13F-HR1wfc_13f-3312012.txt

UNITED STATES

SECURITIES AND EXCHANGE COMMISSION

Washington, D.C. 20549

FORM 13F

FORM 13F COVER PAGE

Report for the Calendar Year or Quarter Ended: 3/31/2012

Check here if Amendment []; Amendment Number:This Amendment (Check only one.): [] is a restatement [] adds new holdings entries.

Institutional Investment Manager Filing this Report:

Name: Wells Fargo & CompanyAddress: 420 Montgomery Street San Francisco, CA 94163

13F File Number: 028-00165

The institutional investment manager filing this report and the person by whomit is signed hereby represent that the person signing the report isauthorized to submit it, that all information contained herein is true,correct and complete, and that it is understood that all required items,statements, schedules, lists, and tables, are considered integral partsof this form.

Person Signing this Report on Behalf of Reporting Manager:

Name: Jane E. WashingtonTitle: VP Trust Operations, Management Support ServicesPhone: 612-667-4922

Filing Contact: Stuart Portoghese (612) 667-0161

Signature, Place and Date of Signing

/s/ Jane E. Washington Mpls, MN April 30, 2012

Report Type (Check only one.):

[X] 13F Holdings Report[ ] 13F Notice[ ] 13F Combination Report

List of Other Managers Reporting for this Manager:

I AM SIGNING THIS REPORT AS REQUIRED BY THE SECURITIES EXCHANGE ACT OF 1934.

FORM 13F SUMMARY PAGE

Report Summary:

Number of Other Included Managers: 14

Form 13F Information Table Entry Total: 18,868

Form 13F Information Table Value Total: $156,600,400 (in thousands)

List of Other Included Managers:

No. 13F File # Name

02 028-01523 Peregrine Capital Management, Inc.03 028-04413 Wells Capital Management Incorporated04 028-01341 Wells Fargo Bank, N.A.05 028-11406 Wells Fargo Funds Management, LLC07 028-10450 Nelson Capital Management, LLC08 028-12161 Wells Fargo Alternative Asset Management LLC.10 028-05723 Metropolitan West Capital Management, LLC.13 028-13117 Wells Fargo Advisors Financial Network, LLC.15 028-13118 Wells Fargo Advisors, LLC17 028-12628 Wells Fargo Delaware Trust Company, N.A.24 028-13600 Wells Fargo Securities, LLC35 028-13622 WIH Holdings39 028-06571 Norwest Limited LP, LLLP43 028-14644 Overland Advisors, LLC50 028-03504 WFC Holdings Corporation73 028-06003 Golden Capital Management, LLC

Wells Fargo & Company Form 13F As of 3/31/2012

Item 4: Fair Item 8: Item 3: Market Item 5: Item 6: Voting AuthorityItem 1: Item 2: Cusip Value SHARES/ SH/ PUT/ Invstmt Item 7: (A) (B) (C)Name of Issuer Title of Class Number (000) PRN AMT PRN CALL Dscretn Mgrs Sole Shared None----------------------------- --------------- --------- --------- ---------- --- ---- ------- ------- --------- --------- ---------AAON INC Equity 000360206 10 472 SH DEFINED 15 472 0 0AAON INC Equity 000360206 1 66 SH DEFINED 24 66 0 0AAON INC Equity 000360206 100 4958 SH DEFINED 4 4958 0 0A A R CORPORATION Equity 000361105 3 187 SH DEFINED 15 187 0 0A A R CORPORATION Equity 000361105 7413 406190 SH DEFINED 2 191280 0 214910A A R CORPORATION Equity 000361105 3 139 SH

DEFINED 24 139 0 0A A R CORPORATION Equity 000361105 13 691 SH DEFINED 4 691 0 0ABB LTD -SPONS ADR Equity 000375204 196 9605 SH DEFINED 13 9605 0 0ABB LTD -SPONS ADR Equity 000375204 7714 377975 SH DEFINED 15 377975 0 0ABB LTD -SPONS ADR Equity 000375204 25783 1263251 SH DEFINED 4 1183114 25653 54484ABB LTD -SPONS ADR Equity 000375204 10 490 SH DEFINED 7 490 0 0ABB LTD -SPONS ADR Equity 000375204 375 18361 SH DEFINED 18361 0 0ACCO BRANDS CORP Equity 00081T108 44 3533 SH DEFINED 15 3533 0 0ACCO BRANDS CORP Equity 00081T108 2 193 SH DEFINED 24 193 0 0ACCO BRANDS CORP Equity 00081T108 25888 2086061 SH DEFINED 3 399124 0 1686937ACCO BRANDS CORP Equity 00081T108 13 1054 SH DEFINED 4 726 328 0ACCO BRANDS CORP Equity 00081T108 247 19926 SH DEFINED 5 19926 0 0ACNB CORP Equity 000868109 0 0 SH DEFINED 15 0 0 0ABM INDUSTRIES INC Equity 000957100 68 2800 SH DEFINED 15 2800 0 0ABM INDUSTRIES INC Equity 000957100 5 186 SH DEFINED 24 186 0 0ABM INDUSTRIES INC Equity 000957100 64690 2662148 SH DEFINED 3 547740 0 2114408ABM INDUSTRIES INC Equity 000957100 78 3204 SH DEFINED 4 3204 0 0ABM INDUSTRIES INC Equity 000957100 647 26617 SH DEFINED 5 26617 0 0AEP INDUSTRIES CMNSTOCK 001031103 1 15 SH DEFINED 24 15 0 0AEP INDUSTRIES INC CMN CommonStock 001031103 172 4949 SH DEFINED 4949 0 0AFC ENTERPRISES Equity 00104Q107 15 912 SH DEFINED 13 912 0 0AFC ENTERPRISES Equity 00104Q107 0 1 SH DEFINED 15 1 0 0AFC ENTERPRISES Equity 00104Q107 1 87 SH DEFINED 24 87 0 0AFC ENTERPRISES Equity 00104Q107 170 10043 SH DEFINED 4 7243 0 2800AFLAC INC Call 001055102 920 20000 Sh Call DEFINED 4 20000 0 0AFLAC INC DOMESTIC COMMON 001055102 8789 191098 SH DEFINED 13 191098 0 0AFLAC INC DOMESTIC COMMON 001055102 181979 3956924 SH DEFINED 15 3956924 0 0AFLAC INC DOMESTIC COMMON 001055102 216 4694 SH DEFINED 17 4694 0 0AFLAC INC DOMESTIC COMMON 001055102 2833 61605 SH DEFINED 3 0 0 61605AFLAC INC DOMESTIC COMMON 001055102 142502 3098545 SH DEFINED 4 2938395 106893 53257AFLAC INC DOMESTIC COMMON 001055102 177 3858 SH DEFINED 7 3858 0 0

AFLAC INC DOMESTIC COMMON 001055102 17031 370313 SH DEFINED 73 138229 0 232084AFLAC INC DOMESTIC COMMON 001055102 542 11775 SH DEFINED 11775 0 0AGCO CORP Equity 001084102 52 1106 SH DEFINED 13 1106 0 0AGCO CORP Equity 001084102 2991 63361 SH DEFINED 15 63361 0 0AGCO CORP Equity 001084102 35 746 SH DEFINED 4 278 142 326AGCO CORP Equity 001084102 18 385 SH DEFINED 5 385 0 0AGCO CORP Equity 001084102 2658 56312 SH DEFINED 73 28174 0 28138AGIC CONV & INCOME FD Closed-End Equi 001190107 25 2599 SH DEFINED 13 2599 0 0AGIC CONV & INCOME FD Closed-End Equi 001190107 7006 737450 SH DEFINED 15 737450 0 0AGIC CONV & INCOME FD Closed-End Equi 001190107 11 1160 SH DEFINED 4 1160 0 0AGIC CONV & INCOME FD II Closed-End Equi 001191105 240 27345 SH DEFINED 13 27345 0 0AGIC CONV & INCOME FD II Closed-End Equi 001191105 271 30926 SH DEFINED 15 30926 0 0AGIC EQUITY & CONV INCM Closed-End Equi 00119P102 187 10551 SH DEFINED 13 10551 0 0AGIC EQUITY & CONV INCM Closed-End Equi 00119P102 10118 569375 SH DEFINED 15 569375 0 0AGIC INTL & PERM STRAT Closed-End Equi 00119Q100 85 7732 SH DEFINED 15 7732 0 0AGIC GLOBAL EQUITY & Closed-End Equi 00119R108 282 19789 SH DEFINED 15 19789 0 0AGL RESOURCES INC Equity 001204106 447 11406 SH DEFINED 13 11406 0 0AGL RESOURCES INC Equity 001204106 9468 241419 SH DEFINED 15 241419 0 0AGL RESOURCES INC Equity 001204106 174 4428 SH DEFINED 3 0 0 4428AGL RESOURCES INC Equity 001204106 2075 52905 SH DEFINED 4 49816 2293 796AGL RESOURCES INC Equity 001204106 1152 29365 SH DEFINED 73 1131 0 28234AG MORTGAGE INVESTMENT Equity 001228105 1071 54280 SH DEFINED 15 54280 0 0AG MORTGAGE INVESTMENT Equity 001228105 1123 56878 SH DEFINED 3 55203 0 1675AG MORTGAGE INVESTMENT Equity 001228105 57 2879 SH DEFINED 5 2879 0 0AH BELO CORP CMNSTOCK 001282102 0 65 SH DEFINED 24 65 0 0AH BELO CORP-A COM 001282102 6901 1411266 SH DEFINED 3 78535 0 1332731AH BELO CORP Equity 001282102 0 2 SH DEFINED 15 2 0 0AES CORP Equity 00130H105 416 31813 SH DEFINED 13 31813 0 0AES CORP Equity 00130H105 2056 157350 SH DEFINED 15 157350 0 0AES CORP Equity 00130H105 318 24324 SH DEFINED 3 0 0 24324AES CORP Equity 00130H105 809 61874 SH

DEFINED 4 59561 503 1810AES CORP Equity 00130H105 9798 749647 SH DEFINED 73 283490 0 466157AK STEEL HOLDINGS CORP CMNSTOCK 001547108 4 496 SH DEFINED 13 496 0 0AK STEEL HOLDINGS CORP CMNSTOCK 001547108 1513 200113 SH DEFINED 15 200113 0 0AK STEEL HOLDINGS CORP CMNSTOCK 001547108 117 15504 SH DEFINED 4 8890 0 6614COHEN & STEERS ETF Equity 00162Q106 266 7270 SH DEFINED 13 7270 0 0COHEN & STEERS ETF Equity 00162Q106 2344 64140 SH DEFINED 15 64140 0 0ALPS TRUST ETF Equity 00162Q205 362 9241 SH DEFINED 15 9241 0 0ALPS TRUST ETF Equity 00162Q304 336 7341 SH DEFINED 13 7341 0 0ALPS TRUST ETF Equity 00162Q304 595 12993 SH DEFINED 15 12993 0 0ALPS ETF TR ETF Equity 00162Q809 4 103 SH DEFINED 13 103 0 0ALPS ETF TR ETF Equity 00162Q809 115 2825 SH DEFINED 15 2825 0 0ALERIAN MLP ETF Equity 00162Q866 8064 484627 SH DEFINED 13 484627 0 0ALERIAN MLP ETF Equity 00162Q866 55676 3345906 SH DEFINED 15 3345906 0 0AMAG PHARMACEUTICALS Equity 00163U106 4 250 SH DEFINED 15 250 0 0AMAG PHARMACEUTICALS Equity 00163U106 1 74 SH DEFINED 24 74 0 0AMAG PHARMACEUTICALS Equity 00163U106 83 5224 SH DEFINED 73 1229 0 3995AMC NETWORKS INC A W I DOMESTIC COMMON 00164V103 92 2059 SH DEFINED 13 2059 0 0AMC NETWORKS INC A W I DOMESTIC COMMON 00164V103 692 15496 SH DEFINED 15 15496 0 0AMC NETWORKS INC A W I DOMESTIC COMMON 00164V103 219 4918 SH DEFINED 4 4735 0 183AMC NETWORKS INC A W I DOMESTIC COMMON 00164V103 3 71 SH DEFINED 7 71 0 0AMN HEALTHCARE SERVICES None Provided 001744101 14842 2449193 SH DEFINED 10 1586200 0 862993AMN HEALTHCARE SERVICES None Provided 001744101 1 139 SH DEFINED 24 139 0 0AMN HEALTHCARE SERVICES None Provided 001744101 2757 455023 SH DEFINED 3 25622 0 429401AOL INC Equity 00184X105 7 374 SH DEFINED 13 374 0 0AOL INC Equity 00184X105 90 4693 SH DEFINED 15 4693 0 0AOL INC Equity 00184X105 32 1688 SH DEFINED 4 1053 29 606API TECHNOLOGIES CORP Equity 00187E203 0 83 SH DEFINED 13 83 0 0API TECHNOLOGIES CORP Equity 00187E203 0 83 SH DEFINED 15 83 0 0AT & T INC DOMESTIC COMMON 00206R102 42338 1355699 SH DEFINED 13 1355699 0 0AT & T INC DOMESTIC COMMON 00206R102 369924 11845140 SH DEFINED 15 11845140 0 0

AT & T INC DOMESTIC COMMON 00206R102 1786 57196 SH DEFINED 17 57196 0 0AT & T INC DOMESTIC COMMON 00206R102 18994 608199 SH DEFINED 3 220100 0 388099AT & T INC DOMESTIC COMMON 00206R102 178156 5704615 SH DEFINED 4 4666980 867439 170196AT & T INC DOMESTIC COMMON 00206R102 98 3145 SH DEFINED 5 3145 0 0AT & T INC DOMESTIC COMMON 00206R102 283 9065 SH DEFINED 7 6270 0 2795AT & T INC DOMESTIC COMMON 00206R102 86052 2755424 SH DEFINED 73 604446 0 2150978AT & T INC DOMESTIC COMMON 00206R102 443 14178 SH DEFINED 14178 0 0AT & T INC PUT 00206R102 1827 58500 SH PUT DEFINED 15 58500 0 0AT & T INC PUT 00206R102 375 12000 SH PUT DEFINED 4 12000 0 0ATMI CORP None Provided 00207R101 7713 331018 SH DEFINED 10 214370 0 116648ATMI CORP None Provided 00207R101 225 9649 SH DEFINED 13 9649 0 0ATMI CORP None Provided 00207R101 800 34348 SH DEFINED 15 34348 0 0ATMI CORP None Provided 00207R101 3 111 SH DEFINED 24 111 0 0ATMI CORP None Provided 00207R101 29173 1252077 SH DEFINED 3 43600 0 1208477ATMI CORP None Provided 00207R101 433 18590 SH DEFINED 4 18590 0 0ATMI CORP None Provided 00207R101 59 2536 SH DEFINED 5 2536 0 0ATP OIL & GAS CORP Equity 00208J108 253 34476 SH DEFINED 15 34476 0 0ATP OIL & GAS CORP Equity 00208J108 1 156 SH DEFINED 24 156 0 0ARCA BIOPHARMA INC Equity 00211Y100 0 16 SH DEFINED 15 16 0 0AU OPTRONICS CORP-ADR Equity 002255107 21 4514 SH DEFINED 15 4514 0 0AU OPTRONICS CORP-ADR Equity 002255107 2 339 SH DEFINED 4 86 253 0AVI BIOPHARMA INC Equity 002346104 2 1400 SH DEFINED 13 1400 0 0AVI BIOPHARMA INC Equity 002346104 70 45701 SH DEFINED 15 45701 0 0AVI BIOPHARMA INC Equity 002346104 1 475 SH DEFINED 24 475 0 0AV HOMES INC Equity 00234P102 0 31 SH DEFINED 15 31 0 0AV HOMES INC Equity 00234P102 0 31 SH DEFINED 24 31 0 0AVX CORPORATION NEW Equity 002444107 3 209 SH DEFINED 15 209 0 0AVX CORPORATION NEW Equity 002444107 13645 1029059 SH DEFINED 3 81413 0 947646AVX CORPORATION NEW Equity 002444107 34 2587 SH DEFINED 4 306 0 2281AXT INC Equity 00246W103 169 26600 SH DEFINED 15 26600 0 0AXT INC Equity 00246W103 1 112 SH

DEFINED 24 112 0 0AZZ INC CMNSTOCK 002474104 2 44 SH DEFINED 24 44 0 0AZZ INCORPORATED COMMON STOCK 002474104 176 3405 SH DEFINED 4 3405 0 0AZZ INCORPORATED Equity 002474104 0 0 SH DEFINED 15 0 0 0AARONS INC Equity 002535300 388 14975 SH DEFINED 15 14975 0 0AARONS INC Equity 002535300 187 7231 SH DEFINED 4 4531 0 2700AASTROM BIOSCIENCES INC Equity 00253U305 1 562 SH DEFINED 13 562 0 0ABAXIS INC Equity 002567105 759 26070 SH DEFINED 13 26070 0 0ABAXIS INC Equity 002567105 222 7630 SH DEFINED 15 7630 0 0ABAXIS INC Equity 002567105 2 79 SH DEFINED 24 79 0 0ABAXIS INC Equity 002567105 6 207 SH DEFINED 4 207 0 0ABBOTT LABORATORIES DOMESTIC COMMON 002824100 233943 3816982 SH DEFINED 10 2889064 0 927918ABBOTT LABORATORIES DOMESTIC COMMON 002824100 17947 292815 SH DEFINED 13 292815 0 0ABBOTT LABORATORIES DOMESTIC COMMON 002824100 103799 1693574 SH DEFINED 15 1693574 0 0ABBOTT LABORATORIES DOMESTIC COMMON 002824100 3848 62782 SH DEFINED 17 62782 0 0ABBOTT LABORATORIES DOMESTIC COMMON 002824100 161524 2635404 SH DEFINED 3 1119196 0 1516208ABBOTT LABORATORIES DOMESTIC COMMON 002824100 346439 5652459 SH DEFINED 4 5034823 458706 158930ABBOTT LABORATORIES DOMESTIC COMMON 002824100 31985 521862 SH DEFINED 5 521862 0 0ABBOTT LABORATORIES DOMESTIC COMMON 002824100 5815 94874 SH DEFINED 7 88579 0 6295ABBOTT LABORATORIES DOMESTIC COMMON 002824100 44616 727957 SH DEFINED 73 157662 0 570295ABBOTT LABORATORIES DOMESTIC COMMON 002824100 648 10580 SH DEFINED 10580 0 0ABBOTT LABORATORIES PUT 002824100 1532 25000 SH PUT DEFINED 4 25000 0 0ABERCROMBIE & FITCH CO CL A DOMESTIC COMMON 002896207 4 90 SH DEFINED 13 90 0 0ABERCROMBIE & FITCH CO CL A DOMESTIC COMMON 002896207 1081 21800 SH DEFINED 15 21800 0 0ABERCROMBIE & FITCH CO CL A DOMESTIC COMMON 002896207 26339 530923 SH DEFINED 3 9811 0 521112ABERCROMBIE & FITCH CO CL A DOMESTIC COMMON 002896207 111 2229 SH DEFINED 4 2139 90 0ABERCROMBIE & FITCH CO CL A DOMESTIC COMMON 002896207 1065 21475 SH DEFINED 73 827 0 20648ABERDEEN ASIA PACIFIC Closed-End Equi 003009107 304 41694 SH DEFINED 13 41694 0 0ABERDEEN ASIA PACIFIC Closed-End Equi 003009107 11839 1623951 SH DEFINED 15 1623951 0 0ABERDEEN ASIA PACIFIC Closed-End Equi 003009107 341 46742 SH DEFINED 4 22902 3840 20000ABERDEEN AUSTRALIA Closed-End Equi 003011103 1177 109507 SH DEFINED 15 109507 0 0

ABERDEEN AUSTRALIA Closed-End Equi 003011103 27 2510 SH DEFINED 4 510 0 2000ABERDEEN GLOBAL INCOME Closed-End Equi 003013109 41 2900 SH DEFINED 13 2900 0 0ABERDEEN GLOBAL INCOME Closed-End Equi 003013109 1211 86612 SH DEFINED 15 86612 0 0ABERDEEN ISRAEL FD INC Closed-End Equi 00301L109 15 1101 SH DEFINED 15 1101 0 0ABERDEEN ISRAEL FD INC Closed-End Equi 00301L109 2897 207079 SH DEFINED 3 148310 0 58769ABERDEEN ISRAEL FD INC Closed-End Equi 00301L109 202 14458 SH DEFINED 4 14458 0 0ABERDEEN EMERGING MKTS Closed-End Equi 00301T102 37 1901 SH DEFINED 15 1901 0 0ABERDEEN EMERGING MKTS Closed-End Equi 00301T102 5695 294008 SH DEFINED 3 224583 0 69425ABERDEEN EMERGING MKTS Closed-End Equi 00301T102 8 420 SH DEFINED 4 0 420 0ABERDEEN CHILE FD INC Closed-End Equi 00301W105 4 190 SH DEFINED 13 190 0 0ABERDEEN CHILE FD INC Closed-End Equi 00301W105 70 3628 SH DEFINED 15 3628 0 0ABERDEEN INDONESIA FD Closed-End Equi 00305P106 20 1607 SH DEFINED 13 1607 0 0ABERDEEN INDONESIA FD Closed-End Equi 00305P106 120 9391 SH DEFINED 15 9391 0 0ABERDEEN INDONESIA FD Closed-End Equi 00305P106 3 200 SH DEFINED 7 200 0 0LATIN AMERICA EQUITY FD Closed-End Equi 00306K106 335 9429 SH DEFINED 15 9429 0 0LATIN AMERICA EQUITY FD Closed-End Equi 00306K106 6506 183212 SH DEFINED 3 146575 0 36637ABIOMED INC Equity 003654100 55 2502 SH DEFINED 15 2502 0 0ABIOMED INC Equity 003654100 2 111 SH DEFINED 24 111 0 0ABIOMED INC Equity 003654100 7 295 SH DEFINED 4 295 0 0ABITIBIBOWATER INC Equity 003687209 38 2673 SH DEFINED 15 2673 0 0ABITIBIBOWATER INC Equity 003687209 2398 167902 SH DEFINED 24 167902 0 0ABOVENET INC Equity 00374N107 825 9969 SH DEFINED 15 9969 0 0ABOVENET INC Equity 00374N107 7 82 SH DEFINED 24 82 0 0ABOVENET INC Equity 00374N107 200 2410 SH DEFINED 4 2410 0 0ABOVENET INC Equity 00374N107 146 1769 SH DEFINED 73 416 0 1353ABRAXAS PETE CORP Equity 003830106 75 23935 SH DEFINED 15 23935 0 0ABRAXAS PETE CORP Equity 003830106 1 289 SH DEFINED 24 289 0 0ABRAXAS PETE CORP Equity 003830106 12 4000 SH DEFINED 4 4000 0 0ACACIA RESEARCH ACACIA TEC DOMESTIC COMMON 003881307 8 193 SH DEFINED 13 193 0 0ACACIA RESEARCH ACACIA TEC DOMESTIC COMMON 003881307 442 10601 SH DEFINED 15 10601 0 0ACACIA RESEARCH ACACIA TEC DOMESTIC COMMON 003881307 6 150 SH

DEFINED 24 150 0 0ACACIA RESEARCH ACACIA TEC DOMESTIC COMMON 003881307 219 5241 SH DEFINED 4 5241 0 0ACADIA HEALTHCARE COM 00404A109 4075 250000 SH DEFINED 2 93590 0 156410ACADIA PHARMACEUTICALS Equity 004225108 2 800 SH DEFINED 15 800 0 0ACADIA REALTY TR Equity 004239109 149 6605 SH DEFINED 13 6605 0 0ACADIA REALTY TR Equity 004239109 0 13 SH DEFINED 15 13 0 0ACADIA REALTY TR Equity 004239109 3 149 SH DEFINED 24 149 0 0ACCELRYS INC Equity 00430U103 5 619 SH DEFINED 13 619 0 0ACCELRYS INC Equity 00430U103 2 195 SH DEFINED 24 195 0 0ACCELRYS INC Equity 00430U103 20763 2601872 SH DEFINED 3 630658 0 1971214ACCELRYS INC Equity 00430U103 1 107 SH DEFINED 4 107 0 0ACCELRYS INC Equity 00430U103 270 33843 SH DEFINED 5 33843 0 0ACCESS NATIONAL CORP Equity 004337101 0 0 SH DEFINED 15 0 0 0ACCRETIVE HEALTH INC Equity 00438V103 169 8459 SH DEFINED 15 8459 0 0ACCRETIVE HEALTH INC Equity 00438V103 4789 239830 SH DEFINED 2 86810 0 153020ACCRETIVE HEALTH INC Equity 00438V103 3 140 SH DEFINED 24 140 0 0ACCURAY INC Equity 004397105 104 14800 SH DEFINED 15 14800 0 0ACCURAY INC Equity 004397105 2 240 SH DEFINED 24 240 0 0ACCURAY INC Equity 004397105 141 20000 SH DEFINED 4 20000 0 0ACCURIDE CORP NEW Equity 00439T206 0 1 SH DEFINED 15 1 0 0ACCURIDE CORP NEW Equity 00439T206 1 142 SH DEFINED 24 142 0 0ACETO CORP Equity 004446100 65 6854 SH DEFINED 15 6854 0 0ACETO CORP Equity 004446100 1 93 SH DEFINED 24 93 0 0ACETO CORP Equity 004446100 147 15490 SH DEFINED 73 3646 0 11844ACHILLION PHARMACEUTICAL Equity 00448Q201 486 50749 SH DEFINED 15 50749 0 0ACHILLION PHARMACEUTICAL Equity 00448Q201 2 166 SH DEFINED 24 166 0 0ACI WORLDWIDE INC DOMESTIC COMMON 004498101 4 100 SH DEFINED 13 100 0 0ACI WORLDWIDE INC DOMESTIC COMMON 004498101 678 16839 SH DEFINED 15 16839 0 0ACI WORLDWIDE INC DOMESTIC COMMON 004498101 6 137 SH DEFINED 24 137 0 0ACI WORLDWIDE INC DOMESTIC COMMON 004498101 28497 707655 SH DEFINED 3 91996 0 615659ACI WORLDWIDE INC DOMESTIC COMMON 004498101 638 15853 SH DEFINED 4 15853 0 0

ACME PACKET INC DOMESTIC COMMON 004764106 188 6819 SH DEFINED 13 6819 0 0ACME PACKET INC DOMESTIC COMMON 004764106 4144 150580 SH DEFINED 15 150580 0 0ACME PACKET INC DOMESTIC COMMON 004764106 11293 410340 SH DEFINED 3 11400 0 398940ACME PACKET INC DOMESTIC COMMON 004764106 676 24579 SH DEFINED 4 24498 81 0ACME PACKET INC DOMESTIC COMMON 004764106 5 168 SH DEFINED 5 168 0 0ACME PACKET INC DOMESTIC COMMON 004764106 10 360 SH DEFINED 7 360 0 0ACME UNITED CORP Equity 004816104 0 1 SH DEFINED 15 1 0 0ACORDA THERAPEUTICS INC Equity 00484M106 3 100 SH DEFINED 13 100 0 0ACORDA THERAPEUTICS INC Equity 00484M106 6 220 SH DEFINED 15 220 0 0ACORDA THERAPEUTICS INC Equity 00484M106 4 139 SH DEFINED 24 139 0 0ACORDA THERAPEUTICS INC Equity 00484M106 1954 73600 SH DEFINED 3 73600 0 0ACORDA THERAPEUTICS INC Equity 00484M106 137 5157 SH DEFINED 73 1214 0 3943ACTIVE POWER INC Equity 00504W100 12 15006 SH DEFINED 15 15006 0 0ACTIVE POWER INC Equity 00504W100 0 280 SH DEFINED 24 280 0 0ACTIVE NETWORK INC Equity 00506D100 3 200 SH DEFINED 15 200 0 0ACTIONS SEMICONDUCTOR Equity 00507E107 1 547 SH DEFINED 15 547 0 0ACTIVISION BLIZZARD INC CMNSTOCK 00507V109 95 7420 SH DEFINED 13 7420 0 0ACTIVISION BLIZZARD INC CMNSTOCK 00507V109 4155 324128 SH DEFINED 15 324128 0 0ACTIVISION BLIZZARD INC CMNSTOCK 00507V109 12128 946000 SH DEFINED 3 0 0 946000ACTIVISION BLIZZARD INC CMNSTOCK 00507V109 1963 153130 SH DEFINED 4 147935 1603 3592ACTIVISION BLIZZARD INC CMNSTOCK 00507V109 387 30214 SH DEFINED 73 7094 0 23120ACTUATE CORPORA CMNSTOCK 00508B102 1 134 SH DEFINED 24 134 0 0ACTUANT CORPORATION DOMESTIC COMMON 00508X203 270 9330 SH DEFINED 15 9330 0 0ACTUANT CORPORATION DOMESTIC COMMON 00508X203 5564 191930 SH DEFINED 2 69570 0 122360ACTUANT CORPORATION DOMESTIC COMMON 00508X203 7 241 SH DEFINED 24 241 0 0ACTUANT CORPORATION DOMESTIC COMMON 00508X203 14418 497334 SH DEFINED 3 219219 0 278115ACTUANT CORPORATION DOMESTIC COMMON 00508X203 596 20569 SH DEFINED 4 20069 500 0ACTUANT CORPORATION DOMESTIC COMMON 00508X203 428 14748 SH DEFINED 5 14748 0 0ACTUANT CORPORATION DOMESTIC COMMON 00508X203 202 6977 SH DEFINED 73 1642 0 5335ACTUANT CORPORATION Corporate Bond 00508XAB0 2265 1550000 PRN DEFINED 24 1550000 0 0ACTUANT CORPORATION Corporate Bond 00508XAB0 9 6000 SH

DEFINED 15 6000 0 0ACUITY BRANDS INC Equity 00508Y102 321 5107 SH DEFINED 15 5107 0 0ACUITY BRANDS INC Equity 00508Y102 10 152 SH DEFINED 24 152 0 0ACUITY BRANDS INC Equity 00508Y102 1245 19809 SH DEFINED 4 19809 0 0ACURA PHARMACEUTICALS Equity 00509L703 66 19000 SH DEFINED 13 19000 0 0ACURA PHARMACEUTICALS Equity 00509L703 7 2000 SH DEFINED 15 2000 0 0ACURA PHARMACEUTICALS Equity 00509L703 0 38 SH DEFINED 24 38 0 0ACURA PHARMACEUTICALS Equity 00509L703 65 18850 SH DEFINED 4 18850 0 0ACXIOM CORP CMNSTOCK 005125109 4 284 SH DEFINED 24 284 0 0ACXIOM CORP COM COMMON STOCK 005125109 574 39085 SH DEFINED 4 39085 0 0Acxiom Corp. Common Stock 005125109 196 13325 SH DEFINED 73 3137 0 10188ACXIOM CORP Equity 005125109 0 12 SH DEFINED 15 12 0 0ADA-ES INC Equity 005208103 9 390 SH DEFINED 13 390 0 0ADA-ES INC Equity 005208103 405 16666 SH DEFINED 15 16666 0 0ADA-ES INC Equity 005208103 134 5500 SH DEFINED 4 5500 0 0ADAMS EXPRESS CO Closed-End Equi 006212104 5 481 SH DEFINED 13 481 0 0ADAMS EXPRESS CO Closed-End Equi 006212104 253 23038 SH DEFINED 15 23038 0 0ADAMS EXPRESS CO Closed-End Equi 006212104 3114 283606 SH DEFINED 3 283606 0 0ADAMS EXPRESS CO Closed-End Equi 006212104 356 32458 SH DEFINED 4 27195 5263 0ADAMS RES & ENERGY INC < Equity 006351308 0 1 SH DEFINED 15 1 0 0ADEPT TECHNOLOGY INC Equity 006854202 0 0 SH DEFINED 15 0 0 0ADMINISTRADORA DE FONDE Equity 00709P108 5 68 SH DEFINED 15 68 0 0ADOBE SYSTEMS INC Equity 00724F101 371 10818 SH DEFINED 13 10818 0 0ADOBE SYSTEMS INC Equity 00724F101 8617 251147 SH DEFINED 15 251147 0 0ADOBE SYSTEMS INC Equity 00724F101 2112 61566 SH DEFINED 3 0 0 61566ADOBE SYSTEMS INC Equity 00724F101 6314 184035 SH DEFINED 4 170246 8589 5200ADOBE SYSTEMS INC Equity 00724F101 5 154 SH DEFINED 7 154 0 0ADOBE SYSTEMS INC Equity 00724F101 4634 135051 SH DEFINED 73 4771 0 130280ADTRAN INC Equity 00738A106 8 244 SH DEFINED 13 244 0 0ADTRAN INC Equity 00738A106 254 8151 SH DEFINED 15 8151 0 0ADTRAN INC Equity 00738A106 7 227 SH DEFINED 24 227 0 0

ADTRAN INC Equity 00738A106 421 13503 SH DEFINED 4 13234 269 0ADVANCE AMERICA CASH Equity 00739W107 1452 138412 SH DEFINED 15 138412 0 0ADVANCE AUTO PARTS INC DOMESTIC COMMON 00751Y106 405 4567 SH DEFINED 13 4567 0 0ADVANCE AUTO PARTS INC DOMESTIC COMMON 00751Y106 21040 237553 SH DEFINED 15 237553 0 0ADVANCE AUTO PARTS INC DOMESTIC COMMON 00751Y106 615 6946 SH DEFINED 4 6370 25 551ADVANCE AUTO PARTS INC DOMESTIC COMMON 00751Y106 5 55 SH DEFINED 7 55 0 0ADVANCE AUTO PARTS INC DOMESTIC COMMON 00751Y106 94 1064 SH DEFINED 73 221 0 843ADVANCED SEMICONDUCTOR Equity 00756M404 28 5529 SH DEFINED 15 5529 0 0ADVOCAT INC Equity 007586100 0 1 SH DEFINED 15 1 0 0ADVANTEST CORP ADR COMMON STOCK 00762U200 4 278 SH DEFINED 4 278 0 0ADVANTEST CORP Equity 00762U200 0 1 SH DEFINED 15 1 0 0ADVISORY BOARD CO/THE Equity 00762W107 269 3041 SH DEFINED 13 3041 0 0ADVISORY BOARD CO/THE Equity 00762W107 12 133 SH DEFINED 15 133 0 0ADVISORY BOARD CO/THE Equity 00762W107 5 59 SH DEFINED 24 59 0 0ADVISORY BOARD CO/THE Equity 00762W107 115 1302 SH DEFINED 4 1302 0 0ADVENT CLAYMORE GLOBAL Closed-End Equi 007639107 12 1810 SH DEFINED 13 1810 0 0ADVENT CLAYMORE GLOBAL Closed-End Equi 007639107 196 28514 SH DEFINED 15 28514 0 0ADVENT CLAYMORE GLOBAL Closed-End Equi 007639107 22 3232 SH DEFINED 4 3232 0 0ADVENT CLAYMORE Closed-End Equi 00764C109 317 19415 SH DEFINED 13 19415 0 0ADVENT CLAYMORE Closed-End Equi 00764C109 4366 267717 SH DEFINED 15 267717 0 0ADVENT CLAYMORE Closed-End Equi 00764C109 1 39 SH DEFINED 4 39 0 0ADVENT CLAYMORE ENHANCED Closed-End Equi 00765E104 8 800 SH DEFINED 13 800 0 0ADVENT CLAYMORE ENHANCED Closed-End Equi 00765E104 168 16747 SH DEFINED 15 16747 0 0ADVENT CLAYMORE ENHANCED Closed-End Equi 00765E104 17 1741 SH DEFINED 4 0 1741 0ADVANTAGE OIL & GAS LTD Equity 00765F101 290 86650 SH DEFINED 15 86650 0 0AECOM TECHNOLOGY CORP Equity 00766T100 110 4935 SH DEFINED 13 4935 0 0AECOM TECHNOLOGY CORP Equity 00766T100 11 477 SH DEFINED 15 477 0 0AECOM TECHNOLOGY CORP Equity 00766T100 468 20913 SH DEFINED 4 20098 345 470AEGERION PHARMACEUTICALS Equity 00767E102 7 500 SH DEFINED 15 500 0 0AEGERION PHARMACEUTICALS Equity 00767E102 0 32 SH DEFINED 24 32 0 0ADVISORSHARES TR ETF Equity 00768Y206 50 1600 SH

DEFINED 15 1600 0 0PERITUS HIGH YIELD ETF Equity 00768Y503 24 488 SH DEFINED 13 488 0 0PERITUS HIGH YIELD ETF Equity 00768Y503 574 11529 SH DEFINED 15 11529 0 0CAMBRIA GLOBAL ETF Equity 00768Y800 135 5539 SH DEFINED 15 5539 0 0THE ACTIVE BEAR ETF Equity 00768Y883 646 31352 SH DEFINED 13 31352 0 0THE ACTIVE BEAR ETF Equity 00768Y883 1452 70460 SH DEFINED 15 70460 0 0AEGION CORP Equity 00770F104 14 794 SH DEFINED 13 794 0 0AEGION CORP Equity 00770F104 613 34357 SH DEFINED 15 34357 0 0AEGION CORP Equity 00770F104 2 138 SH DEFINED 24 138 0 0AEGION CORP Equity 00770F104 538 30150 SH DEFINED 3 20050 0 10100AEGION CORP Equity 00770F104 61 3400 SH DEFINED 4 3400 0 0AEROFLEX HOLDIN CMNSTOCK 007767106 1 70 SH DEFINED 24 70 0 0AEROFLEX HLDG CORP COM 007767106 7914 710370 SH DEFINED 2 333430 0 376940AEROPOSTALE INC None Provided 007865108 26109 1207615 SH DEFINED 10 784740 0 422875AEROPOSTALE INC None Provided 007865108 3 119 SH DEFINED 13 119 0 0AEROPOSTALE INC None Provided 007865108 95 4417 SH DEFINED 15 4417 0 0AEROPOSTALE INC None Provided 007865108 15363 710600 SH DEFINED 2 334900 0 375700AEROPOSTALE INC None Provided 007865108 6 283 SH DEFINED 24 283 0 0AEROPOSTALE INC None Provided 007865108 9 412 SH DEFINED 4 0 412 0AEROPOSTALE INC None Provided 007865108 3597 166383 SH DEFINED 73 163260 0 3123ADVANCED MICRO Equity 007903107 23 2895 SH DEFINED 13 2895 0 0ADVANCED MICRO Equity 007903107 1782 222239 SH DEFINED 15 222239 0 0ADVANCED MICRO Equity 007903107 1978 246572 SH DEFINED 3 59090 0 187482ADVANCED MICRO Equity 007903107 44 5458 SH DEFINED 4 5208 250 0ADVANCED MICRO Equity 007903107 205 25533 SH DEFINED 5 25533 0 0ADVANCED MICRO Equity 007903107 1795 223861 SH DEFINED 73 5669 0 218192ADVANCED MICRO Put 007903107 17 2123 Sh Put DEFINED 43 2123 0 0ADVANCED MICRO Put 007903107 17 2121 Sh Put DEFINED 2121 0 0ADVANCED MICRO DEVICES Corporate Bond 007903AL1 285 274000 PRN DEFINED 15 274000 0 0ADVANCED MICRO DEVICES Corporate Bond 007903AN7 9 9000 PRN DEFINED 15 9000 0 0ADVANCED MICRO DEVICES Corporate Bond 007903AN7 4314 4255000 PRN DEFINED 24 4255000 0 0

ADVANCED MICRO DEVICES Corporate Bond 007903AN7 13179 13000000 PRN DEFINED 50 13000000 0 0ADVANCED MICRO DEVICES Corporate Bond 007903AN7 7 7000 SH DEFINED 13 7000 0 0ADVANCED MICRO DEVICES Corporate Bond 007903AN7 1310 1292000 SH DEFINED 15 1292000 0 0AEGON N V Equity 007924103 32 5753 SH DEFINED 13 5753 0 0AEGON N V Equity 007924103 198 35742 SH DEFINED 15 35742 0 0AEGON N V Equity 007924103 415 74561 SH DEFINED 3 74561 0 0AEGON N V Equity 007924103 93 16757 SH DEFINED 4 12016 997 3744ADVANCED ENERGY INDS INC Equity 007973100 1986 151389 SH DEFINED 13 151389 0 0ADVANCED ENERGY INDS INC Equity 007973100 13 983 SH DEFINED 15 983 0 0ADVANCED ENERGY INDS INC Equity 007973100 30 2279 SH DEFINED 24 2279 0 0ADVANCED ENERGY INDS INC Equity 007973100 3919 298701 SH DEFINED 3 16990 0 281711ADVANCED ENERGY INDS INC Equity 007973100 343 26120 SH DEFINED 4 26120 0 0ADVANCED ENERGY INDS INC Equity 007973100 2671 203605 SH DEFINED 73 199767 0 3838ADVENT SOFTWARE INC Equity 007974108 753 29422 SH DEFINED 15 29422 0 0ADVENT SOFTWARE INC Equity 007974108 7 277 SH DEFINED 17 277 0 0ADVENT SOFTWARE INC Equity 007974108 3 116 SH DEFINED 24 116 0 0ADVENT SOFTWARE INC Equity 007974108 104 4056 SH DEFINED 4 2071 385 1600AETERNA ZENARIS INC COMMON STOCK 007975204 214 100000 SH DEFINED 4 100000 0 0AEROSONIC CORP DEL Equity 008015307 215 87200 SH DEFINED 15 87200 0 0AEROVIRONMENT INC Equity 008073108 115 4291 SH DEFINED 13 4291 0 0AEROVIRONMENT INC Equity 008073108 1980 73839 SH DEFINED 15 73839 0 0AEROVIRONMENT INC Equity 008073108 2 59 SH DEFINED 24 59 0 0AEROVIRONMENT INC Equity 008073108 5 200 SH DEFINED 4 200 0 0AES TR III 6.75% PFD Preferred Stock 00808N202 1017 20361 SH DEFINED 15 20361 0 0AES TR III 6.75% PFD Preferred Stock 00808N202 343 6867 SH DEFINED 4 0 0 0AETRIUM INC Equity 00817R103 62 77750 SH DEFINED 15 77750 0 0AETNA INC(NEW) Equity 00817Y108 1234 24610 SH DEFINED 13 24610 0 0AETNA INC(NEW) Equity 00817Y108 47480 946568 SH DEFINED 15 946568 0 0AETNA INC(NEW) Equity 00817Y108 15 290 SH DEFINED 17 290 0 0AETNA INC(NEW) Equity 00817Y108 665 13248 SH DEFINED 3 0 0 13248AETNA INC(NEW) Equity 00817Y108 5106 101804 SH

DEFINED 4 83289 5341 13174AETNA INC(NEW) Equity 00817Y108 13 250 SH DEFINED 7 250 0 0AETNA INC(NEW) Equity 00817Y108 16620 331347 SH DEFINED 73 132048 0 199299AFFILIATED MANAGERS Equity 008252108 340 3041 SH DEFINED 13 3041 0 0AFFILIATED MANAGERS Equity 008252108 4571 40885 SH DEFINED 15 40885 0 0AFFILIATED MANAGERS Equity 008252108 13 118 SH DEFINED 17 118 0 0AFFILIATED MANAGERS Equity 008252108 257 2300 SH DEFINED 24 2300 0 0AFFILIATED MANAGERS Equity 008252108 45948 410949 SH DEFINED 3 174019 0 236930AFFILIATED MANAGERS Equity 008252108 62030 554779 SH DEFINED 4 523799 6392 24588AFFILIATED MANAGERS Equity 008252108 1601 14315 SH DEFINED 5 14315 0 0AFFILIATED MANAGERS Equity 008252108 54 485 SH DEFINED 7 485 0 0AFFILIATED MANAGERS Equity 008252108 1118 10000 SH DEFINED 10000 0 0AFFILIATED MANAGERS Equity 008252108 1577 14100 Sh DEFINED 43 14100 0 0AFFILIATED MANAGERS Equity 008252108 1577 14100 Sh DEFINED 14100 0 0AMG 3.95 08/38 CONVCORP 008252AL2 3920 3564000 PRN DEFINED 24 3564000 0 0Affiliated Managers Group Corp Debt Secur 008252AL2 5775 5250000 PRN DEFINED 50 5250000 0 0AFFYMAX INC Equity 00826A109 40 3400 SH DEFINED 15 3400 0 0AFFYMAX INC Equity 00826A109 1 125 SH DEFINED 24 125 0 0AFFYMAX INC Equity 00826A109 65 5500 SH DEFINED 4 5000 500 0AFFYMETRIX INC Equity 00826T108 0 100 SH DEFINED 13 100 0 0AFFYMETRIX INC Equity 00826T108 14 3191 SH DEFINED 15 3191 0 0AFFYMETRIX INC Equity 00826T108 1 247 SH DEFINED 24 247 0 0AFFYMETRIX INC Equity 00826T108 107 25000 SH DEFINED 4 25000 0 0AGILENT TECH INC CMNSTOCK 00846U101 269 6042 SH DEFINED 13 6042 0 0AGILENT TECH INC CMNSTOCK 00846U101 11079 248922 SH DEFINED 15 248922 0 0AGILENT TECH INC CMNSTOCK 00846U101 35 778 SH DEFINED 17 778 0 0AGILENT TECH INC CMNSTOCK 00846U101 44662 1003418 SH DEFINED 3 136123 0 867295AGILENT TECH INC CMNSTOCK 00846U101 6425 144358 SH DEFINED 4 121949 7375 15034AGILENT TECH INC CMNSTOCK 00846U101 2406 54065 SH DEFINED 5 54065 0 0AGILENT TECH INC CMNSTOCK 00846U101 11 248 SH DEFINED 7 248 0 0AGILENT TECH INC CMNSTOCK 00846U101 4222 94863 SH DEFINED 73 3672 0 91191

AGILENT TECH INC CMNSTOCK 00846U101 116 2603 SH DEFINED 2603 0 0AGNICO-EAGLE MINES LTD Equity 008474108 29 872 SH DEFINED 13 872 0 0AGNICO-EAGLE MINES LTD Equity 008474108 3429 102748 SH DEFINED 15 102748 0 0AGNICO-EAGLE MINES LTD Equity 008474108 111341 3335566 SH DEFINED 3 387655 0 2947911AGNICO-EAGLE MINES LTD Equity 008474108 111 3322 SH DEFINED 4 3209 113 0AGNICO-EAGLE MINES LTD Equity 008474108 230 6897 SH DEFINED 5 6897 0 0AGENUS INC NEW Equity 00847G705 264 40008 SH DEFINED 15 40008 0 0AGILYSYS INC Equity 00847J105 2 169 SH DEFINED 15 169 0 0AGILYSYS INC Equity 00847J105 1 60 SH DEFINED 24 60 0 0AGREE REALTY CORP Equity 008492100 201 8901 SH DEFINED 15 8901 0 0AGREE REALTY CORP Equity 008492100 1 40 SH DEFINED 24 40 0 0AGREE REALTY CORP Equity 008492100 1321 58500 SH DEFINED 3 58500 0 0AGRIA CORP ADR Equity 00850H103 3 3157 SH DEFINED 15 3157 0 0AGRIUM INC Equity 008916108 196 2266 SH DEFINED 13 2266 0 0AGRIUM INC Equity 008916108 6489 75134 SH DEFINED 15 75134 0 0AGRIUM INC Equity 008916108 2708 31358 SH DEFINED 3 6841 0 24517AGRIUM INC Equity 008916108 335 3880 SH DEFINED 4 3506 124 250AIR METHODS CORP < Equity 009128307 2 22 SH DEFINED 13 22 0 0AIR METHODS CORP < Equity 009128307 1824 20910 SH DEFINED 15 20910 0 0AIR METHODS CORP < Equity 009128307 3 40 SH DEFINED 24 40 0 0AIR METHODS CORP < Equity 009128307 1235 14150 SH DEFINED 4 8700 0 5450AIR LEASE CORP Equity 00912X302 4328 179819 SH DEFINED 15 179819 0 0AIR LEASE CORP Equity 00912X302 590 24501 SH DEFINED 4 24501 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 6370 69387 SH DEFINED 13 69387 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 139331 1517776 SH DEFINED 15 1517776 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 225 2448 SH DEFINED 17 2448 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 57701 628551 SH DEFINED 3 81059 0 547492AIR PRODUCTS & CHEMICALS Equity 009158106 21400 233114 SH DEFINED 4 176970 38569 17575AIR PRODUCTS & CHEMICALS Equity 009158106 701 7635 SH DEFINED 5 7635 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 29 321 SH DEFINED 7 321 0 0AIR PRODUCTS & CHEMICALS Equity 009158106 4848 52814 SH

DEFINED 73 2035 0 50779AIR T INC Equity 009207101 8 801 SH DEFINED 15 801 0 0AIR TRANSPORT S CMNSTOCK 00922R105 1 189 SH DEFINED 24 189 0 0AIR TRANSPORT SERV GRP COM 00922R105 5172 893235 SH DEFINED 2 423450 0 469785AIR TRANSPORT SERVICES Equity 00922R105 0 3 SH DEFINED 15 3 0 0AIRGAS INC DOMESTIC COMMON 009363102 855 9614 SH DEFINED 13 9614 0 0AIRGAS INC DOMESTIC COMMON 009363102 1036 11643 SH DEFINED 15 11643 0 0AIRGAS INC DOMESTIC COMMON 009363102 173872 1954272 SH DEFINED 3 826000 0 1128272AIRGAS INC DOMESTIC COMMON 009363102 32150 361361 SH DEFINED 4 360475 300 586AIRGAS INC DOMESTIC COMMON 009363102 10748 120801 SH DEFINED 5 120801 0 0AIRGAS INC DOMESTIC COMMON 009363102 2312 25981 SH DEFINED 73 664 0 25317LUV 5.25 11/16 CONVHYLD 00949PAD0 6172 4884000 PRN DEFINED 24 4884000 0 0AIXTRON SE-SPONSORED ADR Equity 009606104 0 0 SH DEFINED 15 0 0 0AIXTRON SE-SPONSORED ADR Equity 009606104 15 855 SH DEFINED 4 855 0 0AKAMAI TECH INC Equity 00971T101 2340 63773 SH DEFINED 13 63773 0 0AKAMAI TECH INC Equity 00971T101 1704 46424 SH DEFINED 15 46424 0 0AKAMAI TECH INC Equity 00971T101 247 6722 SH DEFINED 3 0 0 6722AKAMAI TECH INC Equity 00971T101 242 6602 SH DEFINED 4 5148 143 1311AKAMAI TECH INC Equity 00971T101 1636 44588 SH DEFINED 73 1718 0 42870AKAMAI TECH INC PUT 00971T101 4 100 SH PUT DEFINED 15 100 0 0AKORN INC Equity 009728106 76 6482 SH DEFINED 15 6482 0 0AKORN INC Equity 009728106 2 197 SH DEFINED 24 197 0 0AKORN INC Equity 009728106 46040 3935050 SH DEFINED 3 342600 0 3592450AKORN INC Equity 009728106 922 78803 SH DEFINED 4 77128 0 1675AKORN INC Equity 009728106 4 310 SH DEFINED 5 310 0 0ALAMO GROUP INC Equity 011311107 15 501 SH DEFINED 15 501 0 0ALAMO GROUP INC Equity 011311107 1 24 SH DEFINED 24 24 0 0ALASKA AIR GROUP INC Equity 011659109 12 325 SH DEFINED 13 325 0 0ALASKA AIR GROUP INC Equity 011659109 1195 33375 SH DEFINED 15 33375 0 0ALASKA AIR GROUP INC Equity 011659109 9 252 SH DEFINED 24 252 0 0ALASKA AIR GROUP INC Equity 011659109 61374 1713410 SH DEFINED 3 183210 0 1530200

ALASKA AIR GROUP INC Equity 011659109 225 6272 SH DEFINED 73 1474 0 4798ALASKA COMMUNICATIONS Equity 01167P101 79 25500 SH DEFINED 13 25500 0 0ALASKA COMMUNICATIONS Equity 01167P101 259 84136 SH DEFINED 15 84136 0 0ALASKA COMMUNICATIONS Equity 01167P101 0 159 SH DEFINED 24 159 0 0ALASKA COMMUNICATIONS Equity 01167P101 4 1150 SH DEFINED 4 1150 0 0ALBANY INTERNATIONAL Equity 012348108 50 2200 SH DEFINED 15 2200 0 0ALBANY INTERNATIONAL Equity 012348108 2 97 SH DEFINED 24 97 0 0ALBANY INTL CORP Corporate Bond 012348AC2 5 5000 SH DEFINED 13 5000 0 0ALBANY MOLECULAR Equity 012423109 0 100 SH DEFINED 13 100 0 0ALBANY MOLECULAR Equity 012423109 0 56 SH DEFINED 15 56 0 0ALBANY MOLECULAR Equity 012423109 0 77 SH DEFINED 24 77 0 0ALBEMARLE CORP Equity 012653101 7 109 SH DEFINED 13 109 0 0ALBEMARLE CORP Equity 012653101 1410 22051 SH DEFINED 15 22051 0 0ALBEMARLE CORP Equity 012653101 37 580 SH DEFINED 17 580 0 0ALBEMARLE CORP Equity 012653101 2957 46261 SH DEFINED 4 44161 399 1701ALBEMARLE CORP Equity 012653101 2184 34160 SH DEFINED 73 19292 0 14868ALCOA INC Call 013817101 106 10600 Sh Call DEFINED 4 10600 0 0ALCOA INC Equity 013817101 1031 102927 SH DEFINED 13 102927 0 0ALCOA INC Equity 013817101 12027 1200304 SH DEFINED 15 1200304 0 0ALCOA INC Equity 013817101 404 40307 SH DEFINED 3 0 0 40307ALCOA INC Equity 013817101 1282 127989 SH DEFINED 4 101465 8140 18384ALCOA INC Equity 013817101 3680 367241 SH DEFINED 73 16780 0 350461ALCOA INC Equity 013817101 239 23864 SH DEFINED 23864 0 0ALCOA INC Corporate Bond 013817AT8 2 1000 PRN DEFINED 24 1000 0 0ALCATEL-LUCENT ADR Equity 013904305 11 4627 SH DEFINED 13 4627 0 0ALCATEL-LUCENT ADR Equity 013904305 679 299190 SH DEFINED 15 299190 0 0ALCATEL-LUCENT ADR Equity 013904305 37 16343 SH DEFINED 4 15914 429 0ALCATEL-LUCENT ADR Equity 013904305 0 100 SH DEFINED 7 0 0 100ALDERON IRON ORE CORP Equity 01434T100 0 30 SH DEFINED 15 30 0 0ALEXANDER AND BALDWIN Equity 014482103 62 1276 SH DEFINED 15 1276 0 0ALEXANDER AND BALDWIN Equity 014482103 13 270 SH

DEFINED 17 270 0 0ALEXANDER AND BALDWIN Equity 014482103 10929 225570 SH DEFINED 2 106420 0 119150ALEXANDER AND BALDWIN Equity 014482103 6104 125985 SH DEFINED 3 114310 0 11675ALEXANDER AND BALDWIN Equity 014482103 1192 24593 SH DEFINED 4 13207 9107 2279ALERE INC Equity 01449J105 10 400 SH DEFINED 13 400 0 0ALERE INC Equity 01449J105 60 2296 SH DEFINED 15 2296 0 0ALERE INC Equity 01449J105 10239 393650 SH DEFINED 2 185180 0 208470ALERE INC Equity 01449J105 59011 2268793 SH DEFINED 3 1084245 0 1184548ALERE INC Equity 01449J105 585 22502 SH DEFINED 4 21471 158 873ALERE INC Equity 01449J105 4117 158289 SH DEFINED 5 158289 0 0ALERE INC PFD CONV PFD COM 01449J204 93 384 SH DEFINED 3 384 0 0ALERE INC 3.00 % PFD PREFERRED STOCK 01449J204 181 749 SH DEFINED 4 0 0 0ALERE INC CONV SER B PFD Preferred Stock 01449J204 0 0 SH DEFINED 15 0 0 0ALEXANDERS I CMNSTOCK 014752109 3 7 SH DEFINED 24 7 0 0ALEXANDERS INCORPORATED Equity 014752109 0 0 SH DEFINED 15 0 0 0ALEXANDRIA REAL ESTATE Equity 015271109 1 16 SH DEFINED 13 16 0 0ALEXANDRIA REAL ESTATE Equity 015271109 33 448 SH DEFINED 15 448 0 0ALEXANDRIA REAL ESTATE Equity 015271109 792 10824 SH DEFINED 4 7813 79 2932ALEXION PHARMACEUTICALS Equity 015351109 1400 15079 SH DEFINED 13 15079 0 0ALEXION PHARMACEUTICALS Equity 015351109 6791 73134 SH DEFINED 15 73134 0 0ALEXION PHARMACEUTICALS Equity 015351109 665533 7167061 SH DEFINED 3 1583432 0 5583629ALEXION PHARMACEUTICALS Equity 015351109 2257 24304 SH DEFINED 4 22748 509 1047ALEXION PHARMACEUTICALS Equity 015351109 38824 418088 SH DEFINED 5 418088 0 0ALEXCO RESOURCE CORP Equity 01535P106 3 400 SH DEFINED 15 400 0 0ALEXA PHARMACEUTICALS Equity 015384100 4 7200 SH DEFINED 15 7200 0 0ALEXA PHARMACEUTICALS Equity 015384100 0 400 SH DEFINED 4 400 0 0ALICO INC CMNSTOCK 016230104 0 13 SH DEFINED 24 13 0 0ALICO INC COMMON STOCK 016230104 5 200 SH DEFINED 4 200 0 0ALICO INC Equity 016230104 0 1 SH DEFINED 15 1 0 0ALIGN TECH INC Equity 016255101 2 61 SH DEFINED 13 61 0 0ALIGN TECH INC Equity 016255101 3794 137707 SH DEFINED 15 137707 0 0

ALIGN TECH INC Equity 016255101 6 215 SH DEFINED 24 215 0 0ALIGN TECH INC Equity 016255101 30710 1114702 SH DEFINED 3 99868 0 1014834ALIGN TECH INC Equity 016255101 19 673 SH DEFINED 4 0 0 673ALIGN TECH INC Equity 016255101 2 79 SH DEFINED 5 79 0 0ALIMERA SCIENCE CMNSTOCK 016259103 0 40 SH DEFINED 24 40 0 0ALLEGHANY CORP NEW Equity 017175100 786 2389 SH DEFINED 15 2389 0 0ALLEGHANY CORP NEW Equity 017175100 6 17 SH DEFINED 17 17 0 0ALLEGHANY CORP NEW Equity 017175100 862 2620 SH DEFINED 4 2321 171 128ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 11872 288372 SH DEFINED 13 288372 0 0ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 2304 55951 SH DEFINED 15 55951 0 0ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 9908 240655 SH DEFINED 3 0 0 240655ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 4375 106278 SH DEFINED 4 104739 365 1174ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 41 991 SH DEFINED 5 991 0 0ALLEGHENY TECHNOLOGIES INC DOMESTIC COMMON 01741R102 1102 26758 SH DEFINED 73 1031 0 25727ALLEGHENY TECHS INC Corporate Bond 01741RAD4 937 752000 PRN DEFINED 24 752000 0 0ALLEGHENY TECHS INC Corporate Bond 01741RAD4 5 4000 PRN DEFINED 4 0 0 0ALLEGHENY TECHS INC Corporate Bond 01741RAD4 10 8000 SH DEFINED 15 8000 0 0ALLEGIANT TRAVEL CO Equity 01748X102 1 26 SH DEFINED 13 26 0 0ALLEGIANT TRAVEL CO Equity 01748X102 1824 33459 SH DEFINED 15 33459 0 0ALLEGIANT TRAVEL CO Equity 01748X102 3 52 SH DEFINED 24 52 0 0ALLEGIANT TRAVEL CO Equity 01748X102 320 5880 SH DEFINED 4 5880 0 0ALLERGAN INC Equity 018490102 1937 20293 SH DEFINED 13 20293 0 0ALLERGAN INC Equity 018490102 22740 238290 SH DEFINED 15 238290 0 0ALLERGAN INC Equity 018490102 11 112 SH DEFINED 17 112 0 0ALLERGAN INC Equity 018490102 184525 1933614 SH DEFINED 3 449501 0 1484113ALLERGAN INC Equity 018490102 165967 1739146 SH DEFINED 4 1687718 43968 7460ALLERGAN INC Equity 018490102 1303 13657 SH DEFINED 5 13657 0 0ALLERGAN INC Equity 018490102 10 100 SH DEFINED 7 100 0 0ALLERGAN INC Equity 018490102 7690 80587 SH DEFINED 73 2941 0 77646ALLETE INC COM Equity 018522300 14 349 SH DEFINED 13 349 0 0ALLETE INC COM Equity 018522300 54 1302 SH

DEFINED 15 1302 0 0ALLETE INC COM Equity 018522300 23589 568554 SH DEFINED 3 39184 0 529370ALLETE INC COM Equity 018522300 768 18521 SH DEFINED 4 15819 1198 1504ALLIANCE BANKSHARES CORP Equity 018535104 13 3100 SH DEFINED 15 3100 0 0ALLIANCE CALIF MUNICIPAL Closed-End Equi 018546101 1335 90743 SH DEFINED 15 90743 0 0ALLIANCE DATA SYSTEMS Equity 018581108 394 3129 SH DEFINED 13 3129 0 0ALLIANCE DATA SYSTEMS Equity 018581108 16054 127456 SH DEFINED 15 127456 0 0ALLIANCE DATA SYSTEMS Equity 018581108 11 85 SH DEFINED 17 85 0 0ALLIANCE DATA SYSTEMS Equity 018581108 264635 2100941 SH DEFINED 3 176993 0 1923948ALLIANCE DATA SYSTEMS Equity 018581108 1901 15090 SH DEFINED 4 14561 143 386ALLIANCE DATA SYSTEMS Equity 018581108 2034 16148 SH DEFINED 5 16148 0 0ALLIANCE DATA SYSTEMS Equity 018581108 20433 162219 SH DEFINED 73 130285 0 31934ADS 1.75 08/13 CONVHYLD 018581AD0 1807 1116000 PRN DEFINED 24 1116000 0 0ALLIANCE HEALTH CMNSTOCK 018606202 0 86 SH DEFINED 24 86 0 0ALLIANCE HEALTHCARE Equity 018606202 0 1 SH DEFINED 15 1 0 0ALLIANCE HOLDINGS GP LP Equity 01861G100 758 17403 SH DEFINED 15 17403 0 0ALLIANCE HOLDINGS GP LP Equity 01861G100 44 1000 SH DEFINED 4 1000 0 0ALLIANCE BERSTEIN NATL Closed-End Equi 01864U106 30 2047 SH DEFINED 13 2047 0 0ALLIANCE BERSTEIN NATL Closed-End Equi 01864U106 762 51478 SH DEFINED 15 51478 0 0ALLIANCE BERSTEIN NATL Closed-End Equi 01864U106 18 1230 SH DEFINED 4 1230 0 0ALLIANCE NEW YORK MUNI Closed-End Equi 018714105 15 1000 SH DEFINED 15 1000 0 0ALLIANCE NEW YORK MUNI Closed-End Equi 018714105 59 4000 SH DEFINED 4 4000 0 0ALLIANCE ONE IN CMNSTOCK 018772103 1 305 SH DEFINED 24 305 0 0ALLIANCE ONE INTERNATIONAL IN COMMON STOCK 018772103 10641 2822639 SH DEFINED 4 1962328 860311 0ALLIANCE ONE INTERNATNL Equity 018772103 0 4 SH DEFINED 15 4 0 0ALLIANCE RESOURCES Equity 01877R108 408 6789 SH DEFINED 13 6789 0 0ALLIANCE RESOURCES Equity 01877R108 1101 18320 SH DEFINED 15 18320 0 0ALLIANCE RESOURCES Equity 01877R108 606 10075 SH DEFINED 4 10075 0 0ALLIANCEBERNSTEIN GLOBAL Closed-End Equi 01879R106 406 27044 SH DEFINED 13 27044 0 0ALLIANCEBERNSTEIN GLOBAL Closed-End Equi 01879R106 3112 207162 SH DEFINED 15 207162 0 0ALLIANCEBERNSTEIN GLOBAL Closed-End Equi 01879R106 280 18663 SH DEFINED 4 15193 0 3470

ALLIANT ENERGY CORP Equity 018802108 64 1475 SH DEFINED 13 1475 0 0ALLIANT ENERGY CORP Equity 018802108 3299 76152 SH DEFINED 15 76152 0 0ALLIANT ENERGY CORP Equity 018802108 645 14899 SH DEFINED 4 9290 3758 1851ALLIANT ENERGY CORP Equity 018802108 21 489 SH DEFINED 7 489 0 0ALLIANT ENERGY CORP Equity 018802108 2271 52417 SH DEFINED 73 29152 0 23265ALLIANT TECHSYSTEMS INC Equity 018804104 558 11136 SH DEFINED 15 11136 0 0ALLIANT TECHSYSTEMS INC Equity 018804104 6698 133640 SH DEFINED 2 62040 0 71600ALLIANT TECHSYSTEMS INC Equity 018804104 304 6075 SH DEFINED 4 4777 223 1075ALLIANT TECHSYSTEMS INC Equity 018804104 339 6758 SH DEFINED 73 1365 0 5393ALLIANT TECHSYSTEMS INC Corporate Bond 018804AK0 1729 1695000 PRN DEFINED 24 1695000 0 0ALLIANT TECHSYSTEMS INC Corporate Bond 018804AK0 1055 1034000 SH DEFINED 15 1034000 0 0ALLIANCEBERSTEIN INC FND Closed-End Equi 01881E101 140 17064 SH DEFINED 13 17064 0 0ALLIANCEBERSTEIN INC FND Closed-End Equi 01881E101 2047 249958 SH DEFINED 15 249958 0 0ALLIANCEBERSTEIN INC FND Closed-End Equi 01881E101 66150 8076932 SH DEFINED 3 3526976 0 4549956ALLIANCEBERSTEIN INC FND Closed-End Equi 01881E101 194 23744 SH DEFINED 4 18065 5679 0ALLIANCEBERNSTEIN HLDG Equity 01881G106 65 4166 SH DEFINED 13 4166 0 0ALLIANCEBERNSTEIN HLDG Equity 01881G106 2234 143096 SH DEFINED 15 143096 0 0ALLIANCEBERNSTEIN HLDG Equity 01881G106 34 2210 SH DEFINED 4 1210 1000 0ALLIANCE BANCORP INC OF Equity 01890A108 0 1 SH DEFINED 15 1 0 0ALLIANCE FINANC CMNSTOCK 019205103 1 17 SH DEFINED 24 17 0 0ALLIANCE FINL CORP Equity 019205103 0 0 SH DEFINED 15 0 0 0ALLIED HEALTHCARE Equity 019222108 610 177718 SH DEFINED 15 177718 0 0ALLIED HEALTHCARE Equity 019222108 4910 1431585 SH DEFINED 3 882869 0 548716ALLIED HEALTHCARE Equity 019222108 1 255 SH DEFINED 4 255 0 0ALLIED HEALTHCARE Equity 019222108 292 85036 SH DEFINED 5 85036 0 0ALLIED MOTION Equity 019330109 8 1054 SH DEFINED 15 1054 0 0ALLIED NEVADA GOLD CORP Equity 019344100 3 92 SH DEFINED 13 92 0 0ALLIED NEVADA GOLD CORP Equity 019344100 20 627 SH DEFINED 15 627 0 0ALLIED NEVADA GOLD CORP Equity 019344100 17 531 SH DEFINED 4 531 0 0ALLOS THERAPEUTICS INC Equity 019777101 0 150 SH DEFINED 15 150 0 0ALLOS THERAPEUTICS INC Equity 019777101 0 276 SH

DEFINED 24 276 0 0ALLSCRIPTS HEALTHCARE SOLUTIO DOMESTIC COMMON 01988P108 103 6210 SH DEFINED 13 6210 0 0ALLSCRIPTS HEALTHCARE SOLUTIO DOMESTIC COMMON 01988P108 491 29573 SH DEFINED 15 29573 0 0ALLSCRIPTS HEALTHCARE SOLUTIO DOMESTIC COMMON 01988P108 20445 1231600 SH DEFINED 3 148100 0 1083500ALLSCRIPTS HEALTHCARE SOLUTIO DOMESTIC COMMON 01988P108 375 22583 SH DEFINED 4 22583 0 0ALLSTATE CORP Equity 020002101 491 14914 SH DEFINED 13 14914 0 0ALLSTATE CORP Equity 020002101 3204 97337 SH DEFINED 15 97337 0 0ALLSTATE CORP Equity 020002101 20623 626448 SH DEFINED 3 11262 0 615186ALLSTATE CORP Equity 020002101 11101 337215 SH DEFINED 4 224250 84083 28882ALLSTATE CORP Equity 020002101 117 3546 SH DEFINED 5 3546 0 0ALLSTATE CORP Equity 020002101 31 943 SH DEFINED 7 573 0 370ALLSTATE CORP Equity 020002101 4356 132335 SH DEFINED 73 4814 0 127521ALMOST FAMILY INC Equity 020409108 13 493 SH DEFINED 15 493 0 0ALMOST FAMILY INC Equity 020409108 1 29 SH DEFINED 24 29 0 0ALNYLAM PHARMACEUTICALS Equity 02043Q107 168 15200 SH DEFINED 15 15200 0 0ALNYLAM PHARMACEUTICALS Equity 02043Q107 4173 376990 SH DEFINED 2 140870 0 236120ALNYLAM PHARMACEUTICALS Equity 02043Q107 2 158 SH DEFINED 24 158 0 0ALON HOLDINGS BLUE ADR Equity 02051Q106 0 0 SH DEFINED 15 0 0 0ALON USA ENERGY INC Equity 020520102 1 100 SH DEFINED 15 100 0 0ALON USA ENERGY INC Equity 020520102 0 40 SH DEFINED 24 40 0 0ALON USA ENERGY INC Equity 020520102 83 9137 SH DEFINED 73 2150 0 6987ALPHA NATURAL RESOURCES Equity 02076X102 385 25314 SH DEFINED 13 25314 0 0ALPHA NATURAL RESOURCES Equity 02076X102 18276 1201564 SH DEFINED 15 1201564 0 0ALPHA NATURAL RESOURCES Equity 02076X102 1 75 SH DEFINED 24 75 0 0ALPHA NATURAL RESOURCES Equity 02076X102 1191 78317 SH DEFINED 3 0 0 78317ALPHA NATURAL RESOURCES Equity 02076X102 10397 683570 SH DEFINED 4 424479 448 258643ALPHA NATURAL RESOURCES Equity 02076X102 839 55175 SH DEFINED 73 2126 0 53049ALPHA NATURAL RESOURCES Corporate Bond 02076XAA0 138 155000 PRN DEFINED 15 155000 0 0ALPHATEC HOLDINGS INC Equity 02081G102 9 4000 SH DEFINED 15 4000 0 0ALPHATEC HOLDINGS INC Equity 02081G102 0 189 SH DEFINED 24 189 0 0ALPINE GLOBAL DYNAMIC Closed-End Equi 02082E106 181 29936 SH DEFINED 15 29936 0 0

ALPINE GLOBAL PREMIER Closed-End Equi 02083A103 99 15202 SH DEFINED 13 15202 0 0ALPINE GLOBAL PREMIER Closed-End Equi 02083A103 2301 353944 SH DEFINED 15 353944 0 0ALPINE GLOBAL PREMIER Closed-End Equi 02083A103 509 78278 SH DEFINED 3 67442 0 10836ALPINE GLOBAL PREMIER Closed-End Equi 02083A103 228 35117 SH DEFINED 4 35117 0 0ALPINE TOTAL DYNAMIC Closed-End Equi 021060108 278 58749 SH DEFINED 13 58749 0 0ALPINE TOTAL DYNAMIC Closed-End Equi 021060108 1076 226937 SH DEFINED 15 226937 0 0ALPINE TOTAL DYNAMIC Closed-End Equi 021060108 10 2200 SH DEFINED 4 2200 0 0ALTAIR NANOTECHNOLOGIES Equity 021373204 0