312.21.71.87.97.8

    Populus grandidentata235.6235.131.625.57.96.335.128.1

    Quercus rubra143.770.725.816.37.03.833.821.1

    Total634.1100.021.8100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Acer rubrum148.1166.627.526.1

    Acer saccharinum------------

    Amelanchier spp.------------

    Betula papyrifera------------

    Fagus grandifolia8.934.41.86.5

    Pinus resinosa105.2123.317.814.4

    Pinus strobus288.9268.645.222.9

    Populus grandidentata5.913.21.22.5

    Quercus rubra25.279.66.520.0

    Total582.2100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 37,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=10)

    Speciesxsdxsdxsdxsd

    Acer rubrum102.280.617.317.91.31.27.79.9

    Betula papyrifera2.27.00.61.80.10.20.61.9

    Fagus grandifolia11.121.61.73.20.10.20.71.3

    Pinus resinosa26.751.13.56.90.61.62.66.3

    Pinus strobus57.876.49.611.50.91.55.27.4

    Populus grandidentata222.2171.835.521.98.76.438.422.7

    Quercus rubra184.497.731.814.78.33.144.917.8

    Total606.7100.020.0100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Acer rubrum268.9228.933.124.4

    Acer saccharinum20.063.210.031.6

    Amelanchier spp.22.248.01.94.1

    Betula papyrifera8.928.11.85.7

    Fagus grandifolia88.9214.98.817.1

    Pinus resinosa11.118.92.86.3

    Pinus strobus191.1125.832.225.4

    Populus grandidentata31.165.62.85.0

    Quercus rubra77.8170.36.611.1

    Total720.0100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 40,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=7)

    Speciesxsdxsdxsdxsd

    Acer rubrum517.5183.667.915.513.68.461.222.3

    Acer saccharum92.190.212.913.92.02.411.413.9

    Betula papyrifera9.525.21.53.90.20.50.82.1

    Fagus grandifolia25.432.53.54.42.35.39.018.9

    Pinus resinosa

    Pinus strobus34.964.03.86.11.02.03.55.6

    Populus grandidentata50.8105.67.816.42.97.012.026.9

    Populus tremuloides6.316.80.61.60.20.60.61.6

    Prunus serotina3.28.40.41.10.10.30.41.1

    Quercus rubra9.517.51.62.80.20.31.32.2

    Total749.2100.022.4100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea3.28.40.20.6

    Acer pensylvanicum3.28.40.51.3

    Acer rubrum577.8499.245.826.6

    Acer saccharum187.3137.522.525.3

    Acer saccharinum9.511.91.01.5

    Acer spicatum231.7345.215.221.2

    Alnus rugosa19.041.41.52.7

    Amelanchier spp.28.658.41.83.7

    Fagus grandifolia25.450.42.23.7

    Pinus resinosa47.6126.03.28.3

    Pinus strobus9.517.51.22.5

    Populus grandidentata19.050.41.23.2

    Quercus rubra60.3159.63.810.1

    Total1222.2100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 41,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Abies balsamea4.49.90.30.70.00.10.10.3

    Acer rubrum377.8231.543.214.210.86.341.922.4

    Acer saccharum231.1140.227.15.15.22.719.45.3

    Betula alleghaniensis8.912.21.21.80.81.22.94.3

    Betula papyrifera4.49.90.51.10.00.10.20.4

    Fagus grandifolia142.297.617.616.27.55.428.121.5

    Ostrya virginica31.143.32.73.70.50.81.42.2

    Pinus strobus4.49.90.30.70.10.20.20.4

    Populus grandidentata26.739.86.210.21.11.55.67.7

    Tsuca canadensis8.912.20.81.20.10.10.30.4

    Total835.6100.025.999.9

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea8.912.21.11.5

    Acer pensylvanicum4.49.90.51.2

    Acer rubrum164.4168.917.520.0

    Acer saccharum404.4284.341.130.0

    Acer saccharinum102.2160.710.116.8

    Acer spicatum88.9175.011.824.0

    Alnus rugosa8.919.91.22.7

    Amelanchier spp.4.49.90.30.7

    Fagus grandifolia173.3249.615.719.1

    Pinus resinosa------------

    Pinus strobus------------

    Populus grandidentata4.49.90.61.4

    Populus tremuloides------------

    Prunus virginiana------------

    Prunus serotina------------

    Quercus rubra------------

    Total964.4100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 42,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Abies balsamea4.49.91.84.10.10.10.81.8

    Acer rubrum204.4193.334.137.14.64.529.140.4

    Acer saccharum8.919.90.92.10.10.20.30.6

    Betula papyrifera62.282.36.67.62.13.15.46.5

    Fagus grandifolia75.6168.97.917.71.63.55.412.2

    Pinus resinosa98.9106.911.912.92.32.98.210.9

    Pinus strobus87.8151.611.619.01.52.75.69.9

    Populus grandidentata200.0202.524.324.913.414.640.338.2

    Tsuca canadensis8.919.90.92.11.43.14.810.7

    Total746.7100.026.9100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea22.249.70.61.4

    Acer pensylvanicum8.919.91.84.1

    Acer rubrum238.9201.229.233.7

    Acer saccharum17.829.01.72.3

    Acer saccharinum------------

    Acer spicatum------------

    Alnus rugosa------------

    Amelanchier spp.------------

    Fagus grandifolia53.3107.311.521.8

    Pinus resinosa128.9172.413.115.6

    Pinus strobus288.9552.022.333.4

    Populus grandidentata204.4457.25.612.5

    Populus tremuloides382.2854.710.523.5

    Prunus virginiana13.329.80.40.8

    Prunus serotina53.3119.31.53.3

    Quercus rubra20.044.71.94.3

    Total1432.2100.0

    Table 3.29. Comparison of vegetation density and relative density in calcareous ecosystem type 44,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=19)

    Speciesxsdxsdxsdxsd

    Acer rubrum212.9138.124.712.83.52.614.310.9

    Acer saccharum18.442.43.48.70.30.61.74.5

    Betula papyrifera86.3122.98.89.52.13.37.210.8

    Fagus grandifolia63.7116.16.411.30.81.52.95.6

    Pinus resinosa5.814.51.02.60.30.91.02.8

    Pinus strobus62.098.98.012.52.03.28.615.1

    Populus grandidentata301.8227.732.521.714.011.944.324.0

    Quercus rubra151.8188.914.914.86.27.719.817.4

    Tsuca canadensis2.37.00.31.00.00.20.20.6

    Total905.0100.029.2100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Acer pensylvanicum5.322.90.41.2

    Acer rubrum69.3153.020.530.7

    Acer saccharum74.6264.212.919.7

    Acer saccharinum43.6122.728.542.0

    Acer spicatum5.825.53.19.3

    Alnus rugosa------------

    Amelanchier spp.3.812.36.016.6

    Fagus grandifolia71.6242.49.116.5

    Ostya virginiana------------

    Pinus resinosa5.322.90.30.9

    Pinus strobus198.5658.918.233.6

    Populus balsamifera2.611.50.20.5

    Populus grandidentata2.611.50.41.3

    Quercus rubra2.611.50.41.3

    Total485.7100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 45,

    on the climatically morderate high-elevation, outwash plain type, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=19)

    Speciesxsdxsdxsdxsd

    Acer rubrum139.393.014.510.02.81.811.07.2

    Acer saccharum333.3194.833.017.05.13.619.913.1

    Amelanchier spp.3.48.30.30.70.00.10.20.4

    Betula papyrifera106.089.410.18.33.22.811.910.0

    Fagus grandifolia179.5162.416.614.54.54.415.914.9

    Ostrya virginica71.8107.46.99.61.12.23.96.5

    Populus grandidentata47.092.15.912.52.24.012.321.9

    Quercus rubra55.654.16.06.03.53.715.015.1

    Tilia americana63.269.66.36.92.32.68.39.7

    Tsuca canadensis5.113.30.51.30.41.01.64.1

    Total1004.3100.025.2100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Acer pensylvanicum20.547.52.35.6

    Acer rubrum68.8117.44.88.8

    Acer saccharum1026.9680.671.622.9

    Acer saccharinum50.981.33.14.9

    Acer spicatum68.8167.65.817.0

    Alnus rugosa1.76.20.20.7

    Amelanchier spp.6.816.70.81.9

    Fagus grandifolia149.6176.411.112.1

    Ostya virginiana3.813.90.20.7

    Pinus resinosa------------

    Pinus strobus------------

    Populus balsamifera------------

    Populus grandidentata------------

    Quercus rubra------------

    Total1397.9100.0

    Table 3.29. Comparison of vegetation density and relative density in ecosystem type 59,

    on the deep outwash plain over moraine landform, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=19)

    Speciesxsdxsdxsdxsd

    Abies balsamea2.27.00.31.00.00.10.10.2

    Acer rubrum186.7110.536.315.15.33.926.716.3

    Betula papyrifera33.379.35.614.00.92.33.69.5

    Fagus grandifolia26.770.14.511.71.13.05.515.4

    Populus grandidentata167.8112.937.329.37.75.543.731.7

    Populus tremuloides28.957.411.022.21.22.613.127.5

    Quercus rubra33.356.75.08.32.04.17.414.9

    Total476.7100.018.1100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea6.715.00.61.4

    Acer pensylvanicum8.915.50.51.0

    Acer rubrum573.3810.335.930.2

    Acer saccharum10.031.60.92.9

    Amelanchier spp.143.9164.99.311.6

    Betula papyrifera4.414.10.10.4

    Fagus grandifolia64.492.28.311.5

    Pinus strobus8.915.51.63.1

    Populus grandidentata11.118.91.42.7

    Populus tremuloides5.015.80.51.4

    Quercus rubra426.7369.940.830.0

    Total1263.3100.0

    Table 3.29. Comparison of vegetation density, relative density in ecosystem type 60,

    on the deep outwash plain over moraine landform, UMBS, Cheboygan Co., northern

    Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=4)

    Speciesxsdxsdxsdxsd

    Acer pensylvanicum13.319.91.72.50.10.20.30.5

    Acer rubrum275.6171.133.119.95.64.514.511.1

    Acer saccharum62.2127.19.519.81.12.53.47.3

    Betula papyrifera53.373.06.59.22.84.17.510.8

    Fagus grandifolia66.749.78.35.61.10.82.81.8

    Populus grandidentata320.074.740.89.827.48.071.514.0

    Total777.8100.038.1100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea------------

    Acer pensylvanicum322.2305.027.327.5

    Acer rubrum638.9711.237.339.8

    Acer saccharum344.4688.914.929.8

    Amelanchier spp.66.7105.83.45.5

    Betula papyrifera------------

    Fagus grandifolia155.6107.312.310.1

    Pinus strobus------------

    Populus grandidentata------------

    Populus tremuloides------------

    Quercus rubra94.4160.64.88.3

    Total1622.2100.0

    Table 3.28. Comparison of overstory vegetation basal area and relative dominance on dune ecosystem type 74,on the

    moderate, high-elevation, outwash plain , UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Acer rubrum26.729.04.47.00.30.41.52.4

    Betula papyrifera4.49.90.51.20.00.10.10.3

    Pinus resinosa284.4351.224.326.56.89.021.022.7

    Pinus strobus57.871.311.516.00.80.94.15.0

    Populus grandidentata253.3151.236.119.58.45.335.221.9

    Quercus rubra177.8198.123.219.78.88.038.027.2

    Total804.4100.025.1100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea17.829.02.03.5

    Acer pensylvanicum------------

    Acer rubrum168.9109.526.229.5

    Acer saccharum------------

    Acer saccharinum------------

    Acer spicatum------------

    Amelanchier spp.------------

    Betula papyrifera------------

    Fagus grandifolia17.839.82.65.8

    Pinus resinosa155.6178.515.419.2

    Pinus strobus595.6719.143.032.2

    Populus balsamifera17.829.01.21.7

    Populus grandidentata------------

    Quercus rubra84.4165.19.615.4

    Total1057.8100.0

    Table 3.28. Comparison of overstory vegetation basal area and relative dominance on glacial lake beach ecosystem type 76, on the

    moderate, high-elevation, outwash plain , UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Acer rubrum302.2219.832.523.94.43.816.012.8

    Betula papyrifera26.736.53.04.20.40.51.62.1

    Fagus grandifolia4.49.90.51.00.00.10.10.3

    Pinus banksiana71.1115.95.79.31.42.04.46.3

    Pinus resinosa146.7232.911.818.76.18.418.525.5

    Pinus strobus62.276.05.26.00.91.03.03.0

    Populus grandidentata351.1160.636.416.712.85.447.523.8

    Quercus rubra48.942.74.93.82.41.58.96.4

    Total1013.3100.028.6100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea------------

    Acer pensylvanicum------------

    Acer rubrum382.2275.157.331.9

    Acer saccharum------------

    Acer saccharinum48.9109.35.211.7

    Acer spicatum------------

    Amelanchier spp.13.319.91.82.8

    Betula papyrifera------------

    Fagus grandifolia26.759.63.98.7

    Pinus resinosa44.487.56.29.1

    Pinus strobus142.2221.416.321.2

    Populus balsamifera13.329.81.33.0

    Populus grandidentata4.49.90.41.0

    Quercus rubra71.1134.67.614.4

    Total746.7100.0

    Table 3.28. Comparison of overstory vegetation basal area and relative dominance on glacial lake beach ecosystem type 77, on the

    moderate, high-elevation, outwash plain , UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Acer rubrum44.454.45.55.90.81.14.56.9

    Acer saccharum304.4374.827.533.66.08.026.634.3

    Betula papyrifera10.022.41.12.40.30.60.92.0

    Fagus grandifolia17.829.01.83.10.71.13.24.4

    Fraxiuns americana43.366.24.97.10.91.43.35.1

    Ostrya virginica97.8183.77.713.51.32.25.47.9

    Pinus banksiana88.9112.210.713.32.33.09.011.0

    Pinus resinosa152.2182.221.629.35.05.823.130.1

    Pinus strobus8.919.91.12.60.91.93.57.9

    Populus grandidentata50.074.36.49.51.82.57.510.4

    Quercus rubra74.471.59.58.82.52.610.49.8

    Tilia americana8.919.90.71.50.10.20.30.6

    Tsuga canadensis22.249.71.63.70.61.42.25.0

    Total892.2100.022.6100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea------------

    Acer pensylvanicum4.49.90.51.1

    Acer rubrum600.0695.835.830.0

    Acer saccharum552.2543.940.040.1

    Acer saccharinum36.746.43.14.9

    Acer spicatum52.272.83.35.2

    Amelanchier spp.22.231.43.15.4

    Betula papyrifera8.919.91.73.7

    Fagus grandifolia8.912.20.91.3

    Pinus resinosa17.829.02.85.5

    Pinus strobus88.9198.85.612.6

    Populus balsamifera------------

    Populus grandidentata------------

    Quercus rubra27.829.93.15.3

    Total1420.0100.0

    Table 3.30. Comparison of vegetation density, relative density, basal area and relative dominance on high lake terrace ecosystem type 96

    on the moderate, high-elevation, outwash plain, UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=5)

    Speciesxsdxsdxsdxsd

    Acer rubrum147.8123.225.219.93.23.412.512.6

    Betula papyrifera17.839.84.08.91.02.13.06.7

    Fagus grandifolia4.49.90.61.40.00.10.20.3

    Pinus resinosa20.044.72.14.70.20.50.81.7

    Pinus strobus88.987.514.613.52.52.511.113.5

    Populus grandidentata280.0187.646.421.416.58.761.022.3

    Quercus rubra48.9109.37.115.92.65.911.525.7

    Total607.8100.026.0100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea------------

    Acer pensylvanicum------------

    Acer rubrum258.9205.921.614.0

    Acer saccharum50.086.64.79.2

    Acer saccharinum4.49.90.51.0

    Acer spicatum4.49.90.51.0

    Amelanchier spp.80.085.58.48.9

    Fagus grandifolia160.0269.612.218.7

    Pinus resinosa158.9331.07.715.7

    Pinus strobus405.6339.333.421.8

    Populus balsamifera13.329.80.92.0

    Populus grandidentata------------

    Quercus rubra97.8171.110.115.1

    Total1233.3100.0

    Table 3.30. Comparison of vegetation density, relative density, basal area and relative dominance on high lake terrace ecosystem type 97

    on the moderate, high-elevation, outwash plain, UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=4)

    Speciesxsdxsdxsdxsd

    Acer pensylvanicum16.733.31.73.30.20.30.50.9

    Acer rubrum161.158.419.77.02.61.69.16.5

    Betula papyrifera138.9130.114.613.23.03.610.713.4

    Fagus grandifolia27.833.33.24.00.30.41.11.4

    Pinus resinosa5.611.10.71.40.10.20.30.5

    Pinus strobus33.352.94.06.70.80.92.42.9

    Populus grandidentata338.9211.944.030.119.712.762.540.1

    Quercus rubra61.171.16.37.32.22.87.710.5

    Tsuga canadensis55.6111.15.811.61.63.15.811.6

    Total766.7100.028.793.7

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea------------

    Acer pensylvanicum------------

    Acer rubrum258.9205.921.614.0

    Acer saccharum50.086.64.79.2

    Acer saccharinum4.49.90.51.0

    Acer spicatum4.49.90.51.0

    Amelanchier spp.80.085.58.48.9

    Fagus grandifolia160.0269.612.218.7

    Pinus resinosa158.9331.07.715.7

    Pinus strobus405.6339.333.421.8

    Populus balsamifera13.329.80.92.0

    Populus grandidentata------------

    Quercus rubra97.8171.110.115.1

    Total1233.3100.0

    Table 3.30. Comparison of vegetation density, relative density, basal area and relative dominance on interlobate ecosystem type 109

    UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=13)

    Speciesxsdxsdxsdxsd

    Acer pensylvanicum3.813.90.62.30.00.10.10.3

    Acer rubrum340.6247.135.320.410.07.028.523.5

    Acer saccharum87.2147.011.017.41.93.06.29.7

    Betula allegheniensis1.76.20.20.70.00.10.10.3

    Betula papyrifera29.568.42.23.51.73.83.56.4

    Fagus grandifolia159.0171.517.315.53.63.710.411.3

    Ostrya virginica5.118.50.62.00.10.30.31.0

    Populus grandidentata281.6239.529.821.720.314.448.727.1

    Populus tremuloides6.824.71.34.60.20.80.83.0

    Prunus serotina1.76.20.41.30.00.20.10.5

    Quercus rubra14.530.61.42.90.82.61.33.4

    Total931.6100.038.9100.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea3.412.30.41.5

    Acer pensylvanicum39.362.34.88.8

    Acer rubrum183.8266.020.229.3

    Acer saccharum354.7576.217.120.5

    Acer saccharinum38.5124.48.027.7

    Amelanchier spp.66.7112.25.311.2

    Betula alleghaniensis1.76.20.31.1

    Betula papyrifera1.91.1------

    Fagus grandifolia354.3320.833.635.1

    Fraxinus americana15.443.90.61.5

    Ostya virginiana265.0922.37.524.3

    Pinus strobus------------

    Populus tremuloides------------

    Prunus serotina------------

    Quercus rubra23.955.51.85.1

    Tilia americana------------

    Viburnum acerifolium3.412.30.41.5

    Total1351.9100.0

    Table 3.30. Comparison of vegetation density, relative density, basal area and relative dominance on interlobate ecosystem type 113

    UMBS, Cheboygan Co., northern Lower Michigan.

    Dominant Overstory

    Density (stems/ha)Rden (%)Basal area (m2/ha)Rldom (%)

    Ecosystem Type

    Plots (n=12)

    Speciesxsdxsdxsdxsd

    Acer rubrum251.9124.032.815.66.33.916.211.3

    Acer saccharum122.2116.216.016.32.43.16.510.3

    Betula allegheniensis5.610.10.71.40.10.10.20.4

    Betula papyrifera14.828.92.14.10.51.11.73.8

    Fagus grandifolia64.898.28.914.12.14.35.911.1

    Fraxinus americana1.96.40.31.00.00.10.10.2

    Ostrya virginica5.619.20.82.90.10.30.20.8

    Populus grandidentata248.1175.430.617.625.615.359.227.6

    Populus tremuloides29.658.73.97.02.96.26.612.1

    Quercus rubra5.610.10.81.50.20.30.40.7

    Tilia americana5.613.80.82.10.20.70.82.4

    Tsuga canadensis13.038.41.95.60.20.60.61.7

    Total750.097.740.197.0

    UnderstoryDensity (stems/ha)Rden (%)

    xsdxsd

    Abies balsamea5.613.80.51.2

    Acer pensylvanicum72.2115.36.69.6

    Acer rubrum201.9369.417.523.2

    Acer saccharum224.1214.125.223.3

    Amelanchier spp.33.347.83.14.6

    Betula alleghaniensis11.125.91.54.0

    Betula papyrifera1.96.40.20.6

    Fagus grandifolia155.6150.717.615.0

    Fraxinus americana63.084.75.26.2

    Ostya virginiana25.960.52.35.3

    Pinus strobus1.96.40.20.6

    Populus tremuloides3.712.80.31.2

    Prunus serotina3.78.60.20.5

    Quercus rubra14.833.32.56.8

    Tilia americana1.96.40.51.7

    Viburnum acerifolium------------

    Total820.483.3

  • Remote SensingIn combination with field data, GIS and Landsat data are being used to identify the patterns of ecosystem change among the selected landforms and ecosystem types.

    The 14 test stands were mapped and combined with other UMBS GIS and Landsat data layers

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Spectral-Temporal Remote SensingA combination of Landsat spectral and temporal (deciduous leaf-on/leaf-off) analysis provides information on a) aspen dominated overstory and b) deciduous vs. coniferous understory and successional pathway over the entire spatial extent of UMBS and region.

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Creating an SNRE UMBS GISMany other ArcGIS layers were created as a result of this project - a few are shown here overlaid on one Landsat band.

    The entire UMBS GIS will eventually be hosted on the ESA Lab website.HydrologyLandform Level EcosystemsTransportation

    McIntyre-Stennis/Ecosystem Managment Initiative

  • Conclusions

    Due to accelerating succession, aspen stands will be replaced by later successional species depending upon the site-specific ecosystem of their occurrence.

    Successional models based on the landscape ecosystem framework will prove useful in determining the spatial occurrence of the aspen resource, rate of decline, and succession to other species at the regional landscape scale.

    McIntyre-Stennis/Ecosystem Managment Initiative

    From this motivation we have developed a research plan

    Each objective corresponds to one of the components of the project we will talk about

    Burt - different successional pathways1. To conifer2. To hardwood3. lowland

    KB will also use to talk about using stands for remote sensing analysisof change