CMB ALUMNI NEWSLETTER SUMMER 2020

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IN THIS ISSUE…

Exceptional Thesis Award

CMB “Lab” Photos

Physiology Program

Student Retreat

Honors & Awards

Thesis Defenses

Remembering CMB Faculty

Thank you to Donors!

ALUMNI NEWSLETTER

Welcome from the CMB Chair To say that this has been an unusual year would be quite an understatement! In March, like countless others across campus and around the world, CMB students and faculty suddenly found themselves figuring out how to complete their work from home in order to help prevent the spread of COVID-19. In-person activities, including classes, professional development seminars, student seminars, lab meetings, and even thesis defenses, were moved online. CMB is currently planning a virtual new student orientation. We are adapting and learning about new tools that will hopefully come in handy in the future. We are learning the value of virtual happy hours and better appreciating the beautiful Madison summer weather and time with our families. As of August, most CMB students have been able to return to their labs at least part of the time, with measures in place like rotating schedules to ensure social distancing and safety. The CMB community has shown great resilience by finding creative ways to teach, learn, and complete research within these unprecedented constraints.

Photo credit: University Communications

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The CMB Alumni Newsletter is the annual newsletter for the Cellular & Molecular Biology Program.

1525 Linden DriveMadison, WI 53706

Email: cmb@bocklabs.wisc.edu Website: cmb.wisc.edu

Program Chair: David Wassarman

Editing, design and production: Lauren Weitkamp & Sarah Weitkamp

Cover photo by University Communications

An empty Memorial Union Terrace in June 2020. Currently, the Terrace is open by reservation to small groups.

Photo credit: University Communications

Despite everything, CMB has had a successful year, which began by welcoming 21 new rotating students and 5 MD/PhD students to the program. We will have an incoming class of 16 students in fall 2020. Last fall’s student retreat gave students a chance to share their research and unwind. The 2019 Exceptional Thesis Award review committee faced tough decisions, and ultimately chose not one but two exceptional theses by Will Olson and Kyle Quinney. As the importance of the work of virologists has been widely recognized, UW scientists have been contributing to research around COVID-19. In particular, the Kirchdoerfer Lab was recognized in January for their work studying proteins from six different types of coronaviruses, and more recently began studying proteins from COVID-19. In addition, in June CMB trainers Nathan Sherer and David O’Connor were selected for the UW/WARF COVID-19 Accelerator Challenge. This year has also brought incredibly sad news of the passing of three cherished faculty members, Heidi Dvinge, Donata Oertel, and Jennie Reed.

The Terrace looks a little different these days, and some of our usual summer activities have been put on hold, but we are excited to give the warmest Wisconsin welcome possible to our new CMB class, and to continue our work, which is now more important than ever. We look forward to the day when we can all see each other in person once again.

David A. Wassarman CMB Program Chair

Right: CMB class of 2019 on the first day of new student orientation

Photo credit: CMB Office

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EXCEPTIONAL THESIS AWARD By Sarah Rempel and Raakhee Shankar CMB Graduate Students

Meet Will Olson and Kyle Quinney, the 2019 recipients of the CMB Exceptional Thesis Award. This award is given in recognition of the distinguished work contributed by a student during the completion and defense of their thesis. Each recipient has earned a $250 reward in addition to the placement of their name on a plaque that can be found in the Bock Laboratories Penthouse.

Will OlsonWilliam (Will) Olson’s path into graduate school was a fairly common one: he enjoyed biology, so he initially thought he would go into medicine. After joining a public health lab during undergrad at University of Oregon, he was drawn to academic research. He started graduate school expecting to follow the classical academic track. His thesis entitled “Understanding the Unique Metabolism and Host Manipulation of Toxoplasma gondii” earned the CMB Exceptional Thesis Award.

During his PhD research in the Knoll Lab, Will focused on two major goals: uncovering novel metabolic pathways of the parasite Toxoplasma gondii, and understanding how T. gondii infection changed the metabolic pathways of their hosts. He built the project on the kind of academic collaboration UW-Madison prides itself on; his project brought together the T. gondii expertise of the Knoll lab with the metabolomics expertise of the Amador-Noguez lab. During infection, T. gondii does a “kiss and spit,” injecting proteins and lipids into the host cell before invasion that alters the host cell’s

metabolism. Will explored which host pathways were affected by this cocktail during the course of infection in order to understand how these changes may support T. gondii growth. During this research, he also discovered a new enzyme in T. gondii: Sedoheptulose Bisphosphatase (SBPase), which had been identified in bacteria, but never in this protozoan parasite. SBPase is part of a secondary pathway to generate the riboses that are required for DNA and RNA Synthesis. Excitingly, Will found SBPase was important for parasite growth and replication but was not present in humans, making it a potential drug target to develop new therapies.

Will Olson, one of the recipients of the 2019 CMB Exceptional Thesis Award

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After this conventional path into academia and clearly excelling at research throughout the years of his PhD work, Will is breaking the mold with his next steps: he has just finished up his first year at Marquette Law School.

Will says, “I’m gonna miss the academic environment. The positive, curious, supportive, interested culture.” Being “surrounded by some of the smartest people in the world on a daily basis is pretty cool.” So how did he end up in law school? He got exposure to patent law through a WARF ambassadorship, and becoming a patent attorney seemed like the perfect fit given his love of writing and talking about science.

“I’ll really enjoy still getting to interact with scientists and getting to hear some of the better work they’ve ever done because they’re trying to patent it.” This summer, he’s interning at the law firm Foley and Lardner in their IP office, learning and lending his biochemical expertise.

When not engaging with science and the law, Will organizes with the group he co-founded, Milwaukee Democratic Action, he cooks (no, he hasn’t made any sourdough while under stay-at-home), and he and his wife Emily walk their two adorable dogs Tillamook and Leinenkugel.

Kyle QuinneyIt comes as no surprise to me that Kyle Quinney is one of the winners of the 2019 CMB Exceptional Thesis Award. Kyle received the award for his thesis, ‘Spatiotemporal assembly of the ESCRT machinery in living cells.’

Kyle’s love for science stems from his love of working through cell biology problems. “They are a nice mix of mostly logical but a slight bit of abstract thinking, and they create beautiful pictures.” As an undergrad at The College of Idaho, Kyle majored in biology and chemistry. During this time, he gained research experience working on several different projects that included research on heat sensing ion channels, heavy metal bone cell toxicity, and a summer internship at MIT where he studied DNA damage. He quickly discovered he enjoyed cell biology research and microscopy. Kyle chose the CMB program to pursue his Ph.D. because it offered him a vast choice of faculty studying a diverse set of topics.

Will’s dogs Tillamook and Leinenkugel

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Kyle’s affinity for microscopy led him to join Jon Audhya’s laboratory. In the Audhya lab, Kyle studied how stimuli, such as growth factors, promote prolonged cellular activity of the ESCRT machinery. Since previous studies in the field had been done using ectopic over-expression of fluorescently tagged ESCRT components, Kyle set out to create functionally tagged components using CRISPR/Cas9 gene editing technology. Using the cell lines he created and advanced microscopy techniques like STED and lattice light sheet microscopy for live cell studies, Kyle was able to visualize the native spatiotemporal distribution of the ESCRT components on endosomes. His study showed that upon stimulating starved cells with growth factor, each of the ESCRT components remained associated with endosomes for a considerably longer duration, acting repeatedly to internalize cargo and down-regulate cell signaling. Kyle’s research led to a better understanding of endogenous ESCRT dynamics.

During his time at the Audhya lab, Kyle mentored several undergraduate students and oversaw projects assigned to rotation students. He also received the Molecular Biosciences Training Grant and was selected to present his work at a Gordon Research Seminar.

One of the most important lessons Kyle learned during his PhD is working through tough problems that do not seem to have a solution. He also developed the skill to communicate his science confidently and effectively, “People really cared a lot about what I was working on when I talked with confidence.”

He advises current graduate students to have a non-science hobby, “Because if you never shut off when things are going poorly, it’s hard to feel mentally fit.” He acknowledges that this advice is hard to follow at times.

Kyle is now a post-doc in Dr. Anna Akhmanova’s lab at Utrecht University in The Netherlands studying kinesin motors and ER morphology and functionality. He chose Dr. Akhmanova’s lab because he wanted to study ‘something halfway different from his PhD work’. Moving to a different county is a big decision, but Kyle wanted to get outside his comfort zone and experience other scientists and places.

Outside work, Kyle likes to spend time with his dog Bumi, play video games, go bouldering, and biking. He is also the person I look to when I want suggestions for good music, podcasts, and TV. Fun fact about Kyle: he knows random facts about most things!

Kyle Quinney, one of the recipients of the 2019 CMB Exceptional Thesis Award

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CMB “LAB” PHOTOS This year, instead of taking our traditional lab photos, CMB graduate student Becky Reese helped to collect photos and stories of students’ home workspaces. We loved getting a window into everyone’s work-from-home setup!

Becky Reese (Fifth Year, Elaine Alarid Lab) The biggest difference between my desk at home and my desk at lab is that I have a window next to my desk at home! To help improve my work from home situation, I invested in a new desk chair and a little clip on desk lamp to light up my space. My second monitor has also been very helpful! Above it is a movie frame from Finding Nemo that my mom got me back when I started college to remind me to "just keep swimming"

Cassie Leech (First Year, John Denu Lab)

Working from home hasn’t been so bad when it means staying in your pajamas with your furry coworkers (aka Boomer). Buying a desk and grabbing my computer from lab were huge helps in making me more productive and better able to separate work time from relax time. Virtual game nights and plenty of Sudoku puzzles have also kept me sane!

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Caitlin Murphy (Recent Grad, Matthew Banks Lab, PGTP)

I defended May 6th from home in my footy sweatpants (feejays?). I painted myself some congratulatory balloons and gave them to myself afterward.

Caitlin working in her lab, pre-COVID

Dan Tremmel (Fifth Year, Jon Odorico Lab)Working from home for me comes with the obvious drawback of having very limited methods of generating new data, but I have gotten a lot of writing done and I don’t mind the sunshine, plants and cats! The downside is not working at a real desk, and the cats want to be on top of my stuff all day.

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Sarah Rempel (Fifth Year, Timothy Gomez Lab)

My home office is a corner of my sewing table (I made the shirt and jeans I'm wearing). For a while I was working solely on my laptop, but I finally got my big monitor from lab, so my screen area has quintupled! This has made it a lot easier to work on my video talk for a now-virtual conference and to work in Photoshop on figures for a paper. Of course one of the most important features of my work-from-home setup is a cup of coffee!

Eating lunch with my son Otis!

Steven Wiesner (Sixth Year, Xin Huang Lab, PGTP)

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ANNOUNCING A NEW FOCUS GROUP IN PHYSIOLOGY

In July 2019, we announced that the CMB Coordinating Committee had voted to add a new focus group in Physiology to serve as the home for physiology studies on campus going forward. Dr. Raunak Sinha (Department of Neuroscience) is serving as the chair for the new focus group.

This decision came following the recent discontinuation of the Physiology Graduate Training Program. In addition to adding a new focus group, CMB is also serving as the administrative home for the continuing Physiology students while they complete their Physiology degrees.

Since the addition of the new focus group, 30 faculty members have joined the focus group, and interest in this area of research has already brought additional prospective students to CMB. The graduate students of the physiology focus group together with Dr. Sinha have been planning the first graduate student organized physiology symposium on campus. The symposium was originally scheduled to be held on May 21-22, 2020 but is currently postponed due to the pandemic. The symposium will feature talks from student, local faculty, and invited faculty speakers, along with a poster session. Please stay tuned for more updates about the symposium.

We welcome the continuing Physiology students and are pleased to be able to offer a new home for physiology studies on campus! The new focus group webpage can be found on the CMB website: https://cmb.wisc.edu/physiology/.

Raunak Sinha, Department of Neuroscience, and new CMB Physiology Focus Group Chair

Physiology of an output neuron in the retina

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CMB STUDENT RETREAT 2019 PHOTO GALLERY

The foggy view of Lake Delton early the morning after retreat

This year’s student retreat took place at Baker’s Sunset Bay Resort in Wisconsin Dells after Heidel House, our usual retreat location, closed in May 2019.

Leland Hyman giving a talk

Ian Begeman playing a petri dish toss game

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CMB STUDENT RETREAT 2019

Zena Jensvold playing pin the safety glasses on the scientist

Jon Schmitz, Debayan De Bakshi, and Marissa Gaskill, the retreat organizers who did a great job!

Truman Do playing the petri dish toss game

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CMB STUDENT RETREAT 2019

The team that won the coveted Golden Pipette used their gift cards to go out to dinner together at Eldorado Grill. They brought the Golden Pipette, and while they were there, a table of employees at Gilson (the company that made the pipette) came and talked to them because they were super excited (and confused) to recognize their pipette at the restaurant. They all took a picture together (above) to remember the moment!

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HONORS & AWARDS 2019-2020

Student Awards Katarina Braun, Class of 2017, Friedrich Lab NIH F30

Morgan Giese, Class of 2017, Huttenlocher LabNSF Graduate Research Fellowship - Honorable Mention

Luis Haddock-Soto, Class of 2019, Friedrich LabSciMed GRS

Jessica Han, Class of 2019, Denu LabBio-Data Science Training Grant

Jack Hunt, Class of 2016, Zhao LabRuth L. Kirschstein NRSA Individual Predoctoral Fellowship (NIH F31)

Justin Jagodinski, Class of 2017, Morris LabUW Institutional for Clinical & Translational Research TL1 Training Award

Zena Jensvold, Class of 2017, Lewis LabNSF Graduate Research Fellowship

Charlotte Kanzler, Class of 2019, Newmark Lab First Prize in the Morgridge Institute for Research Ethics Cartooning Contest

Justin Mabin, Class of 2017, Brow LabNSF Graduate Research Fellowship

Jose Martinez, Class of 2018, Bhattacharyya LabBiotechnology Training Program (BTP)

Charu Mehta, Class of 2014, Bresnick LabUW Stem Cell and Regenerative Medicine Center Graduate Training Award

Zach Morrow, Class of 2017, Sauer LabNSF Graduate Research Fellowship

Alex Pieper, Class of 2017, Sondel LabICTR TL1 Training Grant

Stefan Pietrzak, Class of 2017, SridharanGenomic Sciences Training Program Traineeship

Charlotte Kanzler’s winning comic

A still image from Tiaira Porter’s winning UW-Madison Cool Science Image video

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HONORS & AWARDS 2019-2020

Student Awards (cont’d) Tiaira Porter, Physiology Class of 2016, Moore LabUW-Madison 2020 Cool Science Image Winner Along with grad student Chris Morrow, Tiaira captured a video of the movement of neural stem cells switching from dormancy to activity

Sarah Rempel, Class of 2015, Gomez LabVision Research Training Program

Kevin Swift, Class of 2019, Audhya Lab Molecular and Cellular Pharmacology Training Grant

Andrew Sung, Class of 2018, Pagliarini LabBiology of Aging and Age Related Diseases Training Grant

Dan Tremmel, Class of 2015, Odorico Lab Ruth L. Kirschstein NRSA Individual Predoctoral Fellowship (NIH F31)Debra Hullett, PhD, Best Basic Science Poster Award (UW Dept. of Surgery Research Summit 2020) ALPCO Young Investigator AwardTTS-IPITA Scientific Award to attend IPITA 2019BI-USA Stem Cell Research Award

Faculty Awards Jean-Michel AnéPostdoc Mentoring Award

Anjon AudhyaNamed Associate Dean for Basic Research for the University of Wisconsin–Madison School of Medicine and Public Health2019 Ride Scholar

Andrew Bent2019 UW Vilas Mid-Career Investigator Award

Emery BresnickWARF Named Professorship

Mark Burkard 2019 UW Vilas Mid-Career Investigator Award

Edwin ChapmanWARF Named Professorship

Melissa HarrisonH.I. Romnes Faculty Fellowship

Anna HuttenlocherWARF Named Professorship

Pam Kreeger 2019 Ride Scholar

Dudley Lamming2019 Vilas Faculty Early Career Investigator Awards

Peter Lewis2019 Vilas Faculty Early Career Investigator Awards

Hiroshi MaedaH.I. Romnes Faculty Fellowship

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HONORS & AWARDS 2019-2020

Faculty Awards (cont’d) Shigeki Miyamoto2019 Ride Scholar

Darcie Moore2020 Vallee Scholar

David O’ConnorUW/WARF COVID-19 Accelerator Challenge

Marisa OteguiKellett Mid-Career Award

Avtar Roopra 2019 UW Vilas Mid-Career Investigator Award

Kris SahaNamed Retina Research Foundation Chair at the McPherson Eye Research Institute

Nathan Sherer 2019 Ride ScholarUW/WARF COVID-19 Accelerator Challenge

Ahna SkopIF/THEN Ambassador

Rupa Sridharan 2019 UW Vilas Mid-Career Investigator Award

Chad Vezina2019 UW Vilas Mid-Career Investigator Award

Alumni Honors & Awards Kevin Cope, PhD 2018, Ane LabPhoto Selected for Cover of The Plant Cell

William Olson, PhD 2019, Knoll Lab2019 Exceptional Thesis Award

Kyle Quinney, PhD 2019, Audhya Lab2019 Exceptional Thesis Award

Kelly Tyrrell, MS 2011, Amann Lab NASW 2019 Excellence in Institutional Writing Award

Four CMB trainers, Anjon Audhya, Pam Kreeger, Shigeki Miyamoto, and Nathan Sherer, were among the 2019 Ride Scholars

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Francisco Barros Becker Investigating the mechanisms of random and directed migration of leukocytes in vivoAdvisor: Anna HuttenlocherCurrent Position: Post-doc, University of Washington, Seattle

Tim CatlettAxon guidance in human neurons and Tuberous Sclerosis: caveat mTORAdvisor: Timothy GomezCurrent Position: Post-doc, University of Wisconsin-Madison

Tamara ChamberlinInteractions Among Epithelial Cells and Macrophages to Promote Obesity-Associated Breast CancerAdvisor: Lisa ArendtCurrent Position: Assistant Professor, Midwestern University Veterinary School

Tony DawsonPhosphorylation controls assembly and activity of the influenza virus replication machineryAdvisor: Andrew Mehle

Drew DoeringProtein evolution and genomic innovation in eukaryotic iron acquisitionAdvisor: Chris HittingerCurrent Position: Scientist, Xylome Corporation

Netta GolenbergA citrullination deficient zebrafish model reveals a role for Padi2 in fin regenerationAdvisor: Anna HuttenlocherCurrent Position: Clinical Genomic Scientist, Children’s Hospital of Philadelphia

Amelia HajFunctional consequences of immunogenetic variation in rhesus and cynomolgus macaquesAdvisor: David O’ConnorCurrent Position: Medical Student, UW-Madison SMPH

Lauren HillersThe role of the obese microenvironment in mediating breast cancer progression and metastasisAdvisor: Lisa ArendtCurrent Position: Post-doc, National Cancer Institute

CMB STUDENT THESIS DEFENSES 2019-2020

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Jack HuntMicro to Macro in Nervous System Disorders: Using Fragile X Syndrome Stem Cells as a Drug Discovery Platform and Investigating the Role of Neighborhood Disadvantage in Neurodegeneration and Cognitive DeclineAdvisor: Xinyu ZhaoCurrent Position: Medical Student, UW-Madison SMPH

Siddhant Jain Dysregulation of Polycomb Repressive Complex 2 by oncogenic histone mutations in pediatric brain and bone tumorsAdvisor: Peter Lewis

Jeong-Hee LeeMechanisms of human cytomegalovirus entry into undifferentiated cells establish latencyAdvisor: Rob KalejtaCurrent Position: Post-doc, University of Wisconsin-Madison

Angela MyersDksA binding to E. coli RNA polymerase increases ppGpp binding to Site 1Advisor: Richard GourseCurrent Position: Lecturer, Edgewood College

Michelle Pearson Impacts of drought on carrots and their symbiotic fungi: from field interactions to gene expressionAdvisor: Erin SilvaCurrent Position: Instructor, Spokane Falls Community College

Kyle QuinneySpatiotemporal assembly of the ESCRT machinery in living cellsAdvisor: Anjon AudhyaCurrent Position: Post-doc, Utrecht University

Laura SwansonIdentifying immunologic and metabolic contributors to secondary injuries in a Drosophila model of traumatic brain injuryAdvisor: David WassarmanCurrent Position: Medical Student, UW-Madison SMPH

Jeremy VolkeningDevelopment and application of mass spectrometric technologies for plant molecular biologyAdvisor: Michael SussmanCurrent Position: Partner/Consultant, BASE2BIO, LLC

CMB STUDENT THESIS DEFENSES 2019-2020

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Sarah WilsonDetermining the roles of Toxoplasma gondii patatin-like phospholipases and lipoxygenases during infectionAdvisor: Laura KnollCurrent Position: Validation Scientist, Entrogen

Sihui YangIn vivo genetic analysis of Golgi outposts’ role in microtubule organization in Drosophila sensory neuronsAdvisor: Jill WildongerCurrent Position: Scientist, Caltech

Aurélia Faure How KSHV needs EBV to transform peripheral B cells towards lymphomagenesisAdvisor: Bill Sugden

Anna Kowalkowski Intestinal atresias and development with a focus on the Fibroblast growth factor receptor 2IIIb mouse modelAdvisor: Peter Nichol

Yihe Ma Dentate gyrus and cochlear nuclei circuitry revealed by a genetically-encoded voltage indicatorAdvisor: Meyer Jackson

Caitlin Murphy Synaptic and network responses to afferent inputs in higher order cortex and their modulation by the volatile anesthetic isofluraneAdvisor: Matthew Banks

Jun Wan The regulatory role of the mitotic checkpoint component Mad1 in interphase and tumor progression Advisor: Beth Weaver

CMB STUDENT THESIS DEFENSES 2019-2020

PHYSIOLOGY STUDENT THESIS DEFENSES

2019-2020

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We are saddened by the passing of three CMB faculty trainers, Heidi Dvinge, Donata Oertel, and Jennifer Reed, over the past year. We will miss their enthusiasm for science, dedication to teaching and mentoring, and friendship. Heidi was an Assistant Professor in Biomolecular Chemistry who studied how misregulation of RNA processing contributes to disease. She was

an active participant in the RNA Biology focus group as well as in recruiting students to CMB. Donata was the founding chair of the Department of Neuroscience and the Director of the Physiology Graduate Program. Before her passing, she played a large role in transitioning the Physiology Program into CMB and establishing a Physiology focus group. Jennifer was a faculty member in the

Department of Chemical and Biological Engineering for 13 years. She was a dedicated contributor to CMB, serving as Chair of the Systems Biology focus group for several years. The contributions made by Heidi, Donata, and Jennifer will positively impact others’ lives for many years to come.

REMEMBERING CMB FACULTY By CMB Chair David Wassarman

Heidi DvingePhoto by Robin Davies

Donata Oertel

Jennifer Reed

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CMB COORDINATING COMMITTEE The governing body of CMB, 2019-2020

Chairs

David Wassarman, Program ChairBill Bement, LCMB ChairBarak Blum, Admissions ChairCurtis Brandt, Advising & OrientationDonna Peters, Curriculum ChairCaroline Alexander, Awards Chair

Focus Groups

Caroline Alexander, Cancer Biology Jill Wildonger, Cell Adhesion & CytoskeletonDavid Pagliarini (fall), Dudley Lamming (spring), Cellular & Molecular MetabolismAnne Griep, Developmental Biology & Regenerative MedicineJyoti Watters, ImmunologyGuy Groblewski, Membrane Biology & Protein TraffickingRobert Landick, Molecular & Genome Biology of MicrobesRaunak Sinha, PhysiologyJean-Michel Ane, Plant Biology David Brow, RNA BiologyMegan McClean, Systems BiologyMelissa Harrison, Transcriptional MechanismsPaul Ahlquist, Virology

Students

Anna Heffron, MD/PhD representativeZach MorrowKatie MuellerDan Tremmel

Staff

Lauren Weitkamp, Student Services Coordinator Sarah Bierke, Student Services Coordinator Kim Voss, Assistant Director, Bock LabsMarcia Gratz, Payroll & BenefitsKatherine Spencer, Financial Specialist

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Share Your Story

Alumni, we would love for you to share your story in one of two ways:

• Answer our Alumni Spotlight Questionnaire. Your stories will be extremely helpful to our current students, and will give prospective students an idea of the variety of career paths CMB graduates pursue

• Update us on your current job. Any updates shared will be included on the CMB website under the Alumni Directory.

THANK YOU, ALUMNI!

CMB Donors, 2019-2020 Thomas L. Benjamin

Marc Collett

Craig Crews and Katherine McKenzie

Eleanor R. Erikson

Joan and Kenneth Gelhaus

Bradley Gerberich

Tony Hunter

James Kirui

Kyung Lee

Xiaoqi Liu

Richard Macnair

Magda and Michael McCormick

Phyllis Koton Neel and Benjamin Neel

Jane Shaffer

Daniel and Trudy Simmons

Matthew Stanzler

Deborah Steenland

Laurel Sweet

Ellen Thompson

Stay Connected! Twitter: @CMB_UWMadison

Facebook: https://www.facebook.com/cmb.uwmadison

LinkedIn: CMB Professional Network at UW-Madison

Website: https://cmb.wisc.edu/

Thank you for your support!As one of the largest biological science graduate programs at UW-Madison, CMB has been committed to excellence in graduate education since the 1960’s. If you would like to make a gift to this long standing effort, visit the CMB UW Foundation page.