• Gaseous exchange: uptake of O2 from the environment and the discharge of CO2 to the environment.

• Takes place in respiratory structures.• Movement of the respiratory gases

between the respiratory surface and the external environment is by diffusion.

• Characteristics of respiratory structures:– The respiratory surface is moist– Cells lining the respiratory surface are thin– The respiratory structure has a large surface

area.

• Small organisms do not need any special respiratory structure.

• Respiratory surface for unicellular organism such as Amoeba sp. is the entire plasma membrane.

• Surface area per volume depends on the size of organism.

Respiratory system of insects known as tracheal system; composed of air tubes called tracheae.

Air enters the tracheae through spiracles. Spiracles of insects such as grasshopper have valves which

allow them to open and close, so that air can go in and out of the body.

Tracheae branch into fine tubes called tracheoles.

Large number of tracheoles provides large surface area fordiffusion of gases.

• Air travels through the tracheoles reaches the fluid filled tips, O2 diffuses directly from the tracheoles into the cells, while CO2 diffuses from the cells into the tracheoles.

• Larger insects have air sacs which speed up movement of gases to and from tissue.

• Air is drawn into and out of tracheae by rhythmic movements of abdominal muscles that compress and expand tracheae to speed up diffusion of gases into the cells.

• Insect inhales;– Abdominal muscle relax– Spiracles are open– Air pressure inside is lowered– Air is drawn in

• Insect exhales;– Abdominal muscles contract– Increase air pressure– Forces air out

Respiratory structure: gills

Each gill composed of filaments.

Filaments :• have numerous thin-

walled named lamellae.

• Have thin membranes

• Rich supply of blood capillaries

Gaseous exchange enhanced by countercurrent exchange mechanism.• The blood in the capillaries flow in the opposite

direction to the flow of water.• ensures maximum intake of O2 (80%) by the

blood.• Deoxygenated blood enters the capillaries, it

encounters water with higher O2 content • Thus, along the capillaries, the diffusion

gradient allows the transfer of O2 into the blood.

Ventilation increases the flow of water over respiratory surface by swimming and opening and closing the operculum.

Gaseous exchange occurs through skin and lungs.

Skin:• Thin• Highly permeable to

respiratory gases.• Kept moist by secreting

mucus• Beneath skin, large

number of blood capillaries

Facilitate rapid and efficient exchange of gases between skin and environment.

Lung:• Consists of a pair of thin walled sacs• Thin• Moist• Covered by network of capillaries.

Breathing mechanismsa)Filling buccal cavity with air

Bucco-pharyngeal floor lowered glottis closes nostril open to suck air air is drawn into bucco-pharyngeal cavity.

b)Forcing air into the lungsGlottis open nostrils closed bucco-pharyngeal floor

raised air pressure increased air is forced into the lungs

c)Frog repeat steps (a) and (b) to take in more air, pumping up itself to a considerable size

d)Expirationthe frog opens its nostrils and glottis air flows out of its lungs.

Breathing mechanism involves:• Inhalation• Exhalation

Inhalation:• External intercostal muscles contract

internal intercostal muscles relax rib cage move upwards and outwards diaphragm contracts and flattens increases volume of thoracic cavity reduced air pressure air is drawn into the lungs through nostrils and fill the alveoli.

Exhalation• External intercostal muscles relax

internal intercostal muscles contract rib cage move downwards and inwards diaphragm relax and returns to original shape decreases volume of thoracic cavity increased air pressure air is forced out of the lungs.