· 3 intended for hydrobiologists, entomologists, ecologists, university teachers and students of...

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Annotation

Information on the Hydradephaga of the north-western Black Sea Region(Odessa, Nikolaev, and partly Kherson Areas of Ukraine) is for the first timesummarized in this monograph. Species composition, biotopic distribution, lifecycles and phenology, aspects of ecomorphology, relations of Hydradephagawith other organisms are discussed. Seasonal changes of species composition,abundance, and biomass of the Hydradephaga of the region are described.Recommendations for the protection of Hydradephaga and for the usage ofstenoecic species as bioindicators of water quality are given. The book is

3

intended for hydrobiologists, entomologists, ecologists, university teachers andstudents of biology, as well as for amateur naturalists. 21 figs, 35 tables, 262references.

Vasily G. Dyadichko

4

Contents

List of abbreviations 9

Introduction 10

Acknowledgements 13 1.

Chapter 1. Materials and methods 15 2.

Chapter 2. Description of the studied region

Brief physiographic reviewof the north-western Black Sea Region (NWB) 21

HydradephagaStudied water-bodies (WB) as habitats of Hydradephaga

WB of the Ukrainian part of the Danube`sdelta and adjacent territories 22

.WB of the lower reaches of the Budaksky liman 23

.WB of the lower reaches of the Dniestr river andthe Dnestrovsky liman 24

.WB of Odessa city and its environs 26

.WB of the Bolshoy Kuyalnik river basin andof the Kuyalnitsky liman 28

.WB of the Tiligul river basinand of the Tiligulsky liman 30

.WB of the lower reaches of the riversSasyk and Berezanka and of the Berezansky liman 32

5

WB of the South Bug river basin and its tributaries 33,

.WB of the lower reaches of the Dnieper river,the Dneprovsko-Bugsky liman, and of the Kinburnsky peninsula 35

3. HYDRADEPHAGA

Chapter 3. Species composition of the Hydradephaga of NWB

The history of studying the Hydradephaga of NWB 38

Hydradephaga .

General description of the species composition ofHydradephaga of NWB. New information on the region’s fauna 39

Hydradephaga

Peculiarities of the formation ofHydradephaga species composition underthe influence of some factors of water environment 42

Hydradephaga .Attitude of Hydradephaga to flow of water 44

Hydradephaga Attitude of Hydradephaga to temperature 44

Hydradephaga Attitude of Hydradephaga to salinity 44

Hydradephaga Attitude of Hydradephaga to the type of bottom substratum 45

Hydradephaga ( )Attitude of Hydradephaga to the hydrogenion exponent ( ) 46

Hydradephaga Aquatic and riparian vegetation as a factor influencing theformation of Hydradephaga species composition in NW 46

Hydradephaga

Species composition of Hydradephaga of different WB of the region 50

Springs and streams 50

6

)Steppe intermittent rivers(by the examples of the Tiligul and the Bolshoy Kuyalnik) 52

)Small non-intermittent rivers(by the examples of the Savranka and Kodyma) 56

, , ).Lower reaches of the large lowland rivers (by the exampleof the Danube, Dniester, South Bug, and Dnieper) 57

.Limans of NWB 63

.Stagnant WB of the Kinburnsky peninsula 65

Hydradephaga Faunal similarity and structural organization of Hydradephagataxocenes in NWB 67

Hydradephaga

Some stenoecic species of the Hydradephaga of NWBas bioindicators of water quality 74

Hydradephaga Some general patterns of the formation ofHydradephaga species composition in NWB 79

4. ,

HYDRADEPHAGA

Chapter 4. Seasonal changes of the speciescomposition, numbers and biomass of the Hydradephaga of NWB

Brief background 81

HydradephagaExogenous and endogenous causes of seasonal changes inHydradephaga taxocenes 81

, Hydradephaga

7

Seasonal changes of the species composition, abundance,and biomass of the Hydradephagain different types of rivers of NWB 82

, Hydradephaga

Seasonal changes of the species composition, abundance,and biomass of Hydradephaga in springs and streams of NWB 99

, Hydradephaga

Seasonal changes of the species composition, abundance,and biomass of Hydradephaga in some saline WB of NWB 101

5. HYDRADEPHAGA Chapter 5. Some aspects of the biology of the Hydradephaga of NWB

Hydradephaga Life cycles (LC) of the Hydradephaga of the region 103

Hygrobiidae [= Paelobiidae]Family Hygrobiidae [= Paelobiidae] 103

HaliplidaeFamily Haliplidae 104

Noteridae.Family Noteridae 106

Dytiscidae.Family Dytiscidae 107

Gyrinidae.Family Gyrinidae 111

HydradephagaRelationship between the type of LC and habitatpreferences of the Hydradephaga 112

Size groups in adults of the Hydradephaga species of the region 114

Hydradephaga,

Aspects of ecomorphology and dietary habits ofregional Hydradephaga species in connectionwith their distribution in types of aquatic communities 116

6. ,

8

Chapter 6. Preliminary results of the studies ofpredators, parasites, and commensalsof the Hydradephaga of NWB 126Hydradephaga

The Hydradephaga of NWB as food for predatory invertebrates 126Hydradephaga The Hydradephaga of NWB as food for vertebrates 128

Hydradephaga Taxonomic composition of parasites and commensalsof the Hydradephaga of the region 131Fungi 131Algae 132Ciliophora 133Trematoda 133

Conclusion 136

References 143 1

Appendix 1 166 2

Appendix 2 175 3

Appendix 3 194

Index 196

9

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. .

B D ICS N

W

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, (Adephaga) (Polyphaga) [179]. Adephaga

11 , 8 Hydradephaga

11

. (Amphizoidae), (Meruidae), (Aspidytidae), (Hygrobiidae),

(Haliplidae), (Gyrinidae), (Noteridae) (Dytiscidae) [177, 185, 215, 216, 251, 258].

5 [127, 214].

Paelobiidae [226], . Hydradephaga ,

, 190-220 .

[64].

.

, . :

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( 100) ,

.,

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, « » [34,

38].

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1

Hydradephaga, 1999 – 2008 . ( . 1).

1 1999 2008 .

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2007

2001 – 2008

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, [143, 174].

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(60-80º ),

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(36317, 27.10.2008),

[79], . [104].

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(Ø 0,4 – 1,0 ).

0,5×0,3 .

17

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. 1. : 1 – , 2 – , 3 –, 4 – , 5 – , 6

– , 7 – , 8 – , 9 – , 10 – .

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:N=n/k×L (1)

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:=b/k×L (2)

18

– , 2, b – ,, 1.

[140], 1 .

, – 2 . 80 Hydradepahaga.

( -9, -10) [31, 35, 71, 126, 127, 234, 143, 189, 203, 204, 213, 252,

253, 256, 257]. . . . Hydradephaga

. , . [224, 225, 226, 229, 230, 232, 235, 256, 257].

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19

1 – 2–5 ;2 – 7–18 ;3 – 26–41 .

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[8, 21, 26, 94, 84]:D= p (3),

D – , B – , 2, – , %.

[129]:ICS=2c/a+b (4),

c – , , a b –.

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[102, 129, 170]. , ( )

, [42].

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[87, 88, 119, 249]:W(A B) = c/a ; W(B A) = c/b (5),

W – , (4).

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1 – 25% ;2 – 50% ;3 – 75% ;4 – 100% .

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33

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38

3 HYDRADEPHAGA

., Hydradephaga ,

XIX . [236], . [219-222] . [158],

..

. [98]. Hydradephaga, 1 , 1 , 19

1 . .

. [85] (Hydradephaga) .

. 2 , 22 2 .

. Bidessuspumilus Aubè, 1836, ,

[31].. [71],

Hydradephaga . (

) H. pallidulus, , « » «

», « » . 90 .

. [74].

.. [132]

Hydradephaga, . 35 (5 , 2

, 25 3 ), . Hydradephaga

. [82, 83]. - 2 : H. ruficollis G. marinus.

. [149] 3 , .

39

2001 . 5 «» [127],

, . 3 : H. hermanni,

Haliplus mucronatus Stephens, 1828 H. pallidulus. 115 Hydradephaga.

Adephaga [36, 37].

. [36] . [37] 59 Hydradephaga (6, 50 , 2 1 ),

. . 1971 1986 . ,

., Hydradephaga

, .

Hydradephaga ..

103 Hydradephaga, 30 5 :Hygrobiidae – 1, Haliplidae – 15, Noteridae – 2, Dytiscidae – 75, Gyrinidae – 10

( . 2). 2

Hydradephaga

( ) A , Hygrobiidae Hygrobia (1) 1 0 1

Haliplidae Peltodytes, Haliplus (2) 15 15 0

Noteridae Noterus (1) 2 2 0

Dytiscidae

Liopterus, Bidessus, Hydroglyphus, Herophydrus,Hygrotus, Hydroporus, Suphrodytes, Porhydrus,Graptodytes, Nebrioporus, Hydrovatus, Hyphydrus,Platambus, Agabus, Ilybius, Rhantus, Colymbetes,Laccophilus, Acilius, Graphoderus, Cybister, Dytiscus,Hydaticus (23)

75 66 9

Gyrinidae Aulonogyrus, Gyrinus, Orectochilus (3) 10 7 3. : – , A – , , M –

, .

90 , 1 – A. concinnus ( ). 12

40

(H. hermanni, H. quinquelineatus, H. corpulentus, H. flaviventris, H.erythrocephalus, H. striola, H. obscurus, H. tristis, G. granularis, A. lineatus, G.marinus, G. colymbus) [37, 82, 83, 85,98, 127, 132]. ,

, ( ,, , ). «

» . [71] «» [127],

, 36 Hydradephaga: Haliplus lineolatus Mannerheim,

1844, H. mucronatus Stephens, 1928, H. dalmatinus J. Müller, 1990, Bidessusminutissimus (Germar, 1824), B. delicatulus (Schaum, 1844), B. alienusZimmermann, 1919, Nebrioporus canaliculatus (Lacordaire, 1835), N. airumlus(Kolenati, 1845), N. luctuosus (Aubè, 1838), N. elegans (Panzer, 1794),Hygrotus polonicus (Aubè, 1842), H. marklini (Gyllenhal, 1813), H.nigrolineatus (Steven in Schünherr, 1808), H. caspius (Wehncke, 1873), H.saginatus (Schaum, 1857), H. pectoralis (Motschulsky, 1859), Hydroporusumbrosus (Gyllenhal, 1808), H. obsoletus Aubè, 1836, H. marginatus(Duftschmid, 1805), H. tessellatus Drapiez, 1819, Scarodytes halensis(Fabricius, 1787), Porhydrus obliquesignatus (Bielz, 1852), Hyphydrus sanctusSharp, 1882, Agabus guttatus (Paykull, 1798), A. dilatatus (Brullè, 1832), A.sturmii (Gyllenhal, 1808), A. amoenus Solsky, 1874, A. zimmermanni Scholz,1920, Ilybius cinctus Sharp, 1882, Rhantus suturellus (Harris, 1828), R.notaticollis (Aubè, 1837), R. consputus (Sturm, 1834), Eretes griseus (Fabricius,1781), Hydaticus continentalis J. Balfour-Browne, 1944, Dytiscus persicusWehncke, 1876.

Hydradephaga, . 11

: H. immaculatus, H. sibiricus, H. maculatus, H. discretus, A.paludosus, A. fuscipennis, A. didymus, I. fenestratus, I. similis, R. latitans, R.exsoletus.

3 – H. zacharenkoi, H. musicus H. pallidulus - . A. fuscipennis, A. didymus

R. exsoletus, .

( 1 2). Hydradephaga

[38], [106, 107, 111], [11,

12]) ,

41

, ( . 3)., Hydradephaga

, . 3

Hydradephaga ( )

Hygrobiidae 0 0 1 1 1 0 0 0Haliplidae 15 16 15 10 16 - - -Noteridae 2 2 2 2 2 - - -Dytiscidae 95 122 75 90 121 - - -Gyrinidae 6 6 10 6 13 - - -

118 146 103 108 153 96 115 98

[106] [111] , [37, 82, 83, 85,98, 132, 127]

[12,82]

[11, 12,82] [38]

. – , - — , : –, – , – , –

, – : – , –, – .

, Hydradephaga

, . , Hydradephaga

.

. [72, 75, 261], (1,5-2 . )

, (2-7 . ) – . ,

(Hydradephaga) , ,

, . ,

, . ,

Hydradephaga [38, 131].

() Hydradephaga

, .

42

, Hydradephaga

.,

Hydradephaga .

, .

Hydradephaga .

Hydradephaga .

,, . , « » .

[71] Hydradephaga, .

, , .

. [38, 39],

. Hydradephaga

..

. [106, 107], . . [114],. [12], . [111], . [120, 121],. [141-144], . [4], . [131]

. ,

. Hydradephaga

,.

. . [106]

, ,

43

, . ,

. , . . ,

, [131].,

Hydradephaga .

, . [37].

Hydradephaga ,

-. [12, 38, 39, 106, 115, 143, 234],

, , , –

. , pH,

, .. [38] ,

, ( . ).

Hydradephaga ( ) .

, .

. [38], . [106],. [12], . [131] . [143]

. , , ,

-. .

(

), .

( ) . [38] Hydradephaga .

.

44

,

Hydradephaga . ,

.

Hydradephaga , . Hydradephaga .

:1) – (

). 4:

) – ;) – ;) ( . inundantio – , ) –

, .

. [39, 46, 49, 51, 54,62, 63] ,

;) ( ) – ,

.2) –

. 2 :

) – ;) – ,

( , , (A. lineatus) – ( )

). Hydradephaga .

Hydradephaga :

1) – , ( , 15-20° );

2) – , (15° );3) ,

. Hydradephaga .

:

45

1) ( ) – ( 20‰, 60‰);

2) ( . « ») – ( 15-18‰,

5‰).

, .

. , 2007

H. enneagrammus, ,. ,

.3) (1 H. geminus)

();

4) ( ) 0,5-2‰.

, ,

, , .

. H. ovatus, H.inaequalis, I. subaeneus, H. seminiger, H. transversalis

, , . H.

pallidulus , , . ,

. Hydradephaga .

:1) – ,

;2) – ;3) – , ;4) , .

Hydradephaga.

46

, , , ,

. Hydradephaga ( ).

Hydradephaga. ( . [38],

. [106]) , ,

( . [12]) , ( , )

, , .

. ,

, .

, , .

( 1) Hydradephaga .

( . 2).

Hydradephaga .,

,

Hydradephaga. [12,120, 143, 203, 234],

.,

, .

( . Hydradephaga).

( , , ,),

. , .

.

47

4%8%

13%

18%

22%

33%

?2%

1

21%

9%

62%

?8%

2

3%

57%

36%

1%?

3%

3

6%6%

18%

61%

?9%

4. 2. Hydradephaga (1),

(2), (3) (4) . : – , –

, – , – , – , – , – , – , – , – ,

– , – , – , –, – , – , –

, ? – , .

. [10] . ,

.1) –

) , , . , O. villosus

, . Gyrinidae

(G. caspius, G. suffriani, G. distinctus) (Phragmites spp.)

, . , ,

Hydradephaga , .

48

. ,

Hydradephaga (A.sulcatus, G. cinereus, R. suturalis, H. impressopunctatus). ,

,

(Lemna, Potamogeton) Hydradephaga, , ,

, . Dytiscidae ( Agabus, Ilybius, Hydroporinae),

, .

,

. D dimidiatus D.circumflexus, ( –

) (Carex spp.).,

Hydradephaga,

.2) –

, .

. , H. obliquus, H. variegatus H. maculatus (Charophyta), ( 50 ./ 2).

H. variegatus H. maculatus (Ceratophyllum spp.)

(Myriophyllum spp.), , .

Hydradephaga , (

).

( , ), .

3) – .

,

49

, , , .

4) – ,

. , , , . G. cinereus, 28

2007 ,. . ( )

Stigeoclonium sp., (Bacillariophyta) ( . 3).

. 3. ( ) Stigeoclonium sp.( ) G. cinereus. .

, (0 – 1 )

Hydradephaga, . ,

. , -

.

. ,

. H. fulvicollis, H. decoratus, H. palustris A. undulatus,

., H. maculatus, H. variegatus, H. confluens, H.

parallelogrammus, A. conspersus, A. nebulosus, H. grammicus

50

. 3) Hydradephaga

. 2000-2003 . . ,

(11 ), , , . . ( )

. (15 ),

. Hydradephaga (49 )

.

, Hydradephaga -, .

, ,, , ,

, .

Hydradephaga .

[4, 12, 38, 39, 106, 115, 131, 141] , , ,

Hydradephaga [47, 49, 51,53, 54. 62, 63]. ,

, .

Hydradephaga .

..

, ,

[50, 61]. [128], 3 : -, -, .

[143], . , , .

51

63 Hydradephaga, 4 : H. sibiricus, A. biguttatus, I. chalconatus D.marginalis . 4

: H. lineatocollis, H. discretus, A. paludosus A. bipustulatus.

Hydradephaga ( . 4),

– 2. 4

Hydradephaga ()

Haliplidae 4 2 3 1 8 5 1 1Noteridae 2 0 1 1 1 1 1 1Dytiscidae 33 13 35 21 44 24 28 22Gyrinidae 3 0 0 0 4 2 0 0

42 15 39 23 57 32 30 24. – , –

, Hydradephaga (57 )

. , , ,

( ), : ( ), ( ) ( ) ,

. Hydradephaga

(42 ). (3-4

)., , ,

. ( : H.

lineatocollis, H. discretus, A. biguttatus, P. maculatus) , ,, , H.

geminus, H. planus, L. minutus, R. suturalis C. fuscus.

Hydradephaga 0 35, - [34] 3,5 , .

[143] . , ,

52

. , Hydradephaga

..

H. obliquus, , : H. heydeni, H. sibiricus, H.

palustris, H. memnonius, L. hyalinus, P. maculatus, D. marginalis, G. paykulli.

, . , . [91]

(H. decoratus, H. striola, H.erythrocephalus, A. fuscipennis, Agabus adpressus Aubé, 1837, A. sturmii(Gyllenhal, 1808), Colymbetes paykulli Erichson, 1837)

. . [34] Hydradephaga (H. lineolatus, H.

flavicollis, L. hyalinus, I. chalconatus, D. marginalis, G. marinus) « »

) « » ( )., ,

( 20º ) ., Hydradephaga,

. , ,

. (

).

. [38], « »

. ,

, .

, .

78 Hydradephaga, 76% .

,

53

, Hydradephaga ( . 5, 2).

5 Hydradephaga

( )

Haliplidae 12 11 7 7 5 6 7 8Noteridae 2 2 2 2 2 2 2 2Dytiscidae 41 47 41 39 39 39 38 47Gyrinidae 4 4 4 3 3 3 2 5

59 64 54 51 49 50 49 6270 69

. – , – , – , –

, , H. zacharenkoi, P. lineatus, R. bistriatus D.

circumflexus. H.parallelogrammus H. cuspidatus

( ). Hydradephaga

(64) . , ,

. (A. paludosus H. memnonius),

, R.latitans,

. (59 ), H.

fluviatilis. 54 Hydradephaga,

(ICS 0,89 0,88 ). ,

, ( ,

), Hydradephaga . ,

. 51 .

H. discretus A.biguttatus ( ), H. enneagrammus,

.

54

. Hydradephaga, ,

, . H. fulvicollis, H. heydeni, H. flavicollis, H. fulvus, H.

palustris, H. decoratus, A. undulatus, R. exsoletus, C. striatus G. zonatus. R. exsoletus . ,

..

Hydradephaga (62 ) .

N. ceresyi ( ), H. immaculatus, H. obliquus, H.enneagrammus, A. didymus, A. paludosus, I. ater, R. grapii, G. natator, G.substriatus. A. didymus,

[52] .

, W

( . 6, . 4). C. lateralimarginalis, D. dimidiatus, D. circumflexus A. labiatus,

(10-30 ./ ),

. , R. bistriatus H.grammicus, ,

.

. 4. Hydradephaga

( 75%).

. 6.

6 (W, %)

Hydradephaga

W- 90,0 78,0

92,0 - 79,092,0 92,0 -

. – , –, –

Hydradephaga

,

55

. , ,

, Hydradephaga

.

, , ( ).

Hydradephaga (ICS=0,77)., ,

( . 5). 49 Hydradephaga,

L. hyalinus. Hydradephaga ,

, , ,

. 50 Hydradephaga, C. striatus R. latitans

. 2006 .

A. fuscipennis, [52].

(49 ), .

H. fulvus B. unistriatus.,

Hydradephaga. . ,

,

. Hydradephaga

, .

, , ,

, . ,

Hydradephaga, [17, 19, 32, 33, 38, 39, 115,

76, 151-153]. ,

56

, ,

. .

(). ,

Hydradephaga, ,.

. 56

Hydradephaga ( . 7, 2). 7

Hydradephaga ( )

Haliplidae 5 5 5 2 4 2Noteridae 2 2 1 2 2 1Dytiscidae 22 24 21 14 31 28Gyrinidae 2 1 2 0 0 0

31 32 29 18 37 3146 44

. – , – , – ,, – ,

, . Hydradephaga,

, . , 31 ,, H.

versicolor, P. maculatus O. villosus, . 32

Hydradephaga, (A. undulatus) .

29 , (12 ) . H. lineatocollis, H.sibiricus, H. discretus, H. memnonius, L. haemorrhoidalis, A. bipustulatus, A.paludosus, I. chalconatus, I. fuliginosus, I. subaeneus, G. natator.

H. sibiricus, I. chalconatus, . [85]

, . .

57

, .

., .

, ,, . 18

Hydradephaga. , (37 ).

H. memnonius,

.. ,

Hydradephaga ,. H.

heydeni, H. palustris, H. memnonius, L. hyalinus, I. ater, I. similis, C. striatus,D. marginalis. ,

.,

Hydradephaga,

.

. ( ,

, ). Hydradephaga

75 ( . 8, 2). Hydradephaga , – ,

. 34 , 22

, (« » , )

.

D. circumflexus ( . ). (49 ).

H. lineatocollis,.

58

8 Hydradephaga ()

Haliplidae 2 8 2 5 12 4 1 4 1 1Noteridae 2 2 2 2 2 2 2 2 2 2Dytiscidae 13 39 21 22 38 13 15 21 13 25Gyrinidae 0 5 1 5 6 0 0 0 0 2

17 54 26 34 58 19 18 27 16 3058 61 33 30

. – , – ( , , ,), – ( , ), –

, –

, , ,

22° , 20° . – : H.

fluviatilis, H. immaculatus, H. heydeni H. flavicollis. G. distinctus, G. natator, G. substriatus, G.

suffriani, G. caspius. 10-100

. G. suffriani 2001 2005 , 2006

,

( =100%, N=6-22 ./ 2). 51, 58.

H. fulvicollis, H. pubescens,D. circumcinctus, G. suffriani, G. paykulli ( ),

H. furcatus, H. variegatus, H. zacharenkoi, H. cuspidatus, H.parallelogrammus A. bipustulatus, .

, , R. latitans, .

. ,

[259]. ,

. ,

, .

Hydradephaga

59

. 19 ,

(N. crassicornis N. clavicornis), [127] (G. distinctus, G. suffriani),

[71, 127]. , ,C. lateralimarginalis,

. , .

Hydradephaga .

61 .,

: H. flavicollis, H. fulvicollis, H.heydeni, H. decoratus, H. pubescens, R. grapii, D. circumcinctus G. paykulli.

(60% Hydradephaga)

. ,,

, ,

.

. (2007)

, ,

. Hydradephaga

, .

Hydradephaga, .

58 , 52 , 6(H. tristis, H. obscurus, H. pubescens, H. versicolor, H. quinquelineatus G.marinus) . [132] ( 2).

H. musicus, [56].

(ICS=0,78), ,

(H. decoratus, A. undulatus A. sulcatus).

60

.

) (B. nasutus, H. parallelogrammus, H.confluens, H. pallidulus, H. cuspidatus, G. caspius)

(H. confluens, H. pallidulus).

. –

(Charophyta) – (H. obliquus, H. flavicollis, H. maculatus,

H. variegatus), . , H. flavicollis, H. maculatus H.

variegatus , H. obliquus, .

. 21 Hydradephaga,

29, – 54. H. grammicus,

-250. , , Hydradephaga

, , .

: , . ,

Hydradephaga ( 10 ),

[120] [110].

. , , .

26 Hydradephaga, 1 (L. hyalinus) [132].

(21 ) .. ,

(2-4 ), . (0-1

), 14 . , Hydradephaga

, . ,

61

. ,

, , , – 23 . Hydradephaga

, . ,

. 16 , 7 – .

H. pallidulus. Hydradephaga

, ()

-. ,

, Hydradephaga (56%

), . (H. obliquus, H. flavicollis, H. pubescens, H. obscurus, H.

tristis, H. quinquelineatus, H. versicolor, G. marinus) ,

– . – -

[31]. , , : ,

– , .

, ,

. H. variegatus, H. zacharenkoi, H. maculatus, B. nasutus, H.parallelogrammus, H. confluens, H. pallidulus, H. musicus, H. cuspidatus, L.poecilus, H. grammicus, C. lateralimarginalis, G. caspius, G. distinctus, G.suffriani.

Hydradephaga.,

, .

, Hydradephaga . 31

, 1 – G. marinus -. [82, 83]. 16

Hydradephaga,

62

2). 30 , D.

circumcinctus ( .). . [37, 38]

. 33 Hydradephaga.

19 , 5 – (H. flavicollis, H. fluviatilis, H. immaculatus, L. hyalinus, P

maculatus).

(26%), . , 26%

Hydradephaga. , ,

[38, 131, 143]. 18 ,

– . ,

, : ,

, .

.

. 27 Hydradephaga ( 2). H.

flavicollis, H. palustris, L. hyalinus P. maculatus. , Hydradephaga,

. Hydradephaga

,

, . 74%

, . , . Hydradephaga

«» -

[9, 38].

63

. , Hydradephaga,

. ,

( 2). ,

. 26

Hydradephaga. , )

(2-4 ),

Hydradephaga.

. , 2 (N. clavicornis C. lateralimarginalis), – 1 (C.lateralimarginalis).

, , .

, Hydradephaga – .

. 2 : A. bipustulatusH. discretus. 3 :

H. enneagrammus, H. geminus -C. lateralimarginalis.

Hydradephaga . 31 ,

, 1 . 2 (G. cinereus

C. lateralimarginalis), – 2 (G. caspius G.distinctus). H. enneagrammus,

4 (H. geminus, H. planus, A.conspersus R. suturalis). , ,

6 : H. lineatocollis, H.zacharenkoi, H. cuspidatus, H. discretus, H. planus A. biguttatus.

Hydradephaga , , ( . 9).

( ) Hydradephaga. C.

lateralimarginalis. 22 .

,

64

Hydradephaga, .

9 Hydradephaga (

)

1 2 1 2 1 2 1 2 1 2Haliplidae 1 0 0 0 2 0 0 3 0 1 0 0Noteridae 2 0 0 0 2 0 0 1 0 1 0 1Dytiscidae 18 5 2 2 17 6 3 21 10 11 1 19Gyrinidae 2 0 1 1 3 0 0 0 1 0 0 2

23 5 3 3 24 6 3 25 1 2226 3 31 25 11 13 22

. : – , – , –, – , – , – , –

(1 – , 2 – ( – , –

, , – , – ,

).

11 Hydradephaga,.

, ,,

.

, Hydradephaga () .

. , 4 : H.

geminus, A. conspersus H.enneagrammus N. ceresyi.

H. impressopunctatus, H.inaequalis R. suturalis, H.confluens. 24 Hydradephaga,

H. memnonius. ( )

3 :H. geminus, G. caspiusR. suturalis.

.

65

, .

(3 ) Hydradephaga . , .

25 .H. obliquus,

.Hydradephaga ,

. 13 , .

,

.,

Hydradepahaga, ,

. ,

, « »

[40]. . Hydradepahaga

, .

, . ,

– C. lateralimarginalis.

Hydradephaga , , .

H. obliquus, H.

enneagrammus N. ceresyi, ,

, (H.lineatocollis, H. discretus, H. memnonius, A. biguttatus).

.

66

Hydradephaga. , .

, , , .

, , .

. [36, 37, 38], ,, Hydradephaga .

61 , 35 , 26 [37].

2 (G. natator N. ceresyi) . 10, 2)

10 Hydradephaga (

)

Haliplidae 4 5 2 4 2 4 0 2 0Noteridae 2 2 2 2 1 2 0 2 0Dytiscidae 23 37 21 39 21 30 4 15 3Gyrinidae 2 1 0 0 1 1 0 1 1

31 45 25 45 25 37 4 20 447 46 38 21

. : – , – , –, – , –

. – , – [37].

, Hydradephaga (47 ) .

A. sulcatus, A. lineatus G. natator. (46 ), A.

canaliculatus, A. undulatus, H. fulvicollis, H. erythrocephalus, H. palustris, H.tristis, L. hyalinus, R. bistriatus, R. grapii.

, ( 39 21 ),

, . H. corpulentus, H. pallidulus, H.

flaviventris, H. enneagrammus, N. ceresyi, A. conspersus, H. grammicus.

,

. H. furcatus, N. crassicornis, L. haemorrhoidalis,B. unistriatus, H. ovatus, H. inaequalis, S. dorsalis, A. labiatus, I.quadriguttatus, C. fuscus, G. austriacus, H. grammicus, H. seminiger, H.transversalis, D. dimidiatus.

67

, ,

. Hydradephaga (4 )

, ,

. , ,

Hydradephaga () .

, , , ,

. . . [37,

38] , , – , .

H. variegatus, H. zacharenkoi, B. nasutus, H.cuspidatus, H. corpulentus, H. pallidulus, H. confluens, H. flaviventris, H.enneagrammus, H. parallelogrammus, N. ceresyi, L. poecilus, A. conspersus, A.lineatus, H. grammicus, C. lateralimarginalis, G. caspius.

H. fulvicollis, S. dorsalis, H. erythrocephalus, H. palustris,H. pubescens, H. striola, H. tristis, L. hyalinus, R. grapii, G. zonatus. H.corpulentus, H. flaviventris, S. dorsalis, H. erythrocephalus, H. striola A.lineatus .

, Hydradephaga. ,

, , ( ) ,

.

Hydradephaga . . [93],

, .

. [119], Hydradephaga. W [249]

, . ,

. ,

68

.

. ( ,

) , .

[87, 88]. Hydradephaga .

. , , - ( -)

, , ) [42, 63, 87, 88, 119, 139, 150, 164].

Hydradephaga ICS

. 11). 11

(ICS) Hydradephaga

2 3 4 5 6 7 8 9 10 11 121 0,66 0,81 0,81 0,66 0,83 0,61 0,62 0,77 0,11 0,72 0,442 - 0,67 0,64 0,85 0,71 0,75 0,56 0,63 0,14 0,60 0,463 - 0,90 0,59 0,92 0,64 0,68 0,83 0,13 0,74 0,444 - 0,56 0,88 0,61 0,70 0,85 0,12 0,77 0,445 - 0,63 0,72 0,58 0,53 0,14 0,51 0,456 - 0,68 0,67 0,83 0,14 0,79 0,477 - 0,64 0,65 0,14 0,68 0,548 - 0,71 0,08 0,67 0,469 - 0,08 0,83 0,5310 - 0,10 0,1711 - 0,64

. : 1 – , 2 – , 3 – , 4– , 5 – , 6 –

, 7 – , 8 – , 9 – ( , , .,), 10 –

, 11 – , 12 – .

, ICS Hydradephaga

. Hydradephaga ,

69

,

. ICS 0,17 .

, .

ICS: « – » (0,90),

– » (0,85), – » (0,92), «

– » (0,88), « – » (0,85).

Hydradephaga –

. , ,

. « –»

, . « –

» (

), ,

. « – » ,

, ,

, . . « –» ,

, , .

, , .

ICS 0,57., Hydradephaga

. , , , , .

[143], ICS 0,45, 0,78. , Hydradephaga

, ( )

70

, , ,

. , ICS

Hydradephaga, .

, ,

, .

( . 5, 6).

1 2

3. 5. ,

Hydradephaga . 1 – 0,15, 2 – 0,6, 3 – 0,8.. 11

, 0,15 ,

. , 15% Hydradephaga

, .

, . ICS 0,6,

,

71

. , 0,8 ,

« – » « – –

– – »).

:-

Hydradephaga, ;-

( . [10]) ,

;-

, ;- ,

, ,

. ( .);

- ,

. Hydradephaga;- -

.,

, , , ,

, , ICS

( 7, 8, 10, 11 12 . 5 0,8).

, ( )

,

72

.

, W ( . 12).

12 (W, %) Hydradephaga

1 2 3 4 5 6 7 8 9 10 11 121 - 89,2 77,9 73,7 94,1 81,5 85,7 73,9 74,6 57,1 76,8 71,42 52,4 - 51,5 47,4 88,2 55,4 77,1 50,0 49,3 42,9 50,0 53,63 84,1 94,6 - 85,5 88,2 93,8 94,3 84,8 83,6 71,4 82,1 75,04 88,9 97,3 95,6 - 91,2 95,4 97,1 93,5 91,0 71,4 91,1 82,15 50,8 81,1 44,1 40,8 - 47,7 71,4 50,0 40,3 42,9 41,1 50,06 84,1 97,3 89,7 81,6 91,2 - 97,1 80,4 82,1 71,4 85,7 78,67 47,6 73,0 48,5 44,7 73,5 52,3 - 56,5 49,3 42,9 55,4 60,78 54,0 62,2 57,4 56,6 67,6 56,9 74,3 - 59,7 28,6 60,7 60,79 79,4 89,2 82,4 80,3 79,4 84,6 94,3 87,0 - 42,9 91,1 89,310 6,3 8,1 7,4 6,6 8,8 7,7 8,6 4,3 4,5 - 5,4 10,711 68,3 75,7 67,6 67,1 67,6 73,8 88,6 73,9 76,1 42,9 - 96,412 31,7 40,5 30,9 30,3 41,2 33,8 48,6 37,0 37,3 42,9 48,2 -

. . 11.

, W Hydradephaga ,

63%. Hydradephaga ,

, ICS. , ,

Hydradephaga , W. ( . 6)

, , .

. 6 , W=70% Hydradephaga

. . [119],

. , , ,

, ( 1, 3, 4, 6, 9 11,

). ,

.

73

. 6. , Hydradephaga . –

70%. . 11.

4

6

3 1

11

2

5

7

12

8

10

9

74

, , ,

. « –

– » 2, 5 7),

W .

( 2) ,

6). Hydradephaga ( 8),

(12) , ,

, , . ,

, , .

, ICS W , Hydradephaga

.

.

Hydradephaga . Hydradephaga

. [38, 39]. ,

Hydradephaga, , . , .

, .

[76]. Hydradephaga

[32, 33, 38], [38].

75

, Hydradephaga, . . .

[108, 109,106, 112, 113], [108],

., ,

Hydradephaga .

. . [114, 115]

.. [66, 67] , ,

A. canaliculatus.

. [20] . . , L. poecilus

2 , 1:1.

.. [165] R. suturalis

,

.. . [17, 19, 151, 152, 154] ,

,

. Hydradephaga: Brychius elevatus

(Panzer, 1794), Hydroporus nigrita (Fabricius, 1792), G. granularis,Nebrioporus airumlus (Kolenati, 1845), Deronectes latus (Stephens, 1829), A.paludosus, I. fenestratus, L. hyalinus, L. minutus, P. maculatus.

, G. bilineatus, H. angustatus I. fuliginosus. -

H. ruficollis, N.crassicornis, H. ovatus, H. inaequalis, H. impressopunctatus.

, . [131] . [133,135] -

, , . , . . .

.

76

. [133, 135]

(Heteroptera). ,

.

. ,

. ,, .

Hydradephaga. , . . .

[118] C. paykulli, A.sulcatus, A. canaliculatus H. ruficollis. . [141, 143]

( ,, .).

Dytiscus latissimus Linnaeus, 1758 G.paykulli, – B. elevatus, Oreodytes sanmarkii (C.R. Sahlberg, 1826),D. latus, O. villosus.

, Hydradephaga .

, , ,

, .,

Hydradephaga , : H. obliquus –

, H. lineatocollis,H. discretus, A. paludosus – , , H.fulvicollis, H. furcatus, H. decoratus –

, , , H. immaculatus – ,

, H. heydeni – , , , , H. zacharenkoi

– , , H. flavicollis, H. fulvus, H. maculatus – , ,

(

77

), B. nasutus, B. unistriatus, H. parallelogrammus – , (

), H. confluens – ( ),

, H. pallidulus, H. enneagrammus, N. ceresyi – , H. memnonius – :

, , – ,, , A. biguttatus, I.

chalconatus – , H. fluviatilis, H. versicolor, L. hyalinus,P. maculatus, I. fenestratus – , ,

, , , R. bistriatus –

, , , R. latitans – ,

, ,H. grammicus – , G. caspius, G. paykulli, G.natator, G. substriatus, G. suffriani, G. distinctus –

, , , ( G. paykulli) , O. villosus –

.

. , . )

, ,

Hydradephaga ( . ). 2001-2002 .

H. lineatocollis A. paludosus, , (P. caesus, H. inaequalis, L. poecilus, I. fuliginosus,

I. subaeneus, I. quadriguttatus, R. frontalis, D. circumflexus,Graphoderus Hydaticus).

: N. clavicornis, L. minutus, A. bipustulatus, R. suturalis C. fuscus.,

. O. villosus, ( .

). , (

) .

78

. 7).

. 7. Orectochilus villosus (O.F. Müller, 1776). . ,

. . .

, Hydradephaga,

. ,

. ,

,

Hydradephaga .

. Hydradephaga , , ,

[47].

79

Hydradephaga .Hydradephaga , -

, .

, ,

. ( 100%) ,

, , , .).

,

.

. Hydradephaga

, ( ) (

). , , 75%

Hydradephaga. , ,

. ,

Hydradephaga. Hydradephaga ,

. Hydradephaga

, , ,

.,

. ( )

. ( , , ) ,

. ,

80

, . ( ,

), ,

, ( ,

) , .

.,

Hydradephaga,

. , :

, ,

.,

Hydradephaga . [38]

, , . ,

, .

(, ,

.) .

81

4,

HYDRADEPHAGA -

. Hydradephaga

[36]. 5

. [95]. Hydradephaga .

, Hydradephaga .

. [38] . [106],

( ./ ). . , . – .

Hydradephaga.

Hydradephaga . [131].

. , Hydradephaga .

. . [18].

Hydroporus. [41].

Hydradephaga , . [145]

2 (P. caesus A. sulcatus).

Hydradephaga , Hydradephaga

. :

. . ,

Hydradephaga,

82

, , .

, ,

.

, . ,

.,

–,

, Hydradephaga .

, ,

, .,

, .

.

, Hydradephaga

. –

( . ) . ,

– « » (. [10])

.

,

. . [96], « »,

( . « »).

83

, .

(), ,

. ,

( ) )

( , .), ,

.

[58, 61]: –

– – ,

10,9-12,3 . ( , 2001 .), ,

+5 – +7º , , ,.

–

, 12,4-14,2 , 12-15º .

– – , ,, ,, 14,3-15,4 ,

, , 15º . – ,

, ( – ).

, 24-30º , (Carex spp., Typha spp., Phragmites spp.),

. ( – ) ,

– ). – –

, – . Hydradephaga

. ,

. , – 2001

84

. . (37) , 26 ,

– 24. , (H. zacharenkoi,

H. planus, A. bipustulatus, C. fuscus, A. sulcatus, D. dimidiatus .). H. fulvicollis. –

H. planus Dytiscus, ,

( . 3). (2006), ,

Hydradephaga . , 2006 .

. 2: H. planus A. labiatus. ( )

6 Hydradephaga (H. planus, A. labiatus, C. fuscus, D.dimidiatus, D. circumflexus, C. lateralimarginalis A. bipustulatus, A.conspersus, A. labiatus, C. fuscus, D. dimidiatus, D. circumflexus ).

, 10º , 8

(N. clavicornis, N. crassicornis, H. planus, H. angustatus,H. inaequalis, . impressopunctatus, G. bilineatus, A. labiatus).

, (4 ),

( ) (8,5º ). ,

, .

2007 . . Hydradephaga

( . 13).

13 (N, ./ 2) (B, 2) Hydradephaga

2007 .. .

N B N B N B N BHaliplidae, imago 0,00 0,00 0,04 0,10 0,43 0,86 0,14 0,31Noteridae, imago 0,50 2,17 0,09 0,43 0,00 0,00 0,21 1,66Dytiscidae, imago 0,17 0,17 0,47 2,28 0,00 0,00 0,29 37,96Dytiscidae, larvae 0,00 0,00 0,04 1,09 0,14 2,57 0,06 2,03

0,67 2,33 0,64 3,90 0,57 3,42 0,70 41,97

85

. 0,30 1,10 ./ 2, 2,30 4,62 2.

G. bilineatus, N. crassicornisN. clavicornis. A. labiatusIlybius spp., – .

(D=1,04, 0,85 1,0 ). . .

, (3,43 – 95,522). H. zacharenkoi,

– Ilybius spp. (D=1,13). .

Noteridae, H. zacharenkoi A. labiatus, – D. dimidiatus (D=5,57).

,

. Hydaticus, Graphoderus Rhantus, Haliplidae

(H. obliquus, H. immaculatus, H. variegatus , , H. furcatus, ).

– P. lineatus, .

. 2006 . . 21

Hydradephaga, . 3 , . Adephaga 2

(16 ).

( 12-19º ) .

Hydradephaga (09.04.2006 24 , –

23). ,

, . 2 9 2006 Hydradephaga .

3 , 7 21 , (N.crassicornis, N. clavicornis, H. angustatus, H. impressopunctatus, H.parallelogrammus, H. inaequalis, H. geminus, B. nasutus, L. minutus, R.bistriatus, G. cinereus).

, 2006 . , .

2001 , »

86

7-14 , – 15-20 ., (P.

caesus, H. zacharenkoi, H. parallelogrammus, R. suturalis, A. sulcatus .). G. distinctus

B. nasutus, R. frontalis A. canaliculatus, 2001 , 2006 – 2 .

(22 29.04.2006) G. suffriani.

2007 . ,. Hydradephaga.

( . 14).


2007 .. .

N B N B N B N BHaliplidae, imago 0,33 0,93 0,20 0,64 0,00 0,00 0,36 0,96Gyrinidae, imago 0,00 0,00 0,17 1,73 0,00 0,00 0,54 5,15Noteridae, imago 1,87 8,73 1,73 7,96 0,00 0,00 0,33 2,63Dytiscidae, imago 0,20 15,60 4,63 37,42 0,53 5,33 3,52 95,98Dytiscidae, larvae 0,00 0,00 0,07 2,20 0,00 0,00 0,34 23,48

2,40 25,26 6,80 49,95 0,53 5,33 5,09 128,21

, ,

Hydradephaga . , .,

, . ( )

.. N.

crassicornis, G. cinereus. D (2,49 3,90 ).

B. unistriatus, N. crassicornis, G. bilineatus, H. impressopunctatus

H. inaequalis. Hydradephaga L.haemorrhoidalis (Fabricius, 1787) RhantusHydaticus. N. crassicornis (D=2,49), H.

87

impressopunctatus (D=2,56), L. haemorrhoidalis (D=3,03) R. frontalis(D=3,56). .

G. bilineatus, – R. frontalis(D=2,15).

G. bilineatus, (C. fuscus, Rhantus, Ilybius, A.

canaliculatus, C. lateralimarginalis). D A. canaliculatus (4,45), C. lateralimarginalis (7,11).

– A. fuscipennis,

H. grammicus Ilybius. 2006 . H.

maculatus, 2000-2001 2008 . . – .

Hydradephaga

. ,

. , 20 2006 .

Hydradephaga ( 29 34)., ,

, ,

. ( . 8).

0

510

1520

25

3035

40

25.02

.2006

08.03

.2006

19.03

.2006

02.04

.2006

09.04

.2006

16.04

.2006

22.04

.2006

29.04

.2006

07.05

.2006

20.05

.2006

29.05

.2006

10,9 11,5 12 12,9 13,2 13,7 14 14,3 14,7 15,2 15,5

,

. 8. Hydradephaga . 2006 .

88

2007 . . . . ,

. H. inaequalis, – H. inaequalis

H. impressopunctatus ( D=1,91). Hydradephaga

, (B. nasutus, H. decoratus, H. inaequalis, H.

impressopunctatus, G. bilineatus, L. poecilus), ( . 15),

– (C. lateralimarginalis, D=12,75).

15 (N, ./ 2) (B, 2) Hydradephaga .

2007 .

N B N B N BHaliplidae, imago 1,04 2,66 0,50 1,00 2,82 5,64Haliplidae, larvae 0,17 0,33 0,00 0,00 0,00 0,00Noteridae, imago 17,74 52,55 0,50 1,67 4,62 22,56Dytiscidae, imago 14,26 216,91 2,17 8,50 18,97 84,10Dytiscidae, larvae 0,00 0,00 0,00 0,00 0,26 4,62

33,21 272,45 3,17 11,17 26,67 116,92

Hydradephaga ,

. , ,

.,

, , (P. caesus,

N. clavicornis, N. crassicornis, H. angustatus, H. planus, B. nasutus, H.geminus, L. minutus, L. poecilus, I. obscurus, I. similis, I. fuliginosus, C. fuscus,R. suturalis, H. transversalis, G. cinereus).

, ,

, –.

. 2007 . . ,

, , ,

89

. Hydradephaga ,

( . 16).


2007 .

N B N B N BHaliplidae, imago 0,39 1,24 2,70 6,30 0,00 0,00Noteridae, imago 0,50 1,75 0,25 1,00 0,00 0,00Dytiscidae, imago 1,27 174,70 18,98 944,68 2,03 1232,81Dytiscidae, larvae 0,81 120,61 0,00 0,00 0,00 0,00

2,97 298,30 21,93 951,98 2,03 1232,81

C.lateralimarginalis,

. (D=10,59) D. dimidiatus.

H.inaequalis P. lineatus.

C. lateralimarginalis D. dimidiatus(D=23,33 18,6 ).

( , , )

, Hydradephaga ., (H.

inaequalis, C. fuscus, H. transversalis). – A. labiatus C.

lateralimarginalis. – I. obscurus, I. similis, I. fuliginosus, I. subaeneus, R. bistriatus, R.

latitans, H. transversalis H. seminiger, , ,

. , .

, (I. fenestratus, C.lateralimarginalis), – -

. 2000 ., , . ,

(H. zacharenkoi, P. caesus, N. crassicornis, N.clavicornis, H. planus, H. impressopunctatus, H. parallelogrammus, G.bilineatus, H. geminus, H. ovatus, L. minutus, L. poecilus, L. haemorrhoidalis,

90

A. undulatus, A. labiatus, A. bipustulatus, C. fuscus, R. suturalis, D. dimidiatus,A. sulcatus).

2007 . ,. .

. . H. geminus, H.

impressopunctatus, G. cinereus H. transversalis, Hydradephaga.

C. lateralimarginalis. G. cinereus (D=7,38) C.

lateralimarginalis (D=9,17). ( . 17),

Ilybius, C. fuscus R.suturalis. Hydradephaga

G. cinereus, H. transversalis, G. cinereus G. austriacus.


2007 .

N B N BHaliplidae, imago 2,30 5,80 0,40 1,60Haliplidae, larvae 0,00 0,00 0,18 0,66Dytiscidae, imago 1,76 132,92 3,93 273,67Dytiscidae, larvae 8,40 105,05 25,27 262,31

12,46 243,77 29,78 538,23

G. cinereus(D=13,46), C. fuscus (D=10,15) R. suturalis (D=10,17).

, , Adephaga

. –

. 7-10 .,

( 1-1,5 ) , .

(2006) Hydradephaga .

2-5 (2001), .

91

, . , ,

.

( . 9).

05

101520253035

04.03

.2007

07.04

.2007

12.05

.2007

10.06

.2007

07.07

.2007

01.08

.2007

02.09

.2007

13.10

.2007

N

01002003004005006007008009001000

B

N, 2 , 2

0

2

4

6

8

10

12

14

16

04.03.2007 07.04.2007 12.05.2007 01.08.2007N

01002003004005006007008009001000110012001300

B

N, 2 , 2

. 9. Hydradephaga . 2007 .

, . ,

.

.

. .

Hydradephaga,,

(H. impressopunctatus, R. suturalis, Ilybius spp. ).

, Hydradephaga .

, .

. (2001, 2005, 2006,2008) ,

.

92

. ( – ).

. ( – )

,

. ( )

, (2003, 2007). (2001, 2005,

2006, 2008) – , ,

. ( ) , ,

, , .

, , , , .

( – ). , (2006) –

.

. .

Hydradephaga, .

H. fulvicollis, H. pubescens D. circumcinctus. –

R. grapii, R. latitans, H. transversalis, H. seminiger, , Ilybius,

(Rhantus, Hydaticus) (Ilybius). ,

. 2007 .

Hydradephaga 0,10 6,55 ./ 3, – 0,90 31,75 3.

Noteridae ( . 18). N. crassicornis (D=2,54) A. undulatus (D=2,30).

93


2007 .

N B N BHaliplidae, imago 0,00 0,00 0,54 1,91Haliplidae, larvae 0,00 0,00 0,05 0,19Noteridae, imago 0,67 4,67 3,13 14,51Dytiscidae, imago 0,13 1,33 0,60 7,41

0,80 6,00 4,32 24,03

, 4

. (4-6° , –

7-9° , 12° ) , ,

Hydradephaga ( . ).

Hydradephaga, . ,

H. furcatus,.

, ,

, , . ,

P. caesus, H. ruficollis, H. heydeni, H. fluviatilis, H.immaculatus, P. lineatus, H. ovatus, A. undulatus, A. labiatus, I. fenestratus, R.frontalis, R. latitans, C. fuscus D. dimidiatus, C. lateralimarginalis.

, ,

H. inaequalis, I. subaeneus, I. quadriguttatus, I.similis, A. canaliculatus, A. sulcatus).

2007 . Hydradephaga.

, N. crassicornis (D=7,20).

H. fluviatilis, C. lateralimarginalis (D=16,93).

Hydradephaga, .

94

, H. ovatus R. latitans. 19).


2007 .

N B N B N BHaliplidae, imago 3,11 9,40 7,60 10,40 0,90 3,80Noteridae, imago 12,95 60,09 0,00 0,00 2,00 8,40Dytiscidae, imago 2,77 65,02 0,40 288,13 1,90 42,75Gyrinidae, imago 0,13 1,67 0,00 0,00 0,00 0,00

18,96 136,18 8,00 298,53 4,80 54,95

(2002,2004, 2007, 2008 .)

Hydradephaga, .

(2001, 2005, 2006 .) , . 2

(R. latitans, A. labiatus). 2001 2006 H. zacharenkoi G.paykulli. ,

,

. 2007 .

Hydradephaga. P.

caesus H. fluviatilis, C. lateralimarginalis (D=13,72).

H. inaequalis, – C. lateralimarginalis (D=4,90).

Hydradephaga .

. H. inaequalis,

P. lineatus L. poecilus, ( . 20).

C. lateralimarginalis, (D=15,03).

95


2007 .

N B N B N B N BHaliplidae, imago 1,00 3,20 0,10 0,40 0,30 0,60 0,80 2,20Haliplidae, larvae 4,62 12,19 0,00 0,00 1,20 1,00 0,00 0,00Noteridae, imago 0,00 0,00 0,20 1,00 0,00 0,00 0,60 2,40Dytiscidae, imago 0,17 376,67 0,80 4,20 0,10 0,40 6,50 240,30Dytiscidae, larvae 1,58 4,69 0,10 24,00 0,00 0,00 0,00 0,00

7,37 396,75 1,20 29,60 1,60 2,00 7,90 244,90

, ,

. ,

Hydradephaga. 3.08.2007 , , .

, .

, , ,

. – ,

, (H. ruficollis, H. fluviatilis) (G. natator,

G. substriatus, G. distinctus, G. caspius, G. suffriani). 10-100

. - C. lateralimarginalis;

0,3-1,0 . IlybiusI. fenestratus), Rhantus ( R. suturalis) Hydaticus

. 2007

( ) ( ).

96

H. fluviatilis, 1,3 ./ 2 2,8 2 ( . 21).


2007 .

N N BHaliplidae, imago 1,30 2,80 0,50 2,50Dytiscidae, imago 0,00 0,00 6,08 609,67Dytiscidae, larvae 0,00 0,00 0,25 8,75

1,30 2,80 6,83 620,92

. P. lineatus ( 11,75 ./ 2)

– C. lateralimarginalis (B 1490,00 2, D=24,32)., .

. P. lineatus 2007

,

3 .,

Hydradephaga , ( . 10).

0

1

2

3

4

5

6

7

8

9

09.03.2007 13.05.2007 09.06.2007 08.07.2007 01.09.2007

N

0

50

100

150

200

250

300

350

400

450

B

N, 2 , 2

02468

101214161820

09.03.2007 08.04.2007 13.05.200709.06.2007 08.07.2007 28.10.2007

N

050100150200250300350400450500550600650

B

N, 2 , 2

. 10. Hydradephaga 2007 .

, 2 :

( ) ( ). ,

97

.

Hydradephaga .

C. lateralimarginalis. Hydradephaga

. , . – ( .

).

Hydradephaga ( . 11).

4 5. 11. Hydradephaga

: 1 – (W=55%), 2 – ( W=55%), 3 –

W=80%), 4 – ( – , W=60%), 5 – ( , W=100%). : . – , . – ,

. – , . – .

, Hydradephaga,

.

. , Hydradephaga

,

1 2 3

98

. ,

(H. fluviatilis), (D. dimidiatus, A. labiatus) (R. suturalis)

, .

, A. sulcatus G.cinereus.

, .

, Hydradephaga

, .,

. , ,

. ( ) ,

, . . ( 2003 .)

, . W 55 80%

Hydradephaga .

, .

, , . ,

, . Hydradephaga

, . (

) ,

().

. ,

Hydradephaga, .

99

, , ,

Hydradephaga .

, Hydradephaga .

, .

Hydradephaga ,

.

. Hydradephaga

1,15 81,20 ./ 2, – 13,08 1162,00 2. 2007 .

– 1002,402. H. geminus,

Ilybius(D=21,25) A. bipustulatus (D=9,24).

H. geminus, : C. fuscus C. lateralimarginalis.

(D=12,70 15,00). .

. 29,32 ./ 2, 396,80 2.

H. geminus, – C. fuscus (D=12,47) R. suturalis (D=10,13).

, ,

. C. fuscus, A. bipustulatus Ilybius,

H. lineatocollis, A. bipustulatus, H. geminus ( . 22).

Hydradephaga.,

.

H. geminus. , .

100

22 (N, ./ 2) (B, 2)

Hydradephaga 2007 .

N B N B N B N BHaliplidae, imago 3,64 10,91 0,91 2,72 0,93 2,80 0,47 1,88Haliplidae, larvae 0,67 2,00 1,81 3,14 2,56 4,37 0,00 0,00Noteridae, imago 1,00 9,67 1,56 8,07 1,44 8,67 0,00 0,00Dytiscidae, imago 25,95 138,89 11,49 495,05 20,20 287,49 1,11 51,11Dytiscidae, larvae 17,17 514,70 2,55 54,93 4,19 93,48 4,83 124,10

48,43 676,17 18,32 563,90 29,32 396,80 6,41 177,09

, Hydradephaga.

, ( . 23).

23 (N, ./ 2), (B, 2) Hydradephaga

N B III V III V III V

81,20 20,22 1002,40 361,68

N: H.geminus,

B D: Ilybiusspp., larvae

N: H.geminus, B D: C.fuscus

10,30 9,00 327,30 1162,00

N: Ilybiusspp., larvae, A.bipustulatus,

larvae. B D:Ilybius spp.,

larvae

N: N.clavicornis, H.

cuspidatus B D: C.

lateralimarginalis

. : – , – . .

, Hydradephaga

. ,

, . ( . 12).

101

0

10

20

30

40

50

60

18.03.2007 05.05.2007 27.07.2007 20.10.2007

N

0100200300400500600700800

B

N, 2 , 2

. 12. Hydradephaga

.

, Hydradephaga

.

Hydradephaga

,

.

, , , 3 : H. geminus

H. enneagrammus N. ceresyi.

. H. enneagrammus , N.ceresyi – .

, Hydradephaga ( . 24).

24 Hydradephaga

2007 .. .

18.03. 05.05. 02.06. 17.03. 01.07.N B N B N B N B N B

H. enneagrammus 4,80 24,80 0,80 4,00 8,00 18,00 1,2 4,7 31,33 161,99H. enneagrammus,larvae 0,80 7,87 1,60 16,00 3,00 18,00 - - 42,00 252,00

N. ceresyi 0,80 7,87 1,60 16,00 3,00 18,00 - - - -N. ceresyi, larvae 0,93 3,47 4,80 23,20 4,00 22,00 - - - -A. conspersus - - - - 2,00 46,00 - - - -

6,53 36,13 7,20 43,20 17,00 104,00 1,2 4,7 73,33 413,99: – , . –

, . – .

, 2007 . Hydradephaga

.

102

H. enneagrammus (D=4,98). , –

. N. ceresyi (D=4,82).

, . N.

ceresyi, 2 ( 1,60 3,00 ./ 2). Hydradephaga H. enneagrammus,

A. conspersus (D=6,78). ( ),

. , Hydradephaga

, .

,

Hydradephaga .

, . ,

( )

. Hydradephaga

,

, . ,

. Hydradephaga

, , .

. Hydradephaga. ,

,

.

103

5 HYDRADEPHAGA

Hydradephaga . , Hydradephaga.

, .

. [31].

. Hydradephaga. [70] (

), . [173] ( ), . [153]), . . [130] (

), . [157] ( ). , Hydradephaga, . [161-163], .

[117] . [141]. . [195-210] . [253].

, Hydradephaga . , .

. [213, 231, 233, 234].. [231]

Agabini.

[131, 162, 163, 234]. .

Hydradephaga , ,

, . , ,

. 1999-2008 .

, Hydradephaga . .

Hygrobiidae [= Paelobiidae].H. hermanni,

( 1).. [71] . [204] ,

, ,

,

104

, . , H. hermanni .

Haliplidae..

[71, 204] .

P. caesus , . , Elodea

Ceratophyllum [71]. ,

. . 2007 .

, . ,

, [204]. , . ,

, – .H. lineatocollis

– . , , , .

, .

H. fulvicollis – , , – . ,

, , , ( ). ,

, – , –

. () H. fulvicollis –

, , .

H. furcatus – , , 2007 .

. , , , , , .

H. ruficollis H. fluviatilis , , .

( , ,

105

), H. ruficollis 3 . H. ruficollis

Juncus Phragmites, [71].

, . , ,

. . [204],

, . ( ). ,

, , ,

, , .H. heydeni , ,

. , ,

. , ,

. , , H. heydeni .H. immaculatus .

, , (2002). , , , .

, .H. zacharenkoi ,

[241]. , (2000-2001, 2006-2007) .

H. zacharenkoi , ( . 13.). ,

, , .

H. maculatus ,. ,

. H. obliquus, H. variegatus, H. fulvus, H. flavicollis H. sibiricus

, . H. variegatus H. flavicollis

. , , .

106

. 13. Haliplus zacharenkoi Gramma et Prisny, 1973 – . . .

, , , , ,

, , , . , (P. caesus, H. lineatocollis, H. zacharenkoi)

. , P. caesus, H. ruficollis, H. fluviatilis, H.

heydeni H. furcatus , H.lineatocollis – .

(Peltodytes), (Haliplus) [71, 204].

, (

1). Noteridae. 2 :

N. crassicornis N. clavicornis. , . N. crassicornis

, . N. clavicornis – Hydradephaga,

. (

). ,

., , .

– , . [127,

203], . ,.

.

107

1 . ( 1).

Dytiscidae. 5. ( [162],

[203], [234]), .

, 2 [162],

5 .

:1. , 1 (30 ): L.

haemorrhoidalis, B. unistriatus, B. nasutus, H. versicolor, H. decoratus, H.discretus, H. pubescens, H. palustris, H. erythrocephalus, H. striola, H. tristis,S. dorsalis, G. bilineatus, H. cuspidatus, A. paludosus, A. undulatus, R. grapii,R. bistriatus, C. striatus, L. hyalinus, H. seminiger, H. grammicus, G. zonatus,D. dimidiatus, D. marginalis, D. circumflexus, D. circumcinctus, C.lateralimarginalis. , A. didymus R. exsoletus,

. . . [234] ,

1 . ,

, ( H.seminiger , C. lateralimarginalis – ).

, . C. lateralimarginalis

– .

, , 5 , 2-4

( 1). ,

().

(L. haemorrhoidalis, G. bilineatus, A. undulatus, Dytiscus), – , (C.

lateralimarginalis, A. paludosus). Rhantus Hydaticus,

.2. , ,

: H. angustatus, H. ovatus, A.labiatus, R. frontalis, R. latitans, C. fuscus, A. sulcatus, A. canaliculatus, H.transversalis (9 ). H. angustatus

108

2002 . , ,

, , – .

, .,

, H. angustatus. A. labiatus R. latitans

( , ) – [54].

( , ) A. labiatus

(2000, 2006 .). – . A. labiatus

, R. latitans – . Acilius ,

. , 2007 .

. . . ,

.H. ovatus, R. frontalis, C. fuscus H. transversalis

. 2007 . , (C. fuscus) (H. ovatus, H. transversalis),

. 2007 ( C.

fuscus 10 , ). ,

, ., ,

, (Cladocera, Diptera)

. , , H. transversalis

, 13 . H. ovatus, R. frontalis C. fuscus

, (N=6,5-13,9 ./ 2) 1 – 3 .

28 . H. ovatus C. fuscus, , R. frontalis H. transversalis –

, .3. , – ,

: H. enneagrammus, P.

109

lineatus, A. conspersus, A. nebulosus, L. poecilus, G. cinereus, G. austriacus. (7). ,

. A. conspersus, A.nebulosus L. poecilus , ,

– ( , , .), Graphoderus

[203, 234]. P. lineatus, 2007.

, , [63]. ,

2007 , P. lineatus ( ,

) , .

4. , , (5 ): H. geminus, H. inaequalis, H.

impressopunctatus, H. parallelogrammus, H. confluens. , :

, ( H. confluens), ( ),

. , ,

, .

, . H. confluens ,

– , .5. R. suturalis ,

(2006) , – .

, . ( ,), . ,

R. suturalis , . . C. fuscus,

, [162, 201, 234] ,

( . ), [178] , . .

A. fuscipennis [162, 234]. ,

– . ,A. fuscipennis ,

.

110

. .,,

, .I. chalconatus (Panzer, 1796) [234].

. [162] , (Ilybius wasastjernae (Sahlberg, 1824))

, , ,, .

. ,I. chalconatus,

. 7 :

P. maculatus, I. fenestratus, I. fuliginosus, I. ater, I. similis, I. quadriguttatus, I.subaeneus. P. maculatus ,

[203].Ilybius –

, – ,

. , , ,

, . – .

, – , . I. fenestratus ,

, – , , , , ,

. – ,

, .

, ( . 4).

. 6 , [234]

, : H. planus, H. memnonius, N. ceresyi, A. bipustulatus, A. biguttatus, L.

minutus.H. planus, N. ceresyi L. minutus

( 25 23-28º ), –

. , [162] - [234] L. minutus ,

111

. H. planus L. minutus,

, .N. ceresyi – ,

. A. bipustulatus – , ,

, , ,

( , , , .).

, , – .

, – .

A. biguttatus – , ( )

, . ,

. , ,

. 10-15º – .

, , , .H. memnonius ,

. ,

. 2005 . , ,

. H. memnonius ,. . . [234]

, .

Gyrinidae.,

[127, 253]. , , , – .

, – .

( O. villosus – ), 20-40 . . [71] , O. villosus,

,

112

, , ,, .

, , , [71,

127]. ,

. , , , .

Hydradephaga.,

. ,

, , ,, (

) . ,

Ilybius ( 10 ./ 2)., (A. fuscipennis)

( , ), . [234].

, ,

(). Haliplidae, H. ovatus, Dytiscus Cybister)

(, , , ).

()

, . ,

, , , ,

, ., ,

Hydradephaga .

. , Haliplidae, , , ,

Dytiscidae , (Hygrobiidae, Noteridae Gyrinidae) – .

( . 14).

113

70,870,97

0,97

9,71

6,806,80 3,88

1 2 3 4 5 1(4) ?1

6,67

20,00

6,67

40,00

26,67

1 4 5 1(4) ?2

100,00

1 3

78,67

1,33

1,33

9,338,00 1,33

1 2 3 4 5 ?4

. 14. Hydradephaga. 1 – , 2 – Haliplidae, 3 – Hygrobiidae, Noteridae,Gyrinidae, 4 – Dytiscidae. .

, ? – , .

Hydradephaga ,

.

. .

(C. fuscus, R. suturalis, L.minutus)

, , . .

Hydradephaga , . ( , , ) , (H. hermanni, )

( , ). Hydradephaga (

, , , , .),

114

[61]. 10 ,

Ilybius, Rhantus Hydaticus ( )

. , [234].

. , – , .

, . , , H. lineatocollis.

. , ,

Hydradephaga, , .

.. [131] , Hydradephaga

3 (2-5 , 7-18, 26-42 ), .

. ,

. , .

, . ,

( . 25, 1). 25

Hydradephaga

, ,

Haliplidae 1 2,3-4,2 15Noteridae 1 3,5-4,5 2Gyrinidae 1 4,0-8,0 10Dytiscidae: Hydroporinae, Laccophilinae 1 1,7-5,3 35Hygrobiidae 2 8,5-10,0 1Dytiscidae: Copelatinae, Agabinae, Colymbetinae,Dytiscinae: Acilius, Graphoderus, Hydaticus. 2 5,8-18,0 35

Dytiscidae: Dytiscus, Cybister 3 24,0-39,0 5. – . [131]

115

, Haliplidae,Noteridae Gyrinidae –

2,3 – 8,0 . .

Hydroporinae Laccophilinae, 1,7 5,3 .

, 5,8 18,0 . Copelatinae, Agabinae, Colymbetinae,

Dytiscinae: Acilius, GraphoderusHydaticus. Hygrobiidae – H. hermanni. Dytiscinae – Dytiscus Cybister

( 24,0 – 39,0 )., Hydradephaga

(62 36 ).

[131, 194, 242], Hydradephaga

, , . ,

, . , ,

. , 5.07.2008 . .

Hydradephaga , 0,1 .

15-38 2, .

, ,

, ,

. , ,

,. 2005 .

D. dimidiatus ( 2-10 2) ( 0,15 )

, 0,3-0,5 .

, ,

116

. , ,

. ,

, , ,

. [260].

Hydradephaga .» . [3]

, . , . .

[2] -,

. [2]

[3].. [136, 137] .

– «...,

».Hydradephaga

, [71,203, 204, 234].

, , « », « » « -

». . [142] ,

, . Hydradephaga . . [131] H. angustatus, H. striola

Hydroporus umbrosus (Gyllenhal, 1808) . D. latissimus Haliplus confinis Stephens, 1829

2 [143, 203], 1 . ,

Hydradephaga 5 [234]. ,

« ». , , , Hydradephaga

117

[43, 71, 127, 203, 204, 234, 253], [1, 24, 73].

. . [134]

. .

Hydradephaga « », ,

( ). , –,

[71, 127, 131, 142, 213, 253].

. , Hydradephaga [71, 234].

, , .

( ) ,

, – [203, 234].

(Diptera), ,

[143, 201, 203, 234]. , Hydradephaga

« »., Brychius Haliplus

, Peltodytes,

[71, 127, 203, 204, 213, 253].,

Ilybius, Dytiscus semisulcatus O.F. Müller, 1776 [203, 234]. ,

[71, 127, 203, 213]. ,

[71, 127, 203, 234].

« ». , ,,

[4, 143, 203, 234]. ,

118

( Dytiscus,Cybister, Hydaticus), ,

[4, 131, 143]., Hydradephaga,

, [74]. ,

[71, 203, 234]. , Hydradephaga [71, 127, 202, 204, 234],

, , ,

. , ( )

[223, 243]., Hydradephaga

, , . Hydradephaga

[4, 13-15, 38, 71, 131, 133, 143,166, 176, 180, 181, 182, 184, 203, 204, 213, 234, 238, 239, 246-248, 253].

:1. , (

, , , );2. , ,

Haliplus , , Noterus);3. , (Peltodytes Brychius, - Noterus).

Hydradephaga ,

. , .

D. semisulcatus D. latissimus, D. marginalis – .

, . ,

(Heteroptera, Diptera, Ephemeroptera, Plecoptera,Odonata, Coleoptera, Trichoptera .), (Cladocera,Gammaridae, Asellus aquaticus (Linnaeus, 1758)), (Lymnaea,Physidae), , ,

. Dytiscus Thamnophis elegans (Baird et

119

Girard, 1853) 19 [188]. .

. Hydradephaga

. , . [13], D. marginalis 14 21-25 ,

1,4 . A. sulcatus, G. cinereus Agabus sturmii (Gyllenhal, 1808) 225 – 260 808 , 5-11 .

, Hydradephaga ,

, . ,

, .

, Hydradephaga (

). ,

, ., Hydradephaga

,.

, , , ,

. Hydradephaga . 15.

Hydradephaga

. – ,

, [74, 80]. Gyrinidae.

Hydradephaga (

. [3]) , .

, ., ,

, (

120

), – , .

. 15. Hydradephaga. () , . : O –

(Ovum), L1, L2, L3 – (Larvae) , , P – (Pupa), Im – (Imago).

. [137] . [3].

, ,

. Hydradephaga, , A. labiatus, A.

undulatus, Laccophilus, Rhantus Colymbetes, –

Dytiscinae. ,

( ) .

. ,

, .

, , ,

. Colymbetinae Dytiscinae

O

L1 L2 L3

P

Im

LARVAE

, ,,

,

121

, ( ) ( . 16).

1 2 3. 16. : 1 – Graphoderus cinereus, 2 – Cybister lateralimarginalis, 3 – Dytiscus persicus.

. (1,2) . (3).

, .

. ( Haliplidae),

, ( Dytiscidae,Hygrobiidae), ,

( A. undulatus, A. labiatus, Laccophilus, Hydaticus, D. dimidiatus, D. circumflexus),

: , Cladocera ( Acilius Graphoderus),

. [4], Rhantus Colymbetes), , ( C.

lateralimarginalis, . 17). Hydradephaga . ,

, , , ,

« » [127, 203, 204]. , , , ,

, .

122

. 17. Cybister lateralimarginalis – , . . .

–[74], ,

, ).

Hydradephaga ,

, .

Rhantus Colymbetes, Liopterus, Hygrotus, Ilybius, Agabus A. undulatus

A. labiatus), N. ceresyi, H. ovatus, H. cuspidatus, , Hydroporinae P. maculatus.

, .

, [127].

, . , (

123

) (

). N. ceresyi, H. ovatusHygrotus (H. confluens, H. enneagrammus)

, .

Hydradephaga , .

: (Cladocera) (Ostracoda)

( , Diptera), (Ilybius, Agabus) ( 150%

) , 10-12 , ,

(A. aquaticus), (Amphipoda), (Ephemeroptera), (Odonata), (Trichoptera),

(Diptera), (Corixidae). Hydradephaga

– , ,

, .

, – ( . ).

, ,

. H. hermanni, ,

.

, .

, , ( , Cladocera,

(Rotatoria)). . [71] . H. hermanni,

[204], , Tubifex.

Hydradephaga – ,

.

124

, : H. cuspidatus, N. ceresyi, H. ovatus, H.

impressopunctatus, H. parallelogrammus, H. versicolor, H. decoratus, H.inaequalis, Liopterus, Ilybius, Agabus, P. maculatus

Hydroporinae ( Hydroporus, Suphrodytes, Porhydrus,Graptodytes, Bidessus, Hydroglyphus, Herophydrus).

[27, 28, 100, 159, 160]: 0,1 ), (0,1-2,0 ) ( 2,0 ).

Hydradephaga .

– – [159].

Hydradephaga 2,0 ,

, .

Bidessus, Hydroglyphus,Graptodytes,

Haliplus, Peltodytes, Hygrotus, Hydroporus, Hydrovatus, Herophydrus,Porhydrus, Nebrioporus Suphrodytes. ,

Hyphydrus, Liopterus, Ilybius Agabus.

Hydradephaga

, (P.caesus), ( ).

, , , ,

, , . , ,

. Hydradephaga

( ) (). , ,

, – ( , , ), –

, (Cladocera, Ostracoda, Harpacticoida, Cyclopoida,

A. aquaticus). Hydradephaga

. (

125

. [137]) , Acilius,

, [203].

( « » [82]),

Hydradephaga,, ,

( . 3)., ,

Hydradephaga , , .

, Hydradephaga ,

. () ,

. , ( ) ,

.

, . ,

, ( )

. , Hydradephaga

.

126

6.,

Hydradephaga,

, . ( . 18).

. 18. Hydradephaga ().

Hydradephaga . , Hydradephaga

(Hydradephaga). [143, 166, 211, 234, 244],

, .,

. , ,

, .

(Nepa cinerea Linnaeus,1758), (Ranatra linearis (Linnaeus, 1758)), (Ilyocoriscimicoides (Linnaeus, 1758)), (Notonecta glauca Linnaeus, 1758 N. viridis Delcourt,1909) ,

. N. cinerea

HydradephagaPisces

Aves

Mammalia

Heteroptera,Larvae

Odonata,Trichoptera

Larvae Diptera,Ephemeroptera,

Plecoptera

Hymenoptera

Crustacea

Polychaeta,Oligochaeta, Hirudinea,Rotatoria

Algae

Fungi

Apicomplexa

Amphibia,Reptilia

PlantaeVasculares

Mollusca

Acari, Aranei

Nematoda

Nematomorpha

127

Hydroporus, Hygrotus, Agabus, Ilybius Rhantus 10,0, – P. caesus, (H. planus

A. labiatus). ,

, . , Nepidae – R.

linearis, , Hydradephaga

. , ( ) 32 118

8 1-3 , 12,2-150,8% ( . 26).

26 Hydradephaga Ranatra linearis

)N max,

. l, m, M, M/mp,%

H. zacharenkoi, imago 8 2,8 2,0 16,0 13,6N. crassicornis, imago 7 3,7 2,9 20,3 17,2H. inaequalis, imago 6 3,3 2,5 15,0 12,7H. impressopunctatus, imago 6 4,9 8,1 48,6 41,2P. lineatus, imago 7 3,0 2,2 15,4 13,1L. minutus, imago 5 4,4 3,0 15,0 12,7L. poecilus, imago 6 3,6 2,4 14,4 12,2A. labiatus, imago 3 6,2 35,7 107,1 90,8A. bipustulatus, imago 2 10,2 89,0 178,0 150,8R. suturalis, imago 2 11,1 76,7 153,4 130,0I. subaeneus, imago 1 11,5 112,0 112,0 94,9G. austriacus, imago 1 11,8 177,0 177,0 150,0

. N max – ,, l m – , ,

, M – , mp – ,.

I. cimicoidesA. bipustulatus, C. fuscus Ilybius. 1-3 7,0-11,0 3,6-17,0 .

, N. clavicornis L. minutus.

Hydradephaga .

N. glauca N. viridisN. crassicornis H.

impressopunctatus H. inaequalis. [211, 244]

128

40% N. glauca 10% 4-5 N. viridis.

Aeshna Anax imperator (Leach, 1815) 5 – 14

. A. imperatorH. impressopunctatus A. labiatus, , ,

, . . [143],

AeshnaAgabus sturmii (Gyllenhal, 1808) A. congener (Thunberg, 1794). ,

Hydradephaga .Hydroporus

Agabus Argyroneta aquatica(Clerck, 1757). . [143]

Dolomedes fimbriatus (Clerck, 1758).

Hydradephaga.

N. ceresyi, A. bipustulatus, A. labiatus, C. fuscus, R. suturalis, H.transversalis, D. dimidiatus, D. circumflexus, C. lateralimarginalis.

.D. dimidiatus C. lateralimarginalis

(H. seminiger,R. suturalis, A. labiatus) 1-4 . ,

Agabus, Ilybius, Rhantus C. fuscus (1-11 . ).Dytiscus,

, , .

[234]. ,

., Hydradephaga

. (R. linearis, N. glauca),

. Hydradephaga, , ,

(Carabidae) [212, 234].

Hydradephaga .

: (Coregonidae), (Thymallus), (Brachymystax lenok (Pallas, 1773)), (Perca fluviatilis (Linnaeus,

129

1758)), (Carassius), (Cyprinus carpio (Linnaeus, 1758)), (Rutilus rutilus (Linnaeus, 1758)), (Aspius aspius (Linnaeus, 1758)),

(Scardinius erythrophthalmus (Linnaeus, 1758)), (Bliccabjoerkna (Linnaeus, 1758)), (Leuciscus idus (Linnaeus, 1758)) [77, 106,138, 143, 169].

(Esox luceus Linnaeus, 1758), , (L. cephalus(Linnaeus, 1758)), , (Silurus glanis Linnaeus, 1758)

(Belone belone (Linnaeus, 1761)) , ,

., 2004-2008 . ,

Graphoderus C. lateralimarginalis ( . 27). 27

Hydradephaga, .

n n1 Hydradephaga

5.09.2004 4 2 C. lateralimarginalis (L3) – 2 ., Graphoderus sp. (Im) – 1 .12.09.2004 2 1 Graphoderus sp. (Im) – 1 .28.08.2005 2 1 C. lateralimarginalis: L2 – 4 .; L3 – 1 .24.06.2006 3 1 C. lateralimarginalis: L1 – 2 .; L2 – 1 .; L3 – 2 .

13.07.2008 5 3 C. lateralimarginalis: L2 – 3 .; L3 – 1 ., Graphoderus sp. (L3)– 1 .

. n – , n1 – , Hydradephaga, Im – , L1, L2, L3 – –

.

, C. lateralimarginalis, Graphoderus.

, , . ) C. lateralimarginalis

Hydradephaga. , (30-80 )

. Graphoderus ,

( Hydradephaga) [186].

( , ), 2004-2008 . .

Haliplus P.caesus.

D. dimidiatusC. lateralimarginalis.

130

, ,, 2004

.,

2001-2008 . , . ( )

15 Hydradephaga: P. caesus, H. fluviatilis, H.ruficollis, N. crassicornis, H. impressopunctatus, H. inaequalis, H. ovatus, A.labiatus, I. fuliginosus, I. quadriguttatus, R. suturalis, R. latitans, L. minutus, L.poecilus, Hydaticus sp. ,

Hydaticus, Graphoderus C. lateralimarginalis.

.

. 32 , 2003 . ,

R. suturalis.,

. ,

, .

, . Hydradephaga.

Rana, R. suturalis.

Hydradephaga Emys orbicularis (Linnaeus,1758). . ( ),

. [31, 105, 143, 166].

. . Hydradephaga (Ciconia), (Ardea

cinerea Linnaeus, 1758, Nycticorax nycticorax (Linnaeus, 1758), Isobrychusminutus Linnaeus, 1758, Botaurus stellaris (Linnaeus, 1758)), (Stercorarius parasiticus (Linnaeus, 1758)) (Anatidae), (Tringaglareola Linnaeus, 1758, T. erythropus (Pallas, 1764) Philomachus pugnax(Linnaeus, 1758)) [6, 86, 143, 146, 234]. ,

, . [29],

131

(Turdus merula Linnaeus, 1758).A. cinerea, Egretta alba

(Linnaeus, 1758) (Ciconia iconia (Linnaeus, 1758)) D. circumflexus, D. dimidiatus C. lateralimarginalis.

. ,

. Hydradephaga [143]

[234].

Hydradephaga. Hydradephaga ,

, , (Gregarinida: Bothriopsides sp., Ancyrophorauncinata Lèger, 1892), (Ciliophora, Suctoria), (Nematoda), (Nematomorpha: Gordius aquaticus Linnaeus, 1758),

(Trematoda: Pleurogenoides medians (Olson, 1876), Plagiorchis sp.), (Hydrachnida) (Hymenoptera) [31, 44, 99, 154,

155, 190-193, 240, 234, 245, 250, 262]. . [116]

.

. , . , . , .

. 2007 . , , ,

.

.Fungi. Hydradephaga

, .

, C. lateralimarginalis D. dimidiatus.

: H. planus, A. bipustulatus, C. fuscusH. seminiger.

, .

, . -

1 .

132

. Deuteromycetes

[=Fungi Imperfecti], C. lateralimarginalis, 9 2007 . ( . 19).

. 19. Deuteromycetes, C. lateralimarginalis.. .

.

C. lateralimarginalis 1%, 2007

30%. , ,

, ,, .

, 9-11 , . ,

6-8 . .

,

, . ,

, .Algae. , ,

Chlorophyta Bacillariophyta , ,

( . 3). ,

[218], , .

133

, 2007 D. dimidiatus. ,

, , , .

Ciliophora. ( . [44])

Hydradephaga. Dytiscus, Hydaticus Graphoderus ( . 20.) Vorticella,Carchaesium Zoothamnium ( . ).

. 20. , Graphoderuscinereus. .

, . ,

, ,

.Trematoda. ,

. Asimphilodora sp., C. lateralimarginalis. 8 C. lateralimarginalis,

28 2007 ,

134

. , Hydradephaga ,

.,

Hydradephaga , ,, , (

. [10]) .,

, . Hydradephaga

. ,

[13, 14, 31, 38, 203]. , Rhantus, Colymbetes, Agabus, Ilybius, Hydaticus, Acilius , .

[7, 45, 122-125, 143, 167, 168, 175, 183, 187, 203, 217, 227, 228, 234, 237].,

( . 21).

Hydradephaga:

. 21. Hydradephaga

, .

. . [41], Hydroporus sp.

135

, 46

. ,

.,

. , , , .

,

.

136

, 103 Hydradephaga, 30 5 : Hygrobiidae – 1, Haliplidae – 15,

Noteridae – 2, Dytiscidae – 75, Gyrinidae – 10 . 90 , 1 – A. concinnus , 12

. 11 Hydradephaga , 3 – H. zacharenkoi, H. musicus H.

pallidulus - , A.fuscipennis, A. didymus, R. exsoletus,

. Hydradephaga

, , , ,

,

. Hydradephaga ,

, .

(63 Hydradephaga) ( 80%) . ,

,

, .

Hydradephaga (H. fulvicollis, H. heydeni, H. decoratus, H. pubescens, H.palustris, H. memnonius, A. fuscipennis, I. ater, R. grapii, R. exsoletus, D.marginalis, D. circumcinctus, G. paykulli .)

, .

, .

Hydradephaga

« »

.

Hydradepahaga, .

, ,

137

« ».

. Hydradepahaga ,

. ,

.-

Hydradephaga , 0,57. ,

. ,, , , , ICS 0,45, 0,78.

Hydradephaga ( )

, ,

, . , ICS

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1. . . . / . ., . . , . . . – . : ,

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(Coleoptera, Hydradephaga) ( , ) / . . //

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166

1 1.1

, Hydradephaga

. º S, Im S, L I L NG

Haliplidae

1Peltodytes caesus(Duftschmid, 1805)

1 1 (?4) IV-XI 1-2 . 0,5-1

2 Haliplus obliquus(Fabricius, 1787) , 1 ? IV-VII ? ?

3 Haliplus lineatocollis(Marsham, 1802) 1 ?5 ?1-2

. 0,5-1

4 Haliplus fulvicollisErichson, 1837 1 ?1

(?4) III-IV ?III-VI ?1-2. 0,5-1

5 Haliplus furcatusSeidlitz, 1887

, 1 ?4III-V

(2007: IX)

? ?1,5-2. 0,5

6 Haliplus fluviatilisAubé, 1836 , 1 1(4) III-X III-

VIII0,5-1

. 1-2

7 Haliplus immaculatusGerhardt,1877 , 1 ?1 IV-XI III-VII 0,5-1

. 1

8 Haliplus heydeniWehcke, 1875

,, 1 ?4 III-X III-X 1-2 . 0,5-1

9 Haliplus ruficollis (DeGeer, 1774)

1 1(4) III-XIIII-

VIII,X-XI

0,5-2. 1-2

10 Haliplus sibiricusMotschulsky, 1860 1 ? III-X ? ? ?

167

1.11 2 3 4 5 6 7 8 9 10 11 12 13

11 Haliplus zacharenkoiGramma et Prisny,1973

,, 1 ?4

,2006 .:IV-XII

?, .IV-V

..

?2 . ?0,5

12 Haliplus flavicollisSturm, 1834 , , 1 ? III-X V-VI

(L3) ? ?

13 Haliplus fulvus(Fabricius, 1801) , 1 ? III-X V-VI

(L3) ? ?

14 Haliplus maculatusMotschulsky, 1860 , 1 ? III-X ? ? ?

15 Haliplus variegatusSturm, 1834 1 ? III-X V-VI

(L3) ? ?

Hygrobiidae

16 *Hygrobia hermanni(Fabricius 1775) ? 2 ?1 ? ? ?1 . ?1

Noteridae

17 Noterus crassicornis(O.F.Muller, 1776) 1 1 III-VI 1

. 1

18 Noterus clavicornis(De Geer, 1774) 1 1 III-VI 1

. 1

Dytiscidae Copelatinae

19Liopterushaemorrhoidalis(Fabricius, 1787)

, , 2 1 IV-VI 0,5. 1

Hydroporinae

20 Bidessus nasutusSharp, 1887

, 1 1 III-X IV-VI 0,5. 1

21 Bidessus unistriatus(Goeze, 1777) , 1 1 III-X IV-VI 0,5

. 1

168

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

22Hydroglyphusgeminus (Fabricius,1792)

1 1 III-IX 1-3. 2-4

23Hydrovatuscuspidatus (Kunze,1818)

, 1 1 IV-X V-VII 2-4. 1

24 Hyphydrus ovatus(Linnaeus, 1761) , 1 1 V-VII,

IX-X1-3

. 1-2

25 Herophydrus musicus(Klug, 1834) 1 1 VII, . ? ? ?1

26Hygrotusimpressopunctatus(Schaller, 1783)

1 1 III-IX 1-2. 2-4

27Hygrotusparallelogrammus(Ahrens, 1812)

, 1 1 IV-IX 1-2. 2-4

28*Hygrotuscorpulentus (Schaum,1864)

? ? 1 1 ? ? ?1

29 Hygrotus confluens(Fabricius, 1787)

, 1 1 III-XI IV-VII 1-2. 2-4

30 Hygrotus pallidulus(Aube, 1850) , 1 1 VII-VIII,

. ?VI-VII ? ?

31Hygrotusenneagrammus(Ahrens, 1833)

1 1 III-XI IV-IX 1-3. 1-3

32 *Hygrotus flaviventris(Motschulsky, 1860) 1 1 ? ? ?

33 Hygrotus decoratus(Gyllenhal, 1810) , 1 1 III-VIII III-V 2-4

. 1

169

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

34 Hygrotus inaequalis(Fabricius, 1777) 1 1 III-IX 1-2

. 2-4

35*Hygrotusquinquelineatus(Zetterstedt, 1828)

? ? 1 1 ? ? ? ?1

36 Hygrotus versicolor(Schaller, 1783) 1 1 VI-VII,

. ?IV-VI

?2-4. 1

37 Suphrodytes dorsalis(Fabricius, 1787) ? 1 1 IV-X ?V-

VII?2-4

. 1

38Hydroporusangustatus Sturm,1835

, 1 1 III-XI IV-V,IX-X

2-3. 1-2

39 Hydroporus planus(Fabricius, 1781)

1 525

.-5.

4-5

40Hydroporuspubescens (Gyllenhal,1808)

1 1 III, . ?III-VI ? 1

41Hydroporus discretusFairmaire et Brisout,1859

, 1 1 III-XI IV-VII 3-5. 1

42*Hydroporuserythrocephalus(Linnaeus, 1758)

1 1 ?III-VI ?2-4. 1

43Hydroporusmemnonius Nicolai,1822

, 1 5 III-VII,.

?3-4. ?

44 *Hydroporus striola(Gyllenhal, 1826) ? 1 1 ?III-VI ?2-4

. 1

45 Hydroporus palustris(Linnaeus, 1761) , 1 1 II-VII,

. ?III-VI ?2-4. 1

170

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

46 *Hydroporusobscurus Sturm, 1835 ? ? ? ? ? 1 1 ? ? ?1 . ?1

47 *Hydroporus tristis(Paykull, 1798) 1 1 ?III-VI ?2-4

. 1

48Graptodytesbilineatus (Sturm,1835)

, 1 1 IV-VI 3-5. 1

49*Graptodytesgranularis (Linnaeus,1767)

1 1 ? ? ? ?1

50 Nebrioporus ceresyi(Aubè, 1838)

1 525

.-5.

4

51 Porhydrus lineatus(Fabricius, 1775) , 1 1 III-IX IV-VII 1,5-4

. 3

Agabinae

52 Agabus biguttatus(Olivier, 1795) , 2 5 IV-X 2-6

. 1-3

53 Agabus bipustulatus(Linnaeus, 1767) 2 5 3-5

. 1-3

54 Agabus didymus(Olivier, 1795) ? ? 2 1 IV, . ? ? ?1

55 Agabus fuscipennis(Paykull, 1798) 2 2 IV-V III-IV 1 . 1

56 Agabus labiatus(Brahm, 1791) 2 1 IV-VI,

VII1,5-3

. 1-2

57 Agabus undulatus(Schrank, 1776) 2 1 IV-VI 2-3

. 1

58 *Agabus lineatusGebler, 1848 ( ) ? ? 2 ? III-VI ? ? ?

171

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

59 Agabus conspersus(Marsham, 1802) 2 1 III-VII 1,5-4

. 3

60 Agabus nebulosus(Forster, 1771) 2 1 III-VII 1,5-4

. 3

61 Agabus paludosus(Fabricius, 1801) , 2 1 III-VII,

. IV-VI 3-5. 1

62 Platambus maculatus(Linnaeus, 1758)

, 2 4 IV-VIII,. ?IX-V 1,5-2

. 0,5

63 Ilybius chalconatus(Panzer, 1796) 2 3 VI-VII,

. ?II-V 1,5-2. 0,5

64 Ilybius fenestratus(Fabricius, 1781)

,, 2 4 IV-X IX-V 1-2 . 0,5

65 Ilybius fuliginosus(Fabricius, 1792) 2 4 V-X VIII-V 1-2 . 0,5

66 Ilybius ater (De Geer,1774) 2 4 IV-X IX-IV 1-2 . 0,5

67 Ilybius quadriguttatus(Lacordaire, 1835) 2 4 V-X, .

III-X IX-V 1-2 . 0,5

68 Ilybius similisThomson, 1856 2 4 V-X, .

III-X IX-V 1-2 . 0,5

69 Ilybius subaeneusErichson, 1837 2 4 V-X IX-V 1-2 . 0,5

Laccophilinae

70 Laccophilus hyalinus(De Geer, 1774)

,,

, 1 1 III-VIII,

. V-VII 2-5. 1

71 Laccophilus minutus(Linnaeus, 1758) 1 5

25.-5.

4-5

172

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

72 Laccophilus poecilusKlug, 1834 1 1 IV-

VIII1,5-4

. 3

Colymbetinae

73 Rhantus grapii(Gyllenhal, 1808) , 2 1 III-X IV-VI 3-6

. 1

74Rhantus suturalis(W.S. MacLeay,1825)

2 1 III-X 1-3. 5

75 Rhantus frontalis(Marsham, 1802)

2 1 III-XIIV-VII,IX-X

1-4. 2

76 Rhantus bistriatus(Bergsträsser, 1778) , 2 1 III-X IV-VII 1,5-4

. 1

77 Rhantus latitansSharp, 1882 , 2 1 III-X III-VI,

VII1-4

. 2

78 Rhantus exsoletus(Forster, 1771) , 2 1 IV-V,

. ?IV-VII

5. ?1

79 Colymbetes fuscus(Linnaeus, 1758) 2 1 II-V,

IX-X4-5

. 2

80 Colymbetes striatus(Linnaeus, 1758) , 2 1 IV-VII,

. III-V 4-5. 1

Dytiscinae

81 Hydaticus seminiger(DeGeer, 1774) 2 1 III ( .

IV)-X IV-VII 3-5. 1

82 Hydaticustransversalis(Pontoppidan, 1763)

2 1 III ( .IV)-XI

IV-VII,VIII-IX

1-4. 2

83 Hydaticus grammicus(Germar, 1827) , ? 2 1 III ( .

IV)-X IV-VI 2-4. 1

173

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

84 Dytiscus dimidiatusBergsträsser, 1778 3 1 III-VI 3-5

. 1

85 Dytiscus marginalisLinnaeus, 1758 3 1 V, . IV-VI 3-5

. 1

86Dytiscuscircumcinctus Ahrens,1811

? 3 1 III-V,. IV-VI 3-5

. 1

87 Dytiscus circumflexusFabricius, 1801 , 3 1 III-V,

VII-XI IV-VI 3-5. 1

88Graphoderusaustriacus (Sturm,1834)

2 1 IV-XI V-VII,IX-X

1-3. 3

89 Graphoderus cinereus(Linnaeus, 1758) 2 1 IV-XI V-VII,

IX-X1-3

. 3

90 Graphoderus zonatus(Hoppe, 1795) , 2 1 IV-V,

. VI-VII 1,5-3. 1

91 Acilius canaliculatus(Nicolai, 1822) 2 1 IV-VI,

VII-IX3-4

. 2

92 Acilius sulcatus(Linnaeus, 1758) 2 1 IV-VI,

VII-IX3-4

. 2

93Cybisterlateralimarginalis (DeGeer, 1774)

3 1 III-XI V-IX 4-7. 1

Gyrinidae Gyrininae

94**Aulonogyrusconcinnus (Klug,1834)

? ? 1 ?1 ? ? ? ?

95 Gyrinus caspiusMénétriés, 1832 1 ?1 IV-X ? ? ?1

174

1.1.1 2 3 4 5 6 7 8 9 10 11 12 13

96 Gyrinus paykulliOchs, 1937

,, 1 ?1 III-X ? ? ?1

97 Gyrinus natatorLinnaeus, 1758 1 ?1 III-X V-VI 3-5

. 1

98 Gyrinus substriatusStephens, 1827 1 ?1 III-X V-VI 3-5

. 1

99 Gyrinus suffrianiScriba, 1855 , 1 ?1 IV-X ? ?3-4

. 1

100 *Gyrinus colymbusErichson, 1837 ? ? ? ? ? 1 ?1 ? ? ? ?1

101 Gyrinus distinctusAubé, 1864 1 ?1 III-X ? ?3-5

. 1

102 *Gyrinus marinusGyllenhal, 1808 ? ? ? ? ? 1 ?1 ? ? ? ?1

Orectochilinae

103 Orectochilus villosus(O.F.Muller, 1776) 1 ?1 VI-VII,

. ?VI-VIII ? 1

. * - , , ** - , .: . – ( – , – , –

, – , – , – ), º - ( – , –, – ), S – (Im – , L – , – , – -

, – , – ), – ( – , – , –, – ), ? – .

: – . , – . , I, L – ( – , – , ), – 1 ( . – , . – , . – ), NG – . . – , . –

, . – , . – , . – .

175

2 2.1

Hydradephaga

Peltodytes caesus (Duftschmidt, 1805) - - - -Haliplus lineatocollis (Marsham, 1802)Haliplus obliquus (Fabricius, 1787) - - - - - -Haliplus ruficollis (De Geer, 1774) - - - - - - -Haliplus fluviatilis Aubé, 1836 - - - - - -Haliplus heydeni Wehncke, 1875 - - - - - -Haliplus sibiricus Motschulsky, 1860 - - - - - - -Haliplus immaculatus Gerhardt,1877 - - - - - -Haliplus zacharenkoi Gramma et Prisny, 1973 - - - -Haliplus maculatus Motschulsky, 1860 - - - - - - -Haliplus variegatus Sturm, 1834 - - - - - -Noterus clavicornis (De Geer, 1774) -Noterus crassicornis (O.F.Muller, 1776) - - - - - - -Liopterus haemorrhoidalis (Fabricius, 1787) - - - - - -Hydroglyphus geminus (Fabricius, 1792)Hydrovatus cuspidatus (Kunze, 1818) -Hyphydrus ovatus (Linnaeus, 1761) - - - - - -Hygrotus inaequalis (Fabricius, 1777) - - - -Hygrotus impressopunctatus (Schaller, 1783) -Hygrotus parallelogrammus (Ahrens, 1812) - - - - - -Hygrotus confluens (Fabricius, 1787) - - - - -Hydroporus angustatus Sturm, 1835 - - - - - - -Hydroporus discretus Fairmaire et Brisout, 1859Hydroporus memnonius Nicolai, 1822Hydroporus palustris (Linnaeus, 1761) - - - - - - -Hydroporus planus (Fabricius, 1781)

176

2.1.1 2 3 4 5 6 7 8 9

Graptodytes bilineatus (Sturm, 1835) - - - - - -Porhydrus lineatus (Fabricius, 1775) - - -Laccophilus hyalinus (De Geer, 1774) - - - - - - -Laccophilus minutus (Linnaeus, 1758)Laccophilus poecilus Klug, 1834 -Agabus biguttatus (Olivier, 1795) - -Agabus bipustulatus (Linnaeus, 1767) -Agabus labiatus (Brahm, 1791) - - - - - - -Agabus undulatus (Schrank, 1776) - - - - -Agabus conspersus (Marsham, 1802) -Agabus nebulosus (Forster, 1771) -Agabus paludosus (Fabricius, 1801) - - - -Platambus maculatus (Linnaeus, 1758) - - - - -Ilybius chalconatus (Panzer, 1796) - - - - - - -Ilybius fenestratus (Fabricius, 1781) - - - - - - -Ilybius fuliginosus (Fabricius, 1792)Ilybius quadriguttatus (Lacordaire, 1835) - -Ilybius similis Thomson, 1856 - - - -Ilybius subaeneus Erichson, 1837 - - - -Rhantus bistriatus (Bergsträsser, 1778) - - - - - -Rhantus frontalis (Marsham, 1802)Rhantus suturalis (W.S. MacLeay, 1825)Colymbetes fuscus (Linnaeus, 1758) -Hydaticus seminiger (DeGeer, 1774) -Hydaticus transversalis (Pontoppidan, 1763) -Dytiscus circumflexus Fabricius, 1801 - - - -Dytiscus dimidiatus Bergsträsser, 1778 - - - - - -Dytiscus marginalis Linnaeus, 1758 - - - - - -Acilius canaliculatus (Nicolai, 1822) - - - - - -

177

2.1.1 2 3 4 5 6 7 8 9

Acilius sulcatus (Linnaeus, 1758) - - - - - -Graphoderus austriacus (Sturm, 1834) -Graphoderus cinereus (Linnaeus, 1758) -Cybister lateralimarginalis (DeGeer, 1774) - - -Gyrinus paykulli Ochs, 1937 - - - - - - -Gyrinus distinctus Aubé, 1864 - - - - -Gyrinus natator Linnaeus, 1758 - - - - - -Gyrinus substriatus Stephens, 1827 - - - - -

39 23 57 32 30 24 42 1563

. – , – . - – , – , – . 2.2

Hydradephaga . .

Peltodytes caesus (Duftschmidt, 1805) - -Haliplus lineatocollis (Marsham, 1802) - - - - -Haliplus obliquus (Fabricius, 1787) - - - - - - - - -Haliplus fulvicollis Erichson, 1837 - - - - - - - -Haliplus furcatus Seidlitz, 1887 - - -Haliplus fluviatilis Aubé, 1836 - - - - - - - - -Haliplus immaculatus Gerhardt,1877 - - - - - - -Haliplus heydeni Wehncke, 1875 - - - - - - -Haliplus ruficollis (De Geer, 1774) - -Haliplus zacharenkoi Gramma et Prisny, 1973Haliplus flavicollis Sturm, 1834 - - - - - - - -Haliplus fulvus (Fabricius, 1801) - - - - - -Haliplus maculatus Motschulsky, 1860 - - - -Haliplus variegatus Sturm, 1834 - - - -Noterus clavicornis (De Geer, 1774) -

178

2.2.1 2 3 4 5 6 7 8 9 10 11

Noterus crassicornis (O.F.Muller, 1776) - -Liopterus haemorrhoidalis (Fabricius, 1787) - -Bidessus nasutus Sharp, 1887 - -Bidessus unistriatus (Goeze, 1777) - - - - -Hydroglyphus geminus (Fabricius, 1792) -Hygrotus decoratus (Gyllenhal, 1810) - - - - - - -Hygrotus inaequalis (Fabricius, 1777) -Hygrotus impressopunctatus (Schaller, 1783) -Hygrotus parallelogrammus (Ahrens, 1812) - -Hygrotus confluens (Fabricius, 1787) - - - - -

Hygrotus enneagrammus (Ahrens, 1833) - - - ) - - - - ( ) -

Hydroporus angustatus Sturm, 1835 - -Hydroporus planus (Fabricius, 1781)Hydroporus discretus Fairmaire et Brisout, 1859 - - - - - - -Hydroporus memnonius Nicolai, 1822 - - - - - - - -Hydroporus palustris (Linnaeus, 1761) - - - - - - -Nebrioporus ceresyi (Aubè, 1838) - - - - - - - - ( ) -Graptodytes bilineatus (Sturm, 1835) - -Porhydrus lineatus (Fabricius, 1775) -Hydrovatus cuspidatus (Kunze, 1818) - -Hyphydrus ovatus (Linnaeus, 1761) - -Agabus biguttatus (Olivier, 1795) - - - - - - - -Agabus didymus (Olivier, 1795) - - - - - - - - -Agabus bipustulatus (Linnaeus, 1767) -Agabus fuscipennis (Paykull, 1798) - - - - - - - -Agabus conspersus (Marsham, 1802) -Agabus nebulosus (Forster, 1771) - -Agabus paludosus (Fabricius, 1801) - - - - - - - -Agabus labiatus (Brahm, 1791) -

179

2.2.1 2 3 4 5 6 7 8 9 10 11

Agabus undulatus (Schrank, 1776) - - - - - -Ilybius fenestratus (Fabricius, 1781) - - - - - - - -Ilybius fuliginosus (Fabricius, 1792) - -Ilybius ater (De Geer, 1774) - - - - - - - - -Ilybius quadriguttatus (Lacordaire, 1835) - -Ilybius similis Thomson, 1856 - -Ilybius subaeneus Erichson, 1837 -Laccophilus hyalinus (De Geer, 1774) - - - - - - - - -Laccophilus minutus (Linnaeus, 1758) -Laccophilus poecilus Klug, 1834 -Rhantus grapii (Gyllenhal, 1808) - - - - - - -Rhantus suturalis (W.S. MacLeay, 1825) -Rhantus frontalis (Marsham, 1802) -Rhantus bistriatus (Bergsträsser, 1778) - - -Rhantus latitans Sharp, 1882 - - - - - - - -Rhantus exsoletus (Forster, 1771) - - - - - - - -Colymbetes fuscus (Linnaeus, 1758) -Colymbetes striatus (Linnaeus, 1758) - - - - - - -Hydaticus seminiger (DeGeer, 1774) - -Hydaticus transversalis (Pontoppidan, 1763) - -Hydaticus grammicus (Germar, 1827) - - - - - - - -Graphoderus austriacus (Sturm, 1834) - -Graphoderus cinereus (Linnaeus, 1758) - -Graphoderus zonatus (Hoppe, 1795) - - - - - -Acilius canaliculatus (Nicolai, 1822) - -Acilius sulcatus (Linnaeus, 1758) - -Dytiscus dimidiatus Bergsträsser, 1778 - -Dytiscus circumflexus Fabricius, 1801 - -Cybister lateralimarginalis (DeGeer, 1774) -Gyrinus caspius Ménétriés, 1832 - - - - - - -

180

2.2.1 2 3 4 5 6 7 8 9 10 11

Gyrinus natator Linnaeus, 1758 - - - - -Gyrinus substriatus Stephens, 1827 - - - - -Gyrinus suffriani Scriba, 1855 - - -Gyrinus distinctus Aubé, 1864 - -

59 64 54 48 6 49 5- 49 58 2478

. – , – , - - . : – , – , – , – , – , –

. 2.3

Hydradephaga . .

Peltodytes caesus (Duftschmid, 1805) -Haliplus lineatocollis (Marsham, 1802) - - - - -Haliplus heydeni Wehncke, 1875 -Haliplus ruficollis (De Geer, 1774)Haliplus fluviatilis Aubé, 1836 - - -Haliplus immaculatus Gerhardt,1877 - - - -Haliplus sibiricus Motschulsky, 1860 - - - - -Noterus clavicornis (De Geer, 1774)Noterus crassicornis (O.F. Müller, 1776) - -Liopterus haemorrhoidalis (Fabricius, 1787) - - -Bidessus unistriatus (Goeze, 1777) - - - - -Hydroglyphus geminus (Fabricius, 1792)Hygrotus inaequalis (Fabricius, 1777)Hygrotus versicolor (Schaller, 1783) - - - - -Hygrotus impressopunctatus (Schaller, 1783)Hygrotus confluens (Fabricius, 1787) - - - - -Hydroporus angustatus Sturm, 1835 - - - -

181

2.3.1 2 3 4 5 6 7

Hydroporus planus (Fabricius, 1781) -Hydroporus discretus Fairmaire et Brisout, 1859 - - - -Hydroporus memnonius Nicolai, 1822 - - -Hydroporus palustris (Linnaeus, 1761) - - - -Graptodytes bilineatus (Sturm, 1835) - -Porhydrus lineatus (Fabricius, 1775) - - - -Platambus maculatus (Linnaeus, 1758) - - - - -Agabus paludosus (Fabricius, 1801) - - - - -Agabus bipustulatus (Linnaeus, 1767) - - -Agabus labiatus (Brahm, 1791) - -Agabus undulatus (Schrank, 1776) - -Ilybius chalconatus (Panzer, 1796) - - - - -Ilybius fenestratus (Fabricius, 1781) - - - -Ilybius fuliginosus (Fabricius, 1792) - - -Ilybius ater (De Geer, 1774) - - - - -Ilybius quadriguttatus (Lacordaire, 1835) - -Ilybius similis Thomson, 1856 - - - -Ilybius subaeneus Erichson, 1837 - - - -Laccophilus hyalinus (De Geer, 1774) - - -Laccophilus minutus (Linnaeus, 1758)Laccophilus poecilus Klug, 1834Rhantus suturalis (W.S. MacLeay, 1825)Rhantus frontalis (Marsham, 1802) -Colymbetes fuscus (Linnaeus, 1758) -Colymbetes striatus (Linnaeus, 1758) - - - - -Hydaticus seminiger (DeGeer, 1774)Hydaticus transversalis (Pontoppidan, 1763)Dytiscus dimidiatus Bergsträsser, 1778 - - -Dytiscus marginalis Linnaeus, 1758 - - - - -Graphoderus austriacus (Sturm, 1834) - - -

182

2.31 2 3 4 5 6 7

Graphoderus cinereus (Linnaeus, 1758) -Acilius canaliculatus (Nicolai, 1822) - - - -Acilius sulcatus (Linnaeus, 1758) - - - -Cybister lateralimarginalis (DeGeer, 1774) - -Hydrovatus cuspidatus (Kunze, 1818) - - - - -Hyphydrus ovatus (Linnaeus, 1761) - - -Gyrinus natator Linnaeus, 1758 - - - - -Gyrinus substriatus Stephens, 1827 - - -Orectochilus villosus (O.F. Müller, 1776) - - - - -

31 32 29 18 37 3156

. – , – , - - . : – , – , – .

2.4 Hydradephaga

Peltodytes caesus(Duftschmid, 1805) -

Haliplus lineatocollis(Marsham, 1802) - - - - - - - - - - - - -

Haliplus obliquus(Fabricius, 1787) - - - - - - - - - - - - - - -

Haliplus fulvicollisErichson, 1837 - - - - - - - - - - - - - -

Haliplus furcatus Seidlitz,1887 - - - - - - - - - - - -

Haliplus ruficollis (DeGeer, 1774) - -

183

2.4.1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Haliplus heydeni Wehncke,1875 - - - - - - - - - - - - -

Haliplus fluviatilis Aubé,1836 - - - - - -

Haliplus immaculatusGerhardt,1877 - - - - - - - - -

Haliplus zacharenkoiGramma et Prisny, 1973 - - - - - - - - - - - - - -

Haliplus flavicollis Sturm,1834 - - - - - - - - - - - -

Haliplus fulvus (Fabricius,1801) - - - - - - - - - - - -

Haliplus variegatus Sturm,1834 - - - - - - - - - - - - -

Haliplus maculatusMotschulsky, 1860 - - - - - - - - -

Noterus clavicornis (DeGeer, 1774)Noterus crassicornis(O.F.Muller, 1776) -

Liopterus haemorrhoidalis(Fabricius, 1787) - - - - - - - - -

Bidessus nasutus Sharp,1887 - - - - - - - - -

Bidessus unistriatus(Goeze, 1777) - - - - - - - -

Hydroglyphus geminus(Fabricius, 1792) -

Herophydrus musicus(Klug, 1834) - - - - - - - - - - - - - - -

184

2.4.1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Hygrotus decoratus(Gyllenhal, 1810) - - - - - - - - - - - - -

Hygrotus inaequalis(Fabricius, 1777) -

Hygrotus versicolor(Schaller, 1783)

( , ) - - - - - - - - - -

Hygrotus quinquelineatus(Zetterstedt, 1828)

( , ) - - - - - - - - - -

Hygrotusimpressopunctatus(Schaller, 1783)

- -

Hygrotusparallelogrammus (Ahrens,1812)

- - - - - - - - - - - - - -

Hygrotus confluens(Fabricius, 1787) - - - - - - - - - - - - - - -

Hygrotus pallidulus (Aube,1850) - - - - - - - - - - - - - - -

Hydroporus angustatusSturm, 1835 - - - - - - - - -

Hydroporus palustris(Linnaeus, 1761) - - - - - - - - - - - - - - -

Hydroporus planus(Fabricius, 1781) - - - -

Hydroporus pubescens(Gyllenhal, 1808) - - - - - - - - - - - - - -

Hydroporus obscurusSturm, 1835 - - - - - - - - - -

Hydroporus tristis(Paykull, 1798) - - - - - - - - - - - - - - -

185

2.4.1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Graptodytes bilineatus(Sturm, 1835) - - - - - - - -

Porhydrus lineatus(Fabricius, 1775) - - - - - - - - -

Hyphydrus ovatus(Linnaeus, 1761) - - - - - - -

Hydrovatus cuspidatus(Kunze, 1818) - - - - - - - - - -

Laccophilus hyalinus (DeGeer, 1774) - - - - - - - - - -

Laccophilus minutus(Linnaeus, 1758)Laccophilus poecilus Klug,1834Agabus bipustulatus(Linnaeus, 1767) - - - - - - - - - - - - -

Agabus labiatus (Brahm,1791) - - - - - - - - - -

Agabus undulatus(Schrank, 1776) - - - - - - - - - -

Platambus maculatus(Linnaeus, 1758) - - - - - - - - - - - - - -

Ilybius fenestratus(Fabricius, 1781) - - - - - - - - -

Ilybius fuliginosus(Fabricius, 1792) - - - - - - - - - - - - - - -

Ilybius quadriguttatus(Lacordaire, 1835) - - - - - - - - - -

Ilybius similis Thomson,1856 - - - - - - - - - - - -

186

2.41 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Ilybius subaeneus Erichson,1837 - - - - - - - - - - - - -

Rhantus grapii (Gyllenhal,1808) - - - - - - - - - - - -

Rhantus suturalis (W.S.MacLeay, 1825)Rhantus frontalis(Marsham, 1802) - - - - - - - -

Rhantus bistriatus(Bergsträsser, 1778) - - - - - - - - - - - - - - -

Rhantus latitans Sharp,1882 - - - - - - - - - -

Colymbetes fuscus(Linnaeus, 1758) - - - -

Hydaticus seminiger(DeGeer, 1774) - - - - - - -

Hydaticus transversalis(Pontoppidan, 1763) - -

Hydaticus grammicus(Germar, 1827) - - - - - ( -250 ) - - - - -

Graphoderus austriacus(Sturm, 1834) - - - - - -

Graphoderus cinereus(Linnaeus, 1758) - - - - - -

Acilius canaliculatus(Nicolai, 1822) - - - - - - - - - -

Acilius sulcatus (Linnaeus,1758) - - - - - - - - - - -

Dytiscus circumcinctusAhrens, 1811 - - - - - - - - - - - - - -

187

2.4.1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17

Dytiscus circumflexusFabricius, 1801 - - - - - - - - - - - - - - -

Dytiscus dimidiatusBergsträsser, 1778 - - - - - -

Cybister lateralimarginalis(De Geer, 1774) - -

Gyrinus caspius Ménétriés,1832 - - - - - - - - -

Gyrinus paykulli Ochs,1937 - - - - - - - - - - - - - -

Gyrinus natator Linnaeus,1758 - - - - - - - - - -

Gyrinus substriatusStephens, 1827 - - - - - - - - - - -

Gyrinus suffriani Scriba,1855 - - - - - - - - - -

Gyrinus distinctus Aubé,1864 - - - - - - - - -

Gyrinus marinus Gyllenhal,1808

( , ) - - - - - - - - -

17 29 54 16 16 22 34 49 51 58 19 19 18 27 17 3075

. – , – , – . : – , –, – . : – . : – . - - , – , –

, – : – [132], – [82, 83], – 250 – - 250 .

188

2.5 Hydradephaga

Peltodytes caesus (Duftschmid,1805) - - - - - - - - - -

Haliplus obliquus (Fabricius,1787) - - - - - - - - - - - -

Haliplus lineatocollis (Marsham,1802) - - - - - - - - - - -

Haliplus furcatus Seidlitz, 1887 - - - - - - - - - - - -Haliplus ruficollis (De Geer, 1774) - - - - - - - - - - -Haliplus zacharenkoi Gramma etPrisny, 1973 - - - - - - - - - - -

Haliplus maculatus Motschulsky,1860 - - - - - - - - - - - -

Noterus clavicornis (De Geer,1774) - - - - - - - -

Noterus crassicornis (O.F. Müller,1776) - - - - - - - - - - -

Liopterus haemorrhoidalis(Fabricius, 1787) - - - - - - - - - - - -

Bidessus nasutus Sharp, 1887 - - - - - - - - - - - -Hydroglyphus geminus (Fabricius,1792) - - -

Hygrotus inaequalis (Fabricius,1777) - - - - - - - - -

Hygrotus impressopunctatus(Schaller, 1783) - - - - - - -

Hygrotus confluens (Fabricius,1787) - - - - - - - - - - - -

Hygrotus enneagrammus (Ahrens,1833) - - - - ) - - - - - - -

189

2.5.1 2 3 4 5 6 7 8 9 10 11 12 13 14

Hydroporus discretus Fairmaire etBrisout, 1859 - - - - - - - - - - -

Hydroporus planus (Fabricius,1781) - - - - - - -

Graptodytes bilineatus (Sturm,1835) - - - - - - - - - - - -

Hydrovatus cuspidatus (Kunze,1818) - - - - - - - - - - -

Hyphydrus ovatus (Linnaeus,1761) - - - - - - - - - - -

Agabus biguttatus (Olivier, 1795) - - - - - - - - - - - -Agabus bipustulatus (Linnaeus,1767) - - - - - - - - - -

Agabus conspersus (Marsham,1802) - - - - - - - - - -

Agabus nebulosus (Forster, 1771) - - - - - - - - - - - -Agabus undulatus (Schrank, 1776) - - - - - - - - - - - -Ilybius fuliginosus (Fabricius,1792) - - - - - - - - - - -

Ilybius quadriguttatus (Lacordaire,1835) - - - - - - - - - - -

Ilybius subaeneus Erichson, 1837 - - - - - - - - - - - -Laccophilus minutus (Linnaeus,1758) - - - - - - -

Laccophilus poecilus Klug, 1834 - - - - - - -Rhantus suturalis (W.S. MacLeay,1825) - - - -

Rhantus frontalis (Marsham,1802) - - - - - - - - - - -

Colymbetes fuscus (Linnaeus,1758) - - - - - - - - -

190

2.5.1 2 3 4 5 6 7 8 9 10 11 12 13 14

Hydaticus seminiger (De Geer,1774) - - - - - - - - - - - -

Hydaticus transversalis(Pontoppidan, 1763) - - - - - - - -

Graphoderus austriacus (Sturm,1834) - - - - - - -

Graphoderus cinereus (Linnaeus,1758) - - - - - - -

Acilius canaliculatus (Nicolai,1822) - - - - - - - - - - -

Acilius sulcatus (Linnaeus, 1758) - - - - - - - - - - - -Dytiscus dimidiatus Bergsträsser,1778 - - - - - - - - - -

Dytiscus circumflexus Fabricius,1801 - - - - - - - - - - - -

Cybister lateralimarginalis (DeGeer, 1774) - - - -

Gyrinus caspius Ménétriés, 1832 - - - - - - - - - -Gyrinus substriatus Stephens,1827 - - - - - - - - - - -

Gyrinus distinctus Aubé, 1864 - - - - - - - - -1 22 11 24 5 6 3 25 3 23 2 3 13

. : – ( ), – , – , – , – , – , – , –

( ), – , – . - – , – , –, – .

191

2.6 Hydradephaga

Peltodytes caesus (Duftschmid, 1805) , , -Haliplus fulvicollis Erichson, 1837 - , - - -Haliplus furcatus Seidlitz, 1887 , - -Haliplus ruficollis (De Geer, 1774) , , - , -Haliplus zacharenkoi Gramma et Prisny, 1973 - - -Haliplus variegatus Sturm, 1834 - - -Noterus clavicornis (De Geer, 1774) , -Noterus crassicornis (O.F.Muller, 1776) , , -Liopterus haemorrhoidalis (Fabricius, 1787) - -Bidessus nasutus Sharp, 1887 , -Bidessus unistriatus (Goeze, 1777) , -Hydroglyphus geminus (Fabricius, 1792) -Hyphydrus ovatus (Linnaeus, 1761) - -Hydrovatus cuspidatus (Kunze, 1818) - -*Nebrioporus ceresyi (Aubè, 1838) - -Graptodytes bilineatus (Sturm, 1835) , , , - -Porhydrus lineatus (Fabricius, 1775) , , - - -Suphrodytes dorsalis (Fabricius, 1787) - -Hydroporus angustatus Sturm, 1835 - - -Hydroporus erythrocephalus (Linnaeus, 1758) - , - - -Hydroporus palustris (Linnaeus, 1761) - , - - -Hydroporus planus (Fabricius, 1781) - -Hydroporus pubescens (Gyllenhal, 1808) , , - - -Hydroporus striola (Gyllenhal, 1826) , , - - -Hydroporus tristis (Paykull, 1798) - , - - -Hygrotus decoratus (Gyllenhal, 1810) - - -Hygrotus inaequalis (Fabricius, 1777) , -

192

2.6.1 2 3 4 5 6

Hygrotus impressopunctatus (Schaller, 1783) , -Hygrotus parallelogrammus (Ahrens, 1812) , Hygrotus pallidulus (Aube, 1850) - - , -Hygrotus confluens (Fabricius, 1787) , , , , -Hygrotus corpulentus (Schaum, 1864) , - , , -Hygrotus enneagrammus (Ahrens, 1833) , -Hygrotus flaviventris (Motschulsky, 1860) - - - , -Laccophilus hyalinus (De Geer, 1774) - , - - -Laccophilus minutus (Linnaeus, 1758) -Laccophilus poecilus Klug, 1834 , -Agabus bipustulatus (Linnaeus, 1767) - - -Agabus conspersus (Marsham, 1802) - - - -Agabus lineatus Gebler, 1848 , - - - -Agabus labiatus (Brahm, 1791) , -Agabus undulatus (Schrank, 1776) - , - - -Ilybius quadriguttatus (Lacordaire, 1835) , , , - -Ilybius subaeneus Erichson, 1837 , , - - -Rhantus bistriatus (Bergsträsser, 1778) - , - - -Rhantus frontalis (Marsham, 1802) - -Rhantus grapii (Gyllenhal, 1808) - , - - -Rhantus suturalis (W.S. MacLeay, 1825) - -Colymbetes fuscus (Linnaeus, 1758) , - -Hydaticus grammicus (Germar, 1827) - - , - -Hydaticus seminiger (DeGeer, 1774) - -Hydaticus transversalis (Pontoppidan, 1763) - -Dytiscus dimidiatus Bergsträsser, 1778 , , - -Graphoderus austriacus (Sturm, 1834) , , , - -Graphoderus cinereus (Linnaeus, 1758) , , - - -Graphoderus zonatus (Hoppe, 1795) , , - - -Acilius canaliculatus (Nicolai, 1822) - , - - -

193

2.6.1 2 3 4 5 6

Acilius sulcatus (Linnaeus, 1758) , - - - -Cybister lateralimarginalis (De Geer, 1774) , - , , -Gyrinus caspius Ménétriés, 1832 - -*Gyrinus natator Linnaeus, 1758 - - - -

47 46 39 21 461

. * - , , - - , – , – , – [37].

194

3

3.1 Hydradephaga .

( 2000-2002 .)

1 2 3 4 5

Gyrinus natator - + + - + - + +G. substriatus - + + - + - + +Peltodytes caesus + + + + + + + +Haliplus lineatocollis + - - + + - + -H. ruficollis + + - + + + - -H. zacharenkoi + - - + - + - -H. fulvicollis - - - - + - + -H. furcatus + + - + + + + -H. fluviatilis + + - + + + - -H. fulvus + + - - + + + -H. variegatus + - - - - + + -Noterus crassicornis + + - + + + + +N. clavicornis + + - + + + + +Hydroporus angustatus + + - + + + - +H. palustris - - - - + - - -H. planus - + + - + - + +Graptodytes bilineatus + + - + + + + -Porhydrus lineatus - - - - + - + -Hygrotus impressopunctatus + + + + + + + +H. parallelogrammus + + - - + + - +H. confluens - + - - - - - +H. inaequalis + + + + + + + +H. decoratus + + - + + + + -H. geminus + + + + + + + +Bidessus nasutus + + - + + - - +Hydrovatus cuspidatus - + - - - - + -Hyphydrus ovatus + + + + + + + -Laccophilus poecilus + + + + + + + +L. minutus + + + + + + + +Liopterus haemorrhoidalis - + + - + + + -Agabus nebulosus - - - + + - + -A. conspersus - - - + + - + -A. bipustulatus - + - - + - + -A. undulatus + + - + + + + -A. labiatus + + - + + + + -Ilybius fenestratus + - - - - + - -I. fuliginosus - + - - - - + -I. subaeneus - + - - - - + -I. obscurus - + - - + + + -I. similis - + - - + + + -

195

3.1.1 2 3 4 5 6 7 8 9

Rhantus grapii - + - + + + + -R. suturalis - + - + + + + -R. frontalis + + - + + + + -R. bistriatus - - - - + - + -R. latitans - - - - + - + -Colymbetes fuscus + + - + + + + -Hydaticus transversalis + + - + + + + -H. seminiger + + - + + + + -Dytiscus dimidiatus + + - + + + + -D. circumflexus - + - + + + + -Acilius sulcatus + + - + + + + -A. canaliculatus + + - + + + + -Graphoderus zonatus - - - - + - + -G. cinereus + + - + + + + -G. austriacus + + - + + + + -Cybister lateralimarginalis + + - + + + + -

34 44 11 35 49 38 47 15. : 1 – (Ceratophyllum spp.), 2

– (Phragmites spp.), 3 – , 4 – (Juncusspp.), (Carex spp.) , 5 – , –

, – .

196

Acari – 126Acilius – 39,108,114,115,121,125,134,173,176,177,179,182,186,190,192,193

- canaliculatus (Nicolai, 1822) - 66,75,76,86,87,93,107- sulcatus (Linnaeus, 1758) – 48,59,66,76,81,84,86,90,93,98,107,119

Adephaga – 2,10,39,85,90Aeshna – 128Agabinae – 114,115Agabus – 39,40,48,52,119,122-124,127,128,134

- adpressus Aubé, 1837 – 52- amoenus Solsky, 1874 – 40- biguttatus (Olivier, 1795) – 51,53,63,77,110,11,138,10,176- bipustulatus (Linnaeus, 1767) – 51,56,58,63,77,84,90,99,100, 110, 111, 127, 128,131,170,176,178,181,185,189,192,194- congener (Thunberg, 1794) – 128- conspersus (Marsham, 1802) – 49,63,64,67,94,101,102,109,171,176,178,189, 192,194- didymus (Olivier, 1795) – 40,54,107,136,170,178- dilatatus (Brullè, 1832) – 40- fuscipennis (Paykull, 1798) – 40,52,55,87,109,112,136,170,178- guttatus (Paykull, 1798) – 40- labiatus (Brahm, 1791) – 54,66,84,85,89,90,93,94,98,107,108,120,121,122,

127,128,130,170,176,178,181,185,192,194- lineatus Gebler, 1848 – 40,44,66,67,170,176,178,192- nebulosus (Forster, 1771) – 49,109,171,176,178,189,194- paludosus (Fabricius, 1801) – 40,51,53,54,56,75,76,77,107,171,176,178,181- sturmii (Gyllenhal, 1808) – 40,52,119,128- undulatus (Schrank, 1776) – 49,54,56,59,66,90,92,93,107,120-122,170,176,

179,181,185,189,192- zimmermanni Scholz, 1920 – 40

Algae – 126,132Amphibia – 126Amphipoda – 123Amphizoidae – 11Anatidae – 130Anax imperator (Leach, 1815) – 128Ancyrophora uncinata Lèger, 1892 – 131Apicomplexa – 126Aranei – 126Ardea cinerea Linnaeus, 1758 – 130Argyroneta aquatica (Clerck, 1757) – 128Asellus aquaticus (Linnaeus, 1758) – 118Asimphilodora sp. – 133,141Aspidytidae – 11

197

Aspius aspius (Linnaeus, 1758) – 129Aulonogyrus – 39,173

- concinnus (Klug, 1834) – 38,136,173Aves – 126Bacillariophyta – 49,132,141Belone belone (Linnaeus, 1761) – 129Bidessus – 38,39,40,124,167,178,180,183,188,191,194

- alienus Zimmermann, 1919 – 40- delicatulus (Schaum, 1844) – 40- minutissimus (Germar, 1824) – 40- nasutus Sharp, 1887 – 60,61,67,76,85,86,88,107,167,178,183,188,191,194- pumilus Aubè, 1836 – 38- unistriatus (Goeze, 1777) – 55,66,76,86,107,167,178,180,183,191

Blicca bjoerkna (Linnaeus, 1758) – 129Botaurus stellaris (Linnaeus, 1758) – 130Bothriopsides sp. – 131Brachymystax lenok (Pallas, 1773) – 128Branchiapoda – 10Brychius elevatus (Panzer, 1794) – 75,117,118Carabidae – 128Carassius – 129Carchaesium – 133,141Carex spp. – 48,83,185Ceratophyllum spp. – 48,104,195Charophyta – 48,60Chlorophyta – 132,141Ciconia – 130,131

- iconia (Linnaeus, 1758) – 131Ciliophora – 131,133Cladocera – 108,118,121,123,129Coleoptera – 10,118Colymbetes – 39,52,121,122,134,172,176,179,181,186,188,192,195

- fuscus (Linnaeus, 1758) – 51,66,77,84,87-90,93,99,100,107-109,113,127,128, 131,140,172,176,179,181,186,189,192,195- paykulli Erichson, 1837 – 52- striatus (Linnaeus, 1758) – 54,55,57,107,172,179,181

Colymbetinae – 114,115,120,172Copelatinae – 114,115,167Coregonidae – 128Corixidae – 123Crustacea – 126Cybister – 39,97,114,115,118,121,122,173,177,179,182,187,190,193,195

- lateralimarginalis (De Geer, 1774) – 54,59,61,63,65,67,84,87-90,93-97,99, 100,107,121,122,128-133,173,177,179,182,187,190,193,195

198

Cyclopoida – 124Cyprinus carpio (Linnaeus, 1758) – 129Deronectes latus (Stephens, 1829) – 75Deuteromycetes [=Fungi Imperfecti] – 132,141Diptera – 108,117,118,123Dolomedes fimbriatus (Clerck, 1758) – 128Dytiscidae – 11,39,41,48,51,53,56,58,64,66,84,86,88-90,93-96,100,107,112-114,121, 136,140Dytiscinae – 114,115,120,172Dytiscus – 39,40,76,84,107,112,114,115,117,118,121,128,133,173,176,179,181,187, 190,192,195

- circumcinctus Ahrens, 1811 – 58,59,62,92,107,136,173,186- circumflexus Fabricius, 1801 – 48,53,54,57,77,84,107,121,128,131,173, 176,179,187,190,195- dimidiatus Bergsträsser, 1778 – 48,54,66,84,85,89,90,93,98,107,115,121, 128,129,131,133,173,176,179,181,187,190,192,195- latissimus Linnaeus, 1758 – 76,116,118- marginalis Linnaeus, 1758 – 51,52,57,107,118,119,136,173,176,181- persicus Wehncke, 1876 – 40,121- semisulcatus O.F. Müller, 1776 – 117,118

Egretta alba (Linnaeus, 1758) – 131Elodea – 104Emys orbicularis (Linnaeus, 1758) – 130Ephemeroptera – 118,123,126Eretes griseus (Fabricius, 1781) – 40Esox luceus Linnaeus, 1758 – 129Fungi – 126,131,132,141Gammaridae – 118Gordius aquaticus Linnaeus, 1758 – 131Graphoderus – 39,77,85,109,114,115,121,129,130,133,173,177,182,186,190,192

- austriacus (Sturm, 1834) – 66,90,105,127,173,177,179,181,186,190,192,195- cinereus (Linnaeus, 1758) – 48,49,63,85,86,88,90,98,109,119,121,133,173, 177,179,182,186,190,192,195- zonatus (Hoppe, 1795) – 54,67,107,173,179,192,195

Graptodytes – 39,124,170,176,178,181,185,189,191,194- bilineatus (Sturm, 1835) – 75,84,85,86,87,88,99,107,170,176,178,181,185, 189,191,194- granularis (Linnaeus, 1767) – 40,75,170

Gregarinida –131Gyrinidae – 11,39,41,47,51,53,56,58,64,66,86,94,111,112,113,114,115,119,136, 140,173Gyrininae – 173Gyrinus – 39,173,174,177,178,180,182,187,190,193,194

- caspius Ménétriés, 1832 – 42,58,60,61,63,64,67,77,95,173,179,187,190,193

199

- colymbus Erichson, 1837 – 40,174- distinctus Aubé, 1864 – 47,58,59,61,63,77,86,95,174,177,180,187,190- marinus Gyllenhal, 1808 – 38,40,52,59,61,174,187- natator Linnaeus, 1758 – 54,56,58,66,77,95,174,177,180,182,187,193,194- paykulli Ochs, 1937 – 52,58,59,76,77,94,136,174,177,187- substriatus Stephens, 1827 – 54,58,77,95,174,177,180,182,187,190,194- suffriani Scriba, 1855 – 47,58,59,61,77,86,95,174,180,187

Haliplidae – 11,39,41,51,53,56,58,64,66,84-86,88-90,93-96,100,104,112-115,121, 136,139,166Haliplus – 39,40,106,117,118,124,129,166,167,175,177,180,182,183,188,191

- confinis Stephens, 1829 – 116- dalmatinus J. Müller, 1990 – 40- flavicollis Sturm, 1834 – 52,54,58,59,60-62,76,105,167,177,183- fluviatilis Aubé, 1836 – 53,58,62,77,93-96,98,104,106,130,166,175,177, 180,183,194- fulvicollis Erichson, 1837 – 49,54,58,59,66,67,76,84,92,104,136,166,177,182, 191,194- fulvus (Fabricius, 1801) – 54,55,76,105,167,177,183,194- furcatus Seidlitz, 1887 – 58,66,76,85,93,104,106,166,177,182,188,191,194- heydeni Wehcke, 1875 – 52,54,57,58,59,76,93,105,106,136,166,175,177, 180,183- immaculatus Gerhardt,1877 – 40,54,58,62,76,85,93,105,166,175,177,180,183- lineatocollis (Marsham, 1802) – 51,56,57,63,65,76,77,99,104,106,114,166, 175,177,180,182,188,194- lineolatus Mannerheim, 1844 – 40,52- maculatus Motschulsky, 1860 – 40,48,49,60,61,76,87,105,167,175,177, 183,188- mucronatus Stephens, 1828 – 39,40- obliquus (Fabricius, 1787) – 48,52,54,60,61,65,76,85,105,166,175,177, 182,188- ruficollis (De Geer, 1774) – 38,75,76,93,95,104-106,130,166,175,177,180, 182,188,191,194- sibiricus Motschulsky, 1860 – 40,51,52,56,105,166,175,180- variegatus Sturm, 1834 – 48,49,58,60,61,67,85,105,167,175,177,183,191,194- zacharenkoi Gramma et Prisny, 1973 – 40,53,58,61,63,67,76,84-86,88,94, 105,106,127,136,167,175,177,183

Harpacticoida – 124Herophydrus – 39,124,168,183

- musicus (Klug, 1834) – 40,59,61,136,168,183Heteroptera – 76,118,126Hirudinea – 126Hydaticus – 39,40,77,85,86,92,95,107,114,115,118,121,130,133,134,172,176,179, 181,186,190,192,195

- continentalis J. Balfour-Browne, 1944 – 40

200

- grammicus (Germar, 1827) – 49,54,60,61,66,67,77,87,107,172,179,186,192- seminiger (DeGeer, 1774) – 45,66,89,92,107,128,131,172,176,179,181,186, 190,192,192- transversalis (Pontoppidan, 1763) – 45,66,88-90,92,107,108,128,172,176, 179,181,186,190,192,195

Hydrachnida – 131Hydradephaga – 10-12,15,16,18,19,22,37,38-63,67-103,106,112-131,133,134, 136-141Hydroglyphus – 39,124,168,175,178,180,183,188,191

- geminus (Fabricius, 1792) – 45,51,63,64,85,88-90,99-101,109,168,175, 178,180,183,188,191,194

Hydroporinae – 48,114,115,122,124,167Hydroporus – 39,40,75,81,116,124,127,128,134,169,170,175,178,181,184,189,191

- angustatus Sturm, 1835 – 75,84,85,88,107,108,116,169,175,178,180,184, 191,194- discretus Fairmaire et Brisout, 1859 – 40,51,53,56,63,65,76,107,169,175,178, 181,189- erythrocephalus (Linnaeus, 1758) – 40,52,66,67,107,169,191- marginatus (Duftschmid, 1805) – 40- memnonius Nicolai, 1822 – 52,53,56,57,64,65,77,110,111,136,169,175, 178,181- nigrita (Fabricius, 1792) – 75- obscurus Sturm, 1835 – 40,59,61,88,89,170,184,194- obsoletus Aubè, 1836 – 40- palustris (Linnaeus, 1761) – 49,52,54,57,62,66,67,107,136,169,175,178,181, 184,191,194- planus (Fabricius, 1781) – 51,63,84,88,89,110,111,127,131,169,175,178,181, 184,189,191,194- pubescens (Gyllenhal, 1808) – 58,59,61,67,92,107,136,169,184,191- sp. – 134- striola (Gyllenhal, 1826) – 40,52,67,107,116,169,191- tessellatus Drapiez, 1819 – 40- tristis (Paykull, 1798) – 40,59,61,66,67,107,170,184,191

Hydrovatus – 39,124,168,175,178,182,185,189,191,194- cuspidatus (Kunze, 1818) –53,58,60,61,63,67,100,107,122,124,168,175,178, 182,185,189,191,194

Hygrobia – 39,167- hermanni (Fabricius 1775) – 39,40,103,104,113,115,123,140,167

Hygrobiidae [=Paelobiidae] – 11,39,41,103,112-115,121,136,140,167Hygrotus – 39,40,122-124,127,169,175,178,180,184,188,191,192,194

- caspius (Wehncke, 1873) – 40- confluens (Fabricius, 1787) – 49,60,61,64,67,77,109,123,168,175,178, 180,184,188,192,194- corpulentus (Schaum, 1864) – 40,66,67,168,192

201

- decoratus (Gyllenhal, 1810) – 49,52,54,59,76,88,107,124,136,168,178,184, 191,194- enneagrammus (Ahrens, 1833) – 45,53,54,63-67,77,101,102,108,123, 168,178,188,192- flaviventris (Motschulsky, 1860) – 40,66,67,168,192- impressopunctatus (Schaller, 1783) – 48,64,75,84-91,109,124,127,128,130, 168,175,178,180,184,188,192,194- inaequalis (Fabricius, 1777) – 45,64,66,75,77,84-86,88,89,93,94,109, 124,127,130,169,175,178,180,184,188,191,194- marklini (Gyllenhal, 1813) – 40- nigrolineatus (Steven in Schünherr, 1808) – 40- pallidulus (Aube, 1850) – 38-40,45,60,61,66,67,77,136,168,184,192- parallelogrammus (Ahrens, 1812) – 49,53,58,60,61,67,77,85,86,89,109, 124,168,175,177,184,192,194- pectoralis (Motschulsky, 1859) – 40- polonicus (Aubè, 1842) – 40- quinquelineatus (Zetterstedt, 1828) – 40,59,61,169,184- saginatus (Schaum, 1857) – 40- versicolor (Schaller, 1783) – 56,59,61,77,101,124,169,180,184

Hymenoptera – 126,131Hyphydrus – 39,40,124,168,175,178,182,185,189,191,194

- ovatus (Linnaeus, 1761) – 45,66,75,89,93,94,107,108,112,122-124,130,168,175,178,182,185,189,191,194

- sanctus Sharp, 1882 – 40Ilybius – 39,40,48,85,87,90-92,95,99,100,110,112,114,117,122-124,128,134,171, 176,179,181,185,186,189,192,195

- ater (De Geer, 1774) – 54,57,110,136,171,179,181- chalconatus (Panzer, 1796) – 51,52,56,77,110,171,176,181- cinctus Sharp, 1882 – 40- fenestratus (Fabricius, 1781) – 40,75,77,89,93,95,110,171,176,179, 181,185, 194- fuliginosus (Fabricius, 1792) – 56,75,77,88,89,110,130,171,176,179,181, 185,189,194- quadriguttatus (Lacordaire, 1835) – 66,77,93,110,130,171,176,179,181,185, 189,192- similis Thomson, 1856 – 40,67,88,89,93,110,171,176,179,181,185,194- subaeneus Erichson, 1837 – 45,56,77,89,93,110,127,171,176,179,181,186, 189,192,194- wasastjernae (Sahlberg, 1824) – 110

Ilyocoris cimicoides (Linnaeus, 1758) – 126Insecta – 10Isobrychus minutus Linnaeus, 1758 – 130Juncus spp. – 105,195Laccophilinae – 114,115,171

202

Laccophilus – 39,120,121,171,172,176,179,181,185,189,192,194- hyalinus (De Geer, 1774) – 52,55,57,60,62,66,67,75,77,107,171,176,179,181, 185,192- minutus (Linnaeus, 1758) – 51,75,77,85,88,89,110,111,113,127,130,140, 171,176,179,181,185,189,192,194- poecilus Klug, 1834 – 61,67,75,77,88,89,94,109,127,130,172,176,179, 181,185,189,192,194

Lemna – 48Leuciscus cephalus (Linnaeus, 1758) – 129

- idus (Linnaeus, 1758) – 129Liopterus – 39,122,124,167,175,178,180,185,188,191,194

- haemorrhoidalis (Fabricius, 1787) – 56,66,86,87,89,107,167,175,178,180, 183,188,191,194

Lymnaea – 118Mammalia – 126Meruidae –11Mollusca – 126Myriophyllum spp.– 48Nebrioporus – 39, 40,75,124,170,178,191

- airumlus (Kolenati, 1845) – 40,75- canaliculatus (Lacordaire, 1835) –40- ceresyi (Aubè, 1838) – 54,64-67,77,101,102,110,111,122-124,128,170, 178,191- elegans (Panzer, 1794) – 40- luctuosus (Aubè, 1838) – 40

Nematoda – 126,131Nematomorpha – 126,131Nepa cinerea Linnaeus, 1758 –126Noteridae – 11,39,41,51,53,56,58,64,66,84-88,91-95,100,106,112-115,136,140,167Noterus – 39,118,167,175,177,178,180,183,188,191,194

- clavicornis (De Geer, 1774) – 59,63,77,84,85,88,89,100,106,127,167, 175,177,180,183,188,191,194- crassicornis (O.F.Muller, 1776) – 59,66,75,84-86,88,89,92,93,106,127,130, 167,175,178,180,183,188,191,194

Notonecta glauca Linnaeus, 1758 – 126,127,128,141- viridis Delcourt,1909 – 126,127,128

Nycticorax nycticorax (Linnaeus, 1758) – 130Odonata – 118,123,126Oligochaeta – 126Orectochilinae – 174Orectochilus – 39,78,174,182

- villosus (O.F.Muller, 1776) – 47,56,76-78,111,138,174,182Oreodytes sanmarkii (C.R. Sahlberg, 1826) – 76Ostracoda – 123,124

203

Paelobiidae [=Hygrobiiade] – 11,103Peltodytes – 39,106,117,118,124,166,175,180,182,188,191,194

- caesus (Duftschmid, 1805) – 77,81,86,88,89,93,94,104,106,124,127,129,130, 166,175

Plantae Vasculares – 126Perca fluviatilis (Linnaeus, 1758) – 128Philomachus pugnax (Linnaeus, 1758) – 130Phragmites – 47,83,105,195Physidae – 118Pisces – 126Plagiorchis sp. – 131,141Platambus – 39,171,176,181,185

- maculatus (Linnaeus, 1758) –51,52,56,62,75,77,110,122,124,171,176,181,185Plecoptera – 118,126Pleurogenoides medians (Olson, 1876) – 131Polychaeta – 126Polyphaga – 10Porhydrus – 39,40,124,170,176,178,181,185,191,194

- lineatus (Fabricius, 1775) – 53,85,89,93,94,109,127,170,176,178,181, 185,191,192,194,- obliquesignatus (Bielz, 1852) – 40

Potamogeton 48Rana – 130,141Ranatra linearis (Linnaeus, 1758) – 126-128,141Reptilia – 126Rhantus – 39,40,85-87,92,95,107,114,120-122,127,128,134,172,176,179,181,186, 189,192,195

- bistriatus (Bergsträsser, 1778) – 53,54,66,78,85,89,172,176,179,186,192,195- consputus (Sturm, 1834) – 40- exsoletus (Forster, 1771) – 40,54,107,136,172,179- frontalis (Marsham, 1802) – 77,86,87,93,107,108,172,176,179,181,186,189, 192,195- grapii (Gyllenhal, 1808) – 54,59,66,67,92,107,136,172,179,186,192,195- latitans Sharp, 1882 – 40,53,55,58,77,89,92-94,107,108,130,172,179,186,195- notaticollis (Aubè, 1837) – 40- suturalis (W.S. MacLeay, 1825) – 48,51,63,64,75,77,86,88,90,91,95,98,99, 109,113,127,128,130,140,172,176,179,181,186,189,192,195- suturellus (Harris, 1828) – 40

Rotatoria – 10,123,126Rutilus rutilus (Linnaeus, 1758) – 129Scardinius erythrophthalmus (Linnaeus, 1758) – 129Scarodytes halensis (Fabricius, 1787) – 40Silurus glanis Linnaeus, 1758 – 129

204

Stercorarius parasiticus (Linnaeus, 1758) – 130Stigeoclonium sp. – 49Suctoria – 131Suphrodytes – 39,124,169,191

- dorsalis (Fabricius, 1787) – 66,67,107,169,191Thamnophis elegans (Baird et Girard, 1853) – 118Thymallus – 128Trematoda – 126,131,133Trichoptera – 118,123,126Tringa erythropus (Pallas, 1764) – 130

- glareola Linnaeus, 1758 – 130Tubifex – 123Turdus merula Linnaeus, 1758 – 131Typha spp. – 83Vorticella – 133,141Zoothamnium – 133,141

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